GENERIC REVISION OF PHORIDAE OF THE NEARCTIC REGION AND PHYLOGENETIC CLASSIFICATION OF PHORIDAE, SCIADOCERIDAE, AND IRONOMYIIDAE (DIPTERA: PHORIDEA)

1992 ◽  
Vol 124 (S164) ◽  
pp. 3-144 ◽  
Author(s):  
Brian V. Brown

AbstractThe Nearctic Region genera of Phoridae are revised, and it is shown that the current classification into three subfamilies is inadequate. The hypothesis that the most closely related taxa to the Phoridae are the Sciadoceridae and Ironomyiidae is supported. Use of these outgroups to polarize character states in the Phoridae allows the reorganization of this family into five subfamilies: Hypocerinae, Phorinae, Aenigmatiinae, Conicerinae, and Metopininae. The newly re-defined Phorinae is the adelphotaxon (sister-group) of the Aenigmatiinae + Conicerinae + Metopininae, among which the relationships are unresolved. The hypocerines are the adelphotaxon of all other extant Phoridae. Within the Hypocerinae the relationships of the 15 included world genera are hypothesized, and three new genera are proposed. The relationships of the eight world phorine genera are only partly resolved. The Aenigmatiinae is organized into two tribes: Aenigmatiini and Diplonevrini. The relationships of the seven world genera of Diplonevrini are hypothesized, whereas those of the Aenigmatiini are not discussed. The six world conicerine genera are revised and their relationships hypothesized. In the Metopininae, two major groups are recognized, the Metopina-group and the Megaselia-group. Beckerina Malloch is the adelphotaxon of these two groups, whereas Rhopica Schmitz and Triphleba Rondani are successive outgroups of all metopinines. The Metopina-group is mostly tropical in distribution and is not discussed further. The Megaselia-group is partially organized into the Gymnophora-subgroup and the Apocephalus-subgroup. Species-level revisions of genera are needed before further progress can be made in this family, because the distribution of character states is insufficiently known. Genera in great need of revision include Peromitra Enderlein, Chaetopleurophora Schmitz, Megaselia Rondani, and Apocephalus Coquillett.

ZooKeys ◽  
2019 ◽  
Vol 815 ◽  
pp. 1-114 ◽  
Author(s):  
Ryan A. St Laurent ◽  
Akito Y. Kawahara

A backbone molecular phylogeny of Mimallonidae, based on 47 species and 515 loci, was recently published. That study resolved some of the major relationships in the family, but taxon sampling was limited and a classification of the family was not formally presented for all species. Here morphological phylogenetic analyses in parsimony and maximum likelihood (ML) frameworks were conducted that included 192 species and 55 morphological characters. A phylogenetic analysis was also conducted on the morphological dataset with a topological constraint based on the 515 locus tree from the previous study. Results show that nearly all species can be confidently placed in a genus using morphological phylogenetics. The presence of a frenulum, a character that was historically used to distinguish major groups of Mimallonidae, varies within and among genera. Based on our phylogenetic results, the classification of Mimallonidae is revised, which now includes 291 species in 41 genera. Descriptions of three new genera are included: Fatellallagen. n., Citrallagen. n., and Lepismallagen. n. The following taxonomic changes were made in the present article: 43 new/revived combinations (in Aceclostria Vuillot, Arcinnus Herbin, Bedosia Schaus, Bedosiallo St Laurent & Kawahara, Cicinnus Blanchard, Citralla, Druentica Strand, Fatellalla, Lacosoma Grote, Lepismalla, Mimallo Hübner, Procinnus Herbin, Psychocampa Grote, Roelmana Schaus, and Thaelia Herbin), two new species-level synonyms (melini Bryk is synonymized with viemanda Schaus, jaruga Jones is synonymized with hamata Walker), one revived synonymy (roscida Dognin is resynonymized with externa Moore), seven new statuses (in Druentica, Macessoga Schaus, and Trogoptera Herrich-Schäffer), six revived statuses (in Aceclostria, Cicinnus, Druentica, Psychocampa, and Zaphanta Dyar), and one new designation of nomen nudum. In order to alleviate nomenclatural problems, twelve lectotypes are designated (for Tolypidaamaryllis (Schaus), Trogopteraalthora Schaus, Adalgisacroesa Schaus, Alheitapulloides (Dognin), LacosomabriasiaSchaus, Lacosomadiederica Schaus, Lacosomaraydela Schaus, Psychocampalacuna (Schaus), Cicinnuscorallina Dognin, Cicinnuslatris Schaus, Cicinnussolvens Schaus, Cicinnustuisana Schaus) as well as a neotype for Mimallodespecta Walker (= Cicinnusdespecta). This paper also provides apomorphies for each genus and a morphological key to genera. Annotations are given to aid researchers in understanding all changes made herein, and images of male and female and their genitalia are present for nearly all type species.


1989 ◽  
Vol 3 (6) ◽  
pp. 721
Author(s):  
DA Duckhouse

Australasian pericomoids, psychodids resembling northern hemisphere species of Pericoma Walker (tribe Pericomini), are mostly members of the tribe Maruinini, here re-defined. Amongst Maruinini, they are like several neotropical genera named by Enderlein (1937), but their actual relationship to Enderlein's genera, and hence their identity, has been a long-standing taxonomic problem. Consideration of extensive new collections and observations made in the southern hemisphere now shows that they consist of the following: Genus Notiocharis Eaton. Tribe Maruinini: genus Didicrum Enderlein, and five new genera, Eremolobulosa, Rotundopteryx, Alloeodidicrurn, Satchellomyia and Ancyroaspis. Of these, the Australian Eremolobulosa is the possible sister group of the European Lobulosa Szabo, and the New Zealand genera Satchellomyia and Ancyroaspis are possible sister groups. Of Enderlein's neotropical genera, five classified by Quate (1963) as synonyms, or in one case a subgenus, of Pericoma (Didicrum, Desmioza, Synmormia, Syntomolaba and Podolepria) are recognised as full genera. A key to Australasian pericomoid genera is provided.


1927 ◽  
Vol 64 (7) ◽  
pp. 308-322 ◽  
Author(s):  
F. R. Cowper Reed

Since the publication of the paper by the present author in 1905 on the Classification of the Phacopidae, a considerable advance has been made in our knowledge of this family as a result of further and better material being obtained, and of new discoveries in different parts of the world. Many new genera and subgenera have been instituted, and modifications or limitations of some of the old terms have been introduced by various authors. The work of Wedekind, Clarke, Rud. and E. Richter, and Kozlowski has specially dealt with questions of classification, but there is still a considerable amount of diversity in the usage and application of the generic and subgeneric names.


2019 ◽  
Vol 50 (5) ◽  
pp. 702-716 ◽  
Author(s):  
Veronica Pereyra ◽  
Adriano Cavalleri ◽  
Claudia Szumik ◽  
Christiane Weirauch

The New World family Heterothripidae (~90 spp., four genera) comprises flower-feeding and ectoparasitic thrips. The monophyly of the group has remained untested and species-level relationships were unknown. Morphological (123 characters) and molecular (28S rDNA D2 and D3-D5, H3, and partial COI) data were compiled to reconstruct phylogenetic relationships of this group. The ingroup was represented by 65 species of the four recognized Heterothripidae genera (Aulacothrips Hood, Heterothrips Hood, Lenkothrips De Santis & Sureda, and Scutothrips Stannard). The monophyly of Heterothripidae was recovered in the total evidence and molecular data only analyses with the ectoparasitic Aulacothrips placed as the sister group of the remaining Heterothripidae. The large genus Heterothrips (>80% of the species-level diversity), which was thoroughly sampled in our analyses (56 species), was recovered as paraphyletic with respect to Scutothrips and Lenkothrips. We conclude that additional morphological and molecular data would be desirable before revising the classification of Heterothripidae


Zootaxa ◽  
2006 ◽  
Vol 1234 (1) ◽  
pp. 1 ◽  
Author(s):  
RICHARD T. THOMPSON

The Holarctic weevil genus Procas Stephens, 1831 is revised for the first time. Six species are recognized: four in the western Palaearctic Region, one in the eastern Palaearctic Region and one in the Nearctic Region. One new species, P. michaelis, sp. n. is described from Spain and one new subspecies, P. picipes levantinus, ssp. n. is described from the Levant. P. picipes (Marsham, 1802), sp. rev. and P. p. steveni (Gyllenhal, 1835), ssp. rev. are released from synonymy with P. armillatus (Fabricius, 1801). The species are separated using, among others, characters derived from sternite 8 of the male which is here used at species level for the first time.        Related genera are discussed. Notodermus Desbrochers, 1875, gen. rev. and Apachiscelus Desbrochers, 1875, gen. rev. are released from synonymy with Procas. Pseudypera Voss, 1936, is returned to synonymy with Notodermus (stat. rev.). Syrdariellia Ter-Minassian, 1978 = Theanellus Reitter, 1912, syn. n. Hypera siccensis (Normand, 1951), comb. n. (ex Procas) = H. pollux (Fabricius, 1796), syn. n. Procas cottyi Perris, 1864 = P. armillatus (Fabricius, 1801), syn. rev. The following new combinations are made in Theanellus: T. alepensis (Pic), comb. n. (ex Procas); T. alternans (Faust, 1885), comb. n. (ex Procas); T. testaceus (Bajtenov, 1974), comb. n. (ex Procas); T. antoinei (Hustache, 1932), comb. n. (ex Procas); T. fastidiosus (Pic, 1904b), comb. n. (ex Procas).        The relationships of the taxa are shown in a simple dendrogram, together with their habitats which vary from very wet to very dry.


2006 ◽  
Vol 91 (11) ◽  
pp. 4235-4236 ◽  
Author(s):  
Pinchas Cohen

Abstract Context: Recent developments in the IGF field have raised questions on whether this is the right time to redefine IGF deficiency. Objective: In this controversy, arguments are made against the need for redefining IGF deficiency at this moment, suggesting instead to wait for further clinical developments. Case: Although a number of rare case reports of IGF deficiency with precise molecular etiologies have been described, the vast majority of the cases remain clinically defined and without a genetic diagnosis. Interventions: Because IGF products are now available for clinical use in IGF-deficient patients, we are still using GH stimulation and static IGF levels as our only clinical diagnostic and classification tools. Positions: We need to develop additional clinical tools, side by side with molecular tools, for the diagnosis and subclassification of IGF deficiency. Chief among these are the IGF-generation test for identification of GH-insensitive patients and genetic panels of polymorphic changes in relevant genes. Conclusions: Until further progress is made in the clinical classification of IGF deficiency, we should not change the current classification, and, when we do, it should be the responsibility of the relevant societies in the field to conduct a consensus statement on the topic first.


1981 ◽  
Vol 12 (3) ◽  
pp. 362-379
Author(s):  
Gaden. Robinson

AbstractTwenty specimens of Tinissa collected by the Danish Noona Dan Expedition (1961–62), the Royal Geographical Society Expedition to Gunung Mulu (1978) and by two individual collectors are recorded. Two new species are described: Tinissa yaloma from New Britain and T. classeyi from Borneo. A revised phylogenetic classification of Tinissa incorporating the new species is proposed. A critique is presented of Kuznetzov & Stekolnikov's classification of the Tineoidea and their alliance of the subfamilies Nemapogoninae and Scardiinae is questioned. The genital musculature of Tinissa is described. The affinities of the Tinissinae are debated and it is concluded that Tinissinae and Scardiinae form a monophyletic group, the Tinissinae being either the sister-group of the Scardiinae or forming a monophyletic group within the Scardiinae.


1989 ◽  
Vol 3 (6) ◽  
pp. 721 ◽  
Author(s):  
DA Duckhouse

Australasian pericomoids, psychodids resembling northern hemisphere species of Pericoma Walker (tribe Pericomini), are mostly members of the tribe Maruinini, here re-defined. Amongst Maruinini, they are like several neotropical genera named by Enderlein (1937), but their actual relationship to Enderlein's genera, and hence their identity, has been a long-standing taxonomic problem. Consideration of extensive new collections and observations made in the southern hemisphere now shows that they consist of the following: Genus Notiocharis Eaton. Tribe Maruinini: genus Didicrum Enderlein, and five new genera, Eremolobulosa, Rotundopteryx, Alloeodidicrurn, Satchellomyia and Ancyroaspis. Of these, the Australian Eremolobulosa is the possible sister group of the European Lobulosa Szabo, and the New Zealand genera Satchellomyia and Ancyroaspis are possible sister groups. Of Enderlein's neotropical genera, five classified by Quate (1963) as synonyms, or in one case a subgenus, of Pericoma (Didicrum, Desmioza, Synmormia, Syntomolaba and Podolepria) are recognised as full genera. A key to Australasian pericomoid genera is provided.


2001 ◽  
Vol 75 (6) ◽  
pp. 1141-1151 ◽  
Author(s):  
Richard A. Fortey

The progress achieved in trilobite systematics over the last 75 years is briefly reviewed. Different approaches to phylogenetics have influenced the way trilobites have been classified. Classical evolutionary taxonomy, the stratigraphical approach, and cladistics have all contributed in different ways to the current classification, which has evolved piecemeal, and is still unsatisfactory is some ways. Nonetheless, progress towards a phylogenetic classification has been made, especially as the result of information from ontogenies provided by well-preserved silificified material. Trilobites are a well-defined clade within a larger arachnomorph group. Agnostida have been excluded from Trilobita, but are perhaps best considered as specialised trilobites, at least until limbs of eodiscids are described. The outstanding problems in classification of each trilobite order are reviewed. Most are concerned with the recognition of the appropriate Cambrian sister taxa, and the discovery of the relevant ontogenies. It is very likely that post-Cambrian clades “root” deeply into the Cambrian. The coherence, or otherwise, of Proetida, Asaphida, Corynexochida and the lichid/odontopleurid groups will be resolved by such studies. The problems of paraphyly in Ptychopariida and Redlichiina may prove more obdurate. The temporal brevity of certain Cambrian family ranges may be partly a taxonomic artefact. The possibility of a late Cambrian gap in the record on some clades should be considered.


1996 ◽  
Vol 70 (4) ◽  
pp. 641-655 ◽  
Author(s):  
Jörg Maletz ◽  
Charles E. Mitchell

New data on the proximal structure of several glossograptid species, including Paraglossograptus holmi, indicate that symmetry between sicula and the first theca is ubiquitous and primitive for the group. This isograptid symmetry is shared with species of Isograptus and Oncograptus, among others, as is a dicalycal second theca, and the initial downward growth of the first several thecal pairs. Consideration of these and other features of the rhabdosome structure leads us to reassess the phylogenetic relations among Glossograptus, Pseudisograptus, Isograptus, and their relatives. The Glossograptidae is a holophyletic clade comprising the common ancestor of Isograptus caduceus and I. victoriae and all its descendants, including the monopleural glossograptines. Incipient monopleural development is exhibited by the otherwise Isograptus-like Bergstroemograptus crawfordi. Derived glossograptids, such as Glossograptus and Kalpinograptus, share a fully monopleural rhabdosome with a prominent proximal bulge formed by the overlap of the early thecae. Contrary to some claims, none possesses a manubrium or manubrium-like structure. The manubrium and the arienigraptid suture are restricted to Arienigraptus hastatus and its descendants. These features unite the Arienigraptidae. The Arienigraptidae, based on their possession of isograptid symmetry and lack of any derived characters shared uniquely with the glossograptids, appear to be a sister group of the Glossograptidae rather than a stem group. These two families appear most closely allied to certain reclined, two-stiped species, such as Isograptus? dilemma, that retain tetragraptid proximal symmetry.


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