The effect of stand structure and stand density on the leaf area–sapwood area relationship of lodgepole pine

1989 ◽  
Vol 19 (3) ◽  
pp. 392-396 ◽  
Author(s):  
Dan C. Thompson

The relationship of sapwood area to leaf area in lodgepole pine was examined across a variety of habitat types and stand densities in northwest Montana. No statistical differences were found between plots with regard to either habitat type or stand density. A nonlinear relationship was found between leaf area and sapwood area. Increasing amounts of sapwood were associated with a decrease in the leaf area–sapwood area ratio. A large amount of within-plot variation in the sapwood area–leaf area relationship was explained by differences between dominant trees and trees of other crown classes. Leaf area (LA) was best estimated by the equation LA = 0.12 × S − 0.0003 × S2 + 0.06 × S × D, where LA is leaf area, S is sapwood area, and D is the crown class (dominant). Differences between dominant and subdominant trees appear to be related to ring width and its associated permeability. Differences in sapwood area–leaf area equations among different studies may be due in part to differences in stand structure.

1991 ◽  
Vol 21 (12) ◽  
pp. 1760-1764 ◽  
Author(s):  
Steven B. Jack ◽  
James N. Long

It is commonly assumed that mature forest stands with closed canopies support constant amounts (weight or area) of foliage, independent of stand density. For stand leaf area to be constant, mean leaf area must be plastic with respect to density. We examined the relationship between density and both leaf area index and mean leaf area for two contrasting tree species, lodgepole pine (Pinuscontorta var. latifolia Engelm.) and subalpine fir (Abieslasiocarpa (Hook.) Nutt.). In lodgepole pine, leaf area index tended to be constant over a wide range of absolute and relative densities, but in subalpine fir, leaf area index increased with density. Consistent with these results, mean leaf area of lodgepole pine was more plastic with respect to density than mean leaf area of subalpine fir. The presumption of stable leaf area index independent of stand density, therefore, may not be as general as usually assumed owing to differential responses of mean leaf area to density. Differences in plasticity between the two species were attributed to differences in relative shade tolerance and the effect of shade on competitive interactions at high densities.


1995 ◽  
Vol 25 (6) ◽  
pp. 970-977 ◽  
Author(s):  
Annikki Mäkelä ◽  
Katri Virtanen ◽  
Eero Nikinmaa

The effects of the average ring width of sapwood, bole length, and stand density on the relationship between foliage biomass and sapwood area at four different stem positions were studied in a data set comprising 20 ca. 35-year-old Scots pine (Pinussylvestris L.) trees in southern Finland. The average ring width within sapwood had no effect on the foliagersapwood ratio inside the crown, but a correlation was found when sapwood was measured at breast height or 20% relative height. The distance of the measuring point from the crown base provided a bigger improvement of foliage biomass prediction from sapwood, a finding emphasizing the significance of the taper of sapwood along the bole. After accounting for the sapwood taper, no differences could be detected between thinned and unthinned stands, nor between breast height and 20% relative height.


1987 ◽  
Vol 17 (2) ◽  
pp. 174-180 ◽  
Author(s):  
M. A. Espinosa Bancalari ◽  
D. A. Perry ◽  
John D. Marshall

The relationship between foliage area and sapwood basal area was studied in three adjacent 22-year-old Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) stands that differed in early growth rates. Sapwood width was fairly constant for most of the stem above the stump, but the number of annual rings in the sapwood decreased gradually with height. Sapwood area also decreased with increasing height in the tree, the stands differing significantly only at breast height. The proportion of heartwood from stump to near the base of the crown was significantly higher for the stand of fastest early growth. Ratios of leaf area to sapwood area were significantly higher for that stand and varied in every stem section, the ratio lower at breast height than at the base of the live crown. At the base of the crown, the ratio of leaf area to sapwood area was 1.33 and 1.57 times greater in the fast-growing stand than in the intermediate- and slow-growing stands, respectively. Leaf area was as closely related to dbh as to sapwood area at breast height. Sapwood area at the crown base was more accurate than sapwood area at breast height for predicting leaf area in the fast stand and was equally accurate in the other two stands. Ratios of leaf area to sapwood area correlated positively with sapwood ring width. However, because sapwood ring width also correlated closely with sapwood area, it did not improve predictive equations. The results suggest that the "pipe model" theory must be modified to account for the internal structure of the "pipe" and that caution should be exercised when using published leaf area to sapwood area ratios.


1988 ◽  
Vol 18 (2) ◽  
pp. 247-250 ◽  
Author(s):  
James N. Long ◽  
Frederick W. Smith

Leaf area to sapwood area ratios for a given species are believed to vary with factors such as site quality, stand density, early stand growth rates, and crown class. Based on data from 55 mature lodgepole pine trees (Pinuscontorta var. latifolia Dougl.) from 10 plots in southeastern Wyoming, we conclude that putative density and site effects on leaf area - sapwood area relations are actually a consequence of the increase in the leaf area to sapwood area ratio with increasing sapwood area. When leaf area is estimated with a nonlinear model that includes tree size and distance to the live crown, the apparent effects of stand density and site index disappear. We consider a constant ratio of leaf area and sapwood cross-sectional area to be inappropriate for the estimation of leaf area aross the range of stand conditions included in most studies of forest ecology.


2005 ◽  
Vol 94 (2) ◽  
pp. 244-255 ◽  
Author(s):  
Quan Wang ◽  
Samuel Adiku ◽  
John Tenhunen ◽  
André Granier

2004 ◽  
Vol 34 (11) ◽  
pp. 2217-2229 ◽  
Author(s):  
Douglas B Mainwaring ◽  
Douglas A Maguire

Basal area and height growth were analyzed for individual trees in uneven-aged ponderosa pine (Pinus ponderosa Dougl. ex Laws.) and lodgepole pine (Pinus contorta Dougl. ex. Loud.) stands in central Oregon. Basal area growth was modeled as a function of other stand and tree variables to address three general objectives: (1) to compare the predictive ability of distance-dependent versus distance-independent stand density variables; (2) to determine the degree to which small trees negatively affect the growth of overstory trees; and (3) to test for differences in growth efficiency between species and between indices of spatial occupancy used to define efficiency (area potentially available, crown projection area, and a surrogate for total tree leaf area). Distance-dependent variables were found to improve growth predictions when added to models with only distance-independent variables, and small trees were found to have a quantifiably negative effect on the growth of larger trees. While volume growth efficiency declined with increasing levels of spatial occupancy for lodgepole pine, ponderosa pine volume growth efficiency was greatest at the highest levels of crown base sapwood area and crown projection area. The behavior in ponderosa pine resulted from the previously recognized correlation between tree height and total leaf area or crown size. The final statistical models distinguished between the positive effect of relative height and the negative effect of increasing tree size.


2012 ◽  
Vol 2012 ◽  
pp. 1-7 ◽  
Author(s):  
Aymer Andrés Vásquez-Ordóñez ◽  
Inge Armbrecht ◽  
Gabriela Pérez-Lachaud

Eucharitidae are parasitoids that use immature stages of ants for their development.KapalaCameron is the genus most frequently collected in the Neotropics, but little is known about the biology and behavior of any of the species of this genus. We aimed to evaluate the effect of habitat type on eucharitid parasitism and to contribute to the knowledge of the host-parasite relationship betweenKapalasp. and the poneromorph antEctatomma ruidum(Roger) in Colombia. TwentyE. ruidumcolonies were extracted from two different habitat types (woodland and grassland), and larvae and cocoons (pupae) were examined in search for parasitoids in different stages of development. Globally, 60% of the colonies were parasitized, with 1.3% of larvae and 4% of pupae parasitized. Planidia (first-instar larvae), pupae, and adults of the parasitoid were observed. All of the pupae and adult parasitoids belonged toKapala iridicolorCameron. All the colonies collected in the woodlands were parasitized and contained more parasitized larvae (2%) and parasitized cocoons (8%) than those collected in grasslands (4/12 parasitized colonies, 0.5% parasitized larvae, 0.8% parasitized cocoons). The relationship observed between habitat type and parasitism prevalence is a novel aspect of the study of eucharitid impact on ant host populations.


2014 ◽  
Vol 4 (16) ◽  
pp. 3218-3235 ◽  
Author(s):  
Anthony P. Walker ◽  
Andrew P. Beckerman ◽  
Lianhong Gu ◽  
Jens Kattge ◽  
Lucas A. Cernusak ◽  
...  

2014 ◽  
Vol 90 (04) ◽  
pp. 489-497 ◽  
Author(s):  
William C. Parker

The relationship of stand structural features with understory light levels, estimated by gap light index (GLI), was investigated in 22 second-growth eastern white (Pinus strobus L.) and red pine (Pinus resinosa Ait.)-dominated stands in central Ontario that encompassed a broad range in density and basal area. Simple, empirical light models were developed to quantify the influence of several stand structural variables on canopy transmittance as estimated by GLI. Models were also derived to facilitate the operational identification of residual basal area, density, and percent canopy closure associated with target understory light levels that optimize the growth of white pine regeneration and its protection from weevil and blister rust when using the uniform shelterwood silvicultural system. Regression models indicated significant negative, nonlinear relationships of GLI with density, basal area, a stand density index, total crown area, and foliar biomass, while GLI was linearly related to percent canopy closure. Application of these models to identify density, basal area, and canopy closure values associated with target light levels for the regeneration and removal cuts of uniform shelterwoods demonstrates the use of this information to help guide management of white pine–red pine forests.


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