A new ceratopsid from the Foremost Formation (middle Campanian) of Alberta

2012 ◽  
Vol 49 (10) ◽  
pp. 1251-1262 ◽  
Author(s):  
Michael J. Ryan ◽  
David C. Evans ◽  
Kieran M. Shepherd

Xenoceratops foremostensis gen. et. sp. nov., a new centrosaurine ceratopsid from the Foremost Formation (Campanian) of Alberta, is described based on frill material from at least three adult-sized individuals collected from a low-density bone bed. The material can be assigned to Centrosaurinae based on features of the preserved squamosal. Although the parietals are incomplete, the shape of the diagnostic parietal can be inferred from several overlapping serial elements. The parietal of the new taxon shares with all other centrosaurines, except Centrosaurus apertus , spike-like ornamentation at the posterolateral (P3) locus under traditional coding methods. At approximately 78 Ma, it is the oldest known Canadian ceratopsid, approximately 0.5 Ma older than Albertaceratops from the lower Oldman Formation of Canada and approximately 1.0 Ma younger than Diabloceratops from the Wahweap Formation of Utah. A phylogenetic analysis resolves the new taxon as the basalmost centrosaurine and places Centrosaurus brinkmani as the sister taxon to Styracosaurus albertensis . The type species of Centrosaurus brinkmani is moved to a new genus.

Zootaxa ◽  
2010 ◽  
Vol 2481 (1) ◽  
Author(s):  
CARLOS MOLINERI

The 12 species previously placed in Tortopus together with 3 species newly described here, are revised and included in a phylogenetic analysis. Based on synapomorphic characters on the nymphs and adults of both sexes, Tortopus is restricted to T. igaranus Needham & Murphy, T. circumfluus Ulmer, T. harrisi Traver, T. zottai (Navás), T. bellus Lugo-Ortiz & McCafferty, and T. arenales sp. nov., and the genus is defined by: female parastyli receptors with long furrows anterior to sockets; penes entirely flattened; male ninth abdominal sternum almost separated in two portions by a median notch; mesosternum with furcasternal protuberances contiguous only on basal corner; and nymphs with two subapical tubercles on mandibular tusks. Tortopsis is newly described for T. bruchianus (Navás), T. limoncocha sp. nov., T. obscuripennis (Domínguez), T. parishi (Banks), T. primus (McDunnough), T. puella (Pictet), T. sarae (Domínguez), T. spatula sp. nov., and T. unguiculatus (Ulmer). Tortopsis is characterized by: R sector of female fore wing without additional veins between R 2 and IR; female parastyli receptors C or V-shaped, with sockets opening towards median line; male gonopore associated with a claw-like structure; penes separated from the base; parastyli more than 5 times length of pedestals; parastyli curved in lateral view; nymphs with a single subapical tubercle on mandibular tusks. The study of available type material permitted inclusion of comparative diagnoses, with figures and redescriptions as needed. The male imago of the type species of Tortopus (T. igaranus Needham & Murphy) is described for the first time, as are the female adults of Tortopus bellus Lugo-Ortiz & McCafferty and T. harrisi Traver. Three new Neotropical species based on male and female adults are described: Tortopus arenales and Tortopsis limoncocha from Ecuador, and Tortopsis spatula from Colombia. Keys to separate the adults and nymphs of the genera of Polymitarcyidae, and for male and female adults of all the species of Tortopus and Tortopsis are presented, as well as line drawings, pictures and SEM photographs of important structures.


PeerJ ◽  
2018 ◽  
Vol 6 ◽  
pp. e5300
Author(s):  
Terry A. Gates ◽  
Khishigjav Tsogtbaatar ◽  
Lindsay E. Zanno ◽  
Tsogtbaatar Chinzorig ◽  
Mahito Watabe

We describe a new iguanodontian ornithopod,Choyrodon barsboldigen. et sp. nov. from the Albian-aged Khuren Dukh Formation of Mongolia based on several partial skeletons interpreted to represent a subadult growth stage based on osteohistological features. This new taxon is diagnosed by many autapomorphies of the maxilla, nasal, lacrimal, opisthotic, predentary, and surangular.Choyrodondisplays an unusual combination of traits, possessing an open antorbital fenestra (a primitive ornithopod trait) together with derived features such as a downturned dentary and enlarged narial fenestra. Histological imaging suggests that the type specimen ofChoyrodonwould have been a subadult at the time of death. Phylogenetic analysis of two different character matrices do not positChoyrodonto be the sister taxon or to be more primitive than the iguanodontianAltirhinus kurzanovi, which is found in the same formation. The only resolved relationship of this new taxon is that it was hypothesized to be a sister-taxon with the North American speciesEolambia caroljonesa. Though discovered in the same formation andChoyrodonbeing smaller-bodied thanAltirhinus, it does not appear that the former species is an ontogimorph of the latter. Differences in morphology and results of the phylogenetic analyses support their distinction although more specimens of both species will allow better refinement of their uniqueness.


2018 ◽  
Vol 58 (2) ◽  
pp. 523-529
Author(s):  
Alexey Reshchikov ◽  
Cornelis Van achterberg

Abstract Gilen gen. nov. (type species G. orientalis sp. nov.) from South East Asia is described and illustrated. The new taxon has a produced mid-longitudinal facial projection, which differentiates it from all other known genera of Ichneumonidae. It belongs to the tribe Perilissini in the subfamily Ctenopelmatinae (Hymenoptera, Ichneumonidae) and is recorded from Laos, Northern Thailand and Vietnam.


2019 ◽  
Vol 94 (3) ◽  
pp. 568-579
Author(s):  
Alexander O. Averianov

AbstractKazachostylops occidentalis Nesov, 1987b, based on partial maxilla and dentary from the upper Paleocene Zhylga locality in South Kazakhstan, is redescribed. A new phylogenetic hypothesis of Arctostylopida is proposed based on phylogenetic analysis of 26 characters and 17 taxa. Kazachostylops is recovered as a sister taxon to the Arctostylopinae, the advanced clade of Asian and North American arctostylopids characterized by pseudohypocone on upper molars and reduced trigonid of lower molars, with the ectolophid being attached labial on the trigonid. Kazachostylops differs from more basal arctostylopids (Asiostylops, Allostylops, Bothriostylops, and Wanostylops) by higher-crowned molars, M1–3 metaconule absent, m1–3 entoconid connected with ectolophid by entolophid, and m2 wider than m1 and m3. Principal component analyses of the upper and lower dentition of arctostylopids show great distinctness of Kazachostylops from other members of the group. The arctostylopid taxa are reviewed, and the new genus Enantiostylops is erected for ‘Sinostylops’ progressus Tang and Yan, 1976 from the lower Eocene of China, because of uniquely concave parastylar area on upper molars.UUID: http://zoobank.org/a46d8f29-fd73-4e59-88dc-fcc55b12d1d3


2019 ◽  
Vol 9 (1) ◽  
Author(s):  
Adele H. Pentland ◽  
Stephen F. Poropat ◽  
Travis R. Tischler ◽  
Trish Sloan ◽  
Robert A. Elliott ◽  
...  

Abstract The Australian pterosaur record is poor by world standards, comprising fewer than 20 fragmentary specimens. Herein, we describe the new genus and species Ferrodraco lentoni gen. et sp. nov., based on the most complete pterosaur specimen ever found in Australia, and the first reported from the Winton Formation (Cenomanian–lower Turonian). The presence of premaxillary and mandibular crests, and spike-shaped teeth with subcircular bases, enable Ferrodraco to be referred to Anhangueria. Ferrodraco can be distinguished from all other anhanguerian pterosaurs based on two dental characters: the first premaxillary and mandibular tooth pairs are small; and the fourth–seventh tooth pairs are smaller than the third and eighth ones. Ferrodraco was included in a phylogenetic analysis of Pterosauria and resolved as the sister taxon to Mythunga camara (upper Albian Toolebuc Formation, Australia), with that clade occupying the most derived position within Ornithocheiridae. Ornithocheirus simus (Albian Cambridge Greensand, England), Coloborhynchus clavirostris (Valanginian Hastings Sands, England), and Tropeognathus mesembrinus (upper Aptian–lower Albian Romualdo Formation, Brazil) were resolved as successive sister taxa, which suggests that ornithocheirids were cosmopolitan during the Albian–Cenomanian. Furthermore, the stratigraphic age of Ferrodraco lentoni (Cenomanian–lower Turonian) implies that anhanguerians might have survived later in Australia than elsewhere.


Zootaxa ◽  
2009 ◽  
Vol 2096 (1) ◽  
pp. 287-298 ◽  
Author(s):  
ELKE WILLEN ◽  
JANINE DITTMAR

A new species of the new genus Keraia gen. nov. from the Guinea Basin is described in the present paper. The new taxon belongs to the Pseudomesochrinae Willen, 1996, which until now has contained only the genus Pseudomesochra T. Scott, 1902. Both genera are prevalent in the deep sea. Whereas Pseudomesochra reaches quite high individual and species numbers in the investigated samples of the DIVA 1, DIVA 2 and ANDEEP deep-sea expeditions, Keraia is found only occasionally and as single specimens. Keraia is characterised among others by a modified antenna exopodite and the shape of P1. Pseudomesochra on the other hand can still be identified as a monophylum by specialised setation on the P1 endopodite and the lack of the inner setae of the first segment of swimming leg exopodites. Other species of Keraia gen. nov. are K. longiseta (Vasconcelos, George & Santos 2008) and the type species K. tamara (Smirnov, 1946). The available records implicate a widespread occurrence of Keraia gen. nov. from the northern to the southern Atlantic, and even reaching both northern and southern polar regions. Up to now all individuals that have been found occur exclusively at deep-sea sites and in very low abundances.


2017 ◽  
Vol 98 (6) ◽  
pp. 1365-1381
Author(s):  
Luiz Ricardo L. Simone

Some Antarctic littorinoideans have a remarkable convergence with Naticoidea in shell and operculum features. Two naticid-like species of that group are studied in their phenotypic features in order to improve their taxonomy and to discuss the meaning of that convergence, as the former are herbivore-detritivore and the latter active predatory organisms. One of the studied species is the littorinidLaevilacunaria antarctica(Martens, 1885). The other belongs to a new genus –Pseudonatica, with the type species also newly described:P. antarctica, the genus is tentatively placed in Zerotulidae. Another Pseudonatica is also described,P. ampullarica, based only on shells collected by Marion-Dufresne French expedition off Brazilian coast, this finding expands the occurrence of zerotulids northwards. Besides the similarities of shell and operculum, other structures of these Antarctic species also show singular similarities with naticoideans, such as the wide foot, the complexity of opercular attachment in pedal opercular pad, the wide oesophageal gland, and the coiled arrangement of the pallial oviduct. The phenotypic characters were coded and inserted in a previous large phylogenetic analysis on Caenogastropoda (Simone, 2011), furnishing a wide basis for discussion on the characters, taxonomic position, evolution and adaptations of these organisms.


Fossil Record ◽  
2001 ◽  
Vol 4 (1) ◽  
pp. 71-82 ◽  
Author(s):  
D. Weyer

<i>Muenstraia</i> n. gen. ist eine der ältesten ahermatypischen Rugosa (Subordo Cyathaxoniina) und umfasst neben der Typusart <i>Muenstraia franconica</i> n. sp. (Ludlovium, Elbersreuther Orthoceratitenkalk, Frankenwald) drei weitere Arten: <i>Muenstraia squarrosa</i> (Sutherland, 1965) (unteres Ludlovium, Henryhouse-Formation, Oklahoma), <i>Muenstraia</i> sp. (oberes Lochkovium, Yukon-Gebiet), <i>Muenstraia thuringica</i> n. sp. (Pragium, Thüringisches Schiefergebirge und Tafilalt). Die Gattung kann von dem isolierten, nur aus Xinjiang bekannten Protozaphrentis Yü, 1957 des hohen Mittelordoviz abgeleitet werden; wichtige Deszendenten im Ludlovian sind <i>Laccophyllum</i> Simpson, 1900 und Sutherlandinia Weyer, 1972. <br><br> Der Bauplan entspricht dem seit Schindewolf (1931) traditionellen Konzept der Gattung <i>Petraia</i> Münster, 1839, die aber nach Revision (Weyer 2000) ihrer wahren Typusart <i>Petraia decussata</i> Münster, 1839 aus dem oberen Famennium einer anderen Entwicklungsreihe angehört (Neaxoninae Hill, 1981, jetzt Petraiidae Koninck, 1872). Für die dadurch namenlos gewordene Familia "Petraiidae" (etwa sensu Hill 1981) werden die bisher als Synonym ruhenden Protozaphrentidae Ivanovskiy, 1959 verfügbar, denen noch <i>Duncanella</i> Nicholson, 1874 sowie die Sutherlandiniinae Weyer, 1972 und die Ditoecholasmatinae Sutherland, 1965 zugeordnet sind. <br><br> Muenstraia, a new genus of Rugosa (Anthozoa) from the Late Silurian and Early Devonian <br><br> The new taxon, one of the most ancient members of the ahermatypic suborder Cyathaxoniina, includes the type species <i>Muenstraia franconica</i> n. sp. (Ludlovian, Elbersreuth <i>Orthoceratites</i>-Limestone Formation. Upper Franconia, Germany) and three further species: <i>Muenstraia squarrosa</i> (Sutherland, 1965) (lower Ludlovian, Henryhouse Formation, Oklahoma, USA), <i>Muenstraia</i> sp. (upper Lochkovian, Yukon Territories, Canada). <i>Muenstraia thuringica</i> n. sp. (middle/upper Pragian, Tentaculitid Limestone Formation, Thuringian Mountains, Germany, and middle Pragian, Tafilalt, Morocco). The genus descends from the isolated Upper Middle Ordovician <i>Protozaphrentis</i> Yü, 1957, only known from Xinjiang in China; it is the ancestor of two new phylogenetic lines starting in the Ludlovian with <i>Laccophyllum</i> Simpson, 1900, and <i>Sutherlandinia</i> Weyer, 1972. <br><br> Morphology and diagnosis are identical with the (since Schindewolf 1931) traditional definition of the genus <i>Petraia</i> Münster, 1839, which represents according to a revision of its real and Upper Famennian type species <i>Petraia</i> <i>decussata</i> Münster, 1839 (Weyer 2000) another phylogenetic line (Neaxoninae Hill, 1981, now Petraiidae Koninck, 1872). Therefore, the valid name of the family "Petraiidae" (sensu Hill 1981) becomes Protozaphrentidae Ivanovskiy, 1959, which comprise also <i>Duncanella</i> Nicholson, 1874, and both the Sutherlandiniinae Weyer, 1972 and Ditoecholasmatinae Sutherland, 1965. <br><br> doi:<a href="http://dx.doi.org/10.1002/mmng.20010040106" target="_blank">10.1002/mmng.20010040106</a>


2021 ◽  
pp. 27-34
Author(s):  
Mark Wilkinson

A new genus and species of rhinatrematid caecilian, Amazops amazops gen. et sp. nov., is described based on a single specimen from Orellana, Ecuador collected in 1990. Among other features the new taxon differs from all other rhinatrematid caecilians in having less than four annular grooves interrupted in the region of the vent and in the squamosal contributing to the bony margin of the orbit. A consideration of its distinctive morphology suggests that it is plausible that the new taxon is the sister taxon of all other rhinatrematid caecilians. That the genus is known from a single specimen, and that this is the first new rhinatrematid species from the Andes described for more than 50 years, highlights the poor sampling (collecting) of rhinatrematid caecilians and limited knowledge of their diversity.


2021 ◽  
Author(s):  
Huang Shi-Ke ◽  
Kevin D. Hyde ◽  
Ausana Mapook ◽  
Sajeewa S.N. Maharachchikumbura ◽  
D. Jayarama Bhat ◽  
...  

Abstract Sordariomycetes is an earlier and one of the widely distributed class of Ascomycota. The class was initially classified based on morphology in having inoperculate and unitunicate asci. With the development of DNA based phylogenetic analysis, several undetermined or polyphyletic members of Sordariomycetes were reclassified. However, not all species belonging to this class have been sequenced and analyzed. There are a number of species, especially those old and poorly studied ones which have never been sequenced before and not even recollected again for further taxonomic verification. One of the main objective in this study is to revise and update the taxonomy of several well-known old and poorly studied species whose classification are still obscure. Herein, we re-examined the type materials and/or authentic specimens together to explore 74 relatively poorly-studied genera, which mainly belong to Boliniales, Calosphaeriales, Chaetosphaeriales, Jobellisiales, and Sordariales classified under Diaporthomycetidae and Sordariomycetidae. We provide descriptions, notes, figures and/or drawings and discussed their phylogenetic relationships. As a result, the monotypic Jobellisiales is transferred from Hypocreomycetidae to Diaporthomycetidae. Based on phylogenetic analysis, the polyphyletic Lasiosphaeriaceae is divided into five families, Bombardiaceae (Apodospora, Bombardia, Bombardioidea and Fimetariella), Lasiosphaeriaceae (Anopodium, Bellojisia, Corylomyces, Lasiosphaeria, Mammaria and Zopfiella), Lasiosphaeridaceae (Lasiosphaeris), Strattoniaceae (Strattonia) and Zygospermellaceae (Episternus and Zygospermella). In addition, a new family Neoschizotheciaceae is established based on Neoschizothecium. Analysis of the type species of Boothiella, Stellatospora, Sulcatistroma and Tengiomyces placed them in Sordariaceae, Chaetomiaceae, Hypocreales and Coronophorales, respectively. We classify the genera lacking molecular data based on their morphology and expect them to be recollected; that is, Kacosphaeria in Calosphaeriales; Arnium, Biconiosporella, Camptosphaeria, Diffractella, Emblemospora, Eosphaeria, Periamphispora, Ramophialophora, Synaptospora and Tripterosporella in Sordariales; Conidiotheca in Sordariomycetes; Copromyces, Effetia, Endophragmiella and Tulipispora are accommodated in Ascomycota. Besides, we establish a new genus Neoschizothecium based on phylogenetic analysis. New combinations proposed include: Camaropella amorpha, Cam. microspora, Cam. plana, Cladorrhinum grandiusculum, Cla. leucotrichum, Cla. terricola, Cla. olerum, Helminthosphaeria plumbea, Immersiella hirta, Jugulospora minor, Lasiosphaeris arenicola, Neoschizothecium aloides, Neo. carpinicola, Neo. conicum, Neo. curvisporum, Neo. fimbriatum, Neo. glutinans, Neo. inaequale, Neo. minicaudum, Neo. selenosporum, Neo. tetrasporum, Neurospora autosteira, Podospora brunnescens, P. flexuosa, P. jamaicensis, P. hamata, P. macrospora, P. spinosa, Strattonia petrogale and Triangularia microsclerotigena, T. nannopodalis, T. praecox, T. samala, T. tarvisina, T. unicaudata, T. yaeyamensis. New epithets are proposed for Apiorhynchostoma apiosporum and Podospora dacryoidea.


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