Systematics, phylogeny and historical biogeography of the Pentagonaster clade (Asteroidea:Valvatida:Goniasteridae)

2007 ◽  
Vol 21 (4) ◽  
pp. 311 ◽  
Author(s):  
Christopher Mah

Morphology-based phylogenetic hypotheses developed for living and fossil goniasterid asteroids have provided several unique opportunities to study bathymetric and biogeographic shifts for an ecologically important group of prominent, megafaunal invertebrates. A cladistic analysis of 18 ingroup taxa employing 65 morphological characters resulted in a single most parsimonious tree. The tree supports assignment of the Atlantic Tosia parva (Perrier, 1881) and the Pacific Tosia queenslandensis Livingstone, 1932 to new, separate genera. The phylogenetic tree supports offshore to onshore bathymetric shifts between basal and derived taxa. The phylogeny is also consistent with historical events surrounding the separation of Antarctica from Australia and South Africa. Buterminaster Blake & Zinsmeister, 1988 from the Eocene La Meseta Formation, Antarctic Peninsula, was included in the phylogenetic analysis and is now supported as the only fossil species in the genus Pentagonaster Gray, 1840. Pentagonaster stibarus H. L. Clark, 1914 is separated from synonymy with P. dubeni Gray, 1847 and resurrected as a valid species. The new genus, Akelbaster, gen. nov., shows unusual new structures that resemble cribiform organs, although their function has not been determined. One specific ingroup lineage, including Tosia and Pentagonaster, attains a much larger adult size than those of its sister-taxa, suggesting that Cope’s rule may apply to asteroids within this clade. Pentagonaster and related genera are revised. Descriptions of four new genera and three new species are presented, including: Akelbaster novaecaledoniae, gen. nov., sp. nov., Ryukuaster onnae, gen. nov., sp. nov., Eknomiaster beccae, sp. nov., Pawsonaster parvus, gen. nov., comb. nov. and Anchitosia queenslandensis, gen. nov., comb. nov.

Zootaxa ◽  
2018 ◽  
Vol 4526 (3) ◽  
pp. 251 ◽  
Author(s):  
VIRGINIA LEÓN-RÈGAGNON ◽  
JANET TOPAN

Lung flukes of the genus Haematoloechus Looss, 1899 are common parasites of anurans worldwide, but the taxonomy of the group has been confusing. In this taxonomic revision, 89 species of Haematoloechus (= Pneumonoeces Looss, 1902, Ostioloides Odening, 1960, Ostiolum Pratt, 1903, Skrjabinoeces Sudarikov, 1950, Neohaematoloechus Odening, 1960, Metahaematoloechus Yamaguti, 1971) are listed. Of these, 70 are considered valid, three are species inquirendae (H. legrandi Mañé–Garzón & Gil, 1959, H. latoricensis Kozák, 1968 & H. vitelloconfluentum (Rai, 1962) Saeed, Al–Barwari & Al-Harmni, 2007), one is a nomen nudum H. sudarikovi Belouss, 1962, 14 are junior synonyms and one belongs to Ostioloides. This publication also describes three new species, H. occidentalis n. sp., H. veracruzanus n. sp. and H. mexicanus n. sp., parasitizing species of Rana Linnaeus in Mexico and redescribes Haematoloechus caballeroi (Skrjabin & Antipin, 1962) Yamaguti, 1971. The phylogenetic hypotheses based on sequences of mitochondrial and ribosomal DNA of Haematoloechus spp. show that genera proposed on the basis of morphological characters are not supported. The host records for species of Haematoloechus, together with the phylogenetic hypothesis of the genus, suggest that this host-parasite association predates the ranid diversification in the Cretaceous. 


2011 ◽  
Vol 9 (4) ◽  
pp. 709-730 ◽  
Author(s):  
Alejandro Londoño-Burbano ◽  
César Román-Valencia ◽  
Donald C. Taphorn

We review species of Parodon Valenciennes, 1850 from the Magdalena, Cauca, Orinoco, Amazonas, Atrato and Caribbean-Guajira River basins of Colombia using meristic and morphological characters. We recognize eight valid species, five previously described: P. apolinari Myers, from the Orinoco River basin; P. buckleyi Boulenger and P. pongoensis (Allen) from the upper Amazon; P. caliensis Boulenger, from the upper Cauca River drainage; and P. suborbitalis Valenciennes, from Lake Maracaibo basin. Three new species are described: P. alfonsoi, from the lower Magdalena River drainage; P. magdalenensis, from the middle Magdalena and upper Cauca River drainages; and P. atratoensis, from the Atrato River basin. We redescribe Parodon suborbitalis using type specimens and topotypes, and designate lectotypes. A taxonomic key is included for identification of the species, as well as geographic distribution maps.


2013 ◽  
Vol 27 (1) ◽  
pp. 129
Author(s):  
Edilson Caron ◽  
Cibele S. Ribeiro-Costa ◽  
Alfred F. Newton

Rove beetles of the genus Piestus Gravenhorst, 1806 are commonly captured under the bark of or inside decaying logs from Neotropical forests. Piestus belongs to the subfamily Piestinae, historically an ill-defined dumping-ground for Staphylinidae defined by plesiomorphic characters, but which has gradually been restricted in concept and currently includes only six additional extant genera worldwide. Piestinae in this restricted sense has been considered a probably monophyletic subfamily, but its status and phylogenetic position, as a possible sister-group of Osoriinae within the recently proposed Oxyteline group of staphylinid subfamilies, are uncertain and need confirmation. The main aim of the present study was to provide a morphological cladistic analysis and complete taxonomic revision of Piestus, which, as the type and most speciose genus of Piestinae, is critical for future phylogenetic studies involving the subfamily. In our study, the monophyly of Piestus is established and phylogenetic relationships among its species are proposed based on 70 adult morphological characters. Piestus is supported by 11 synapomorphies and high branch support. All species of Piestus are revised and the genus is redefined. The genus contains 43 species, including 13 species described here for the first time. The previously proposed subgenera Antropiestus Bernhauer, 1917, Eccoptopiestus Scheerpeltz, 1952, Elytropiestus Scheerpeltz, 1952, Lissopiestus Scheerpeltz, 1952, Piestus s. str., Trachypiestus Scheerpeltz, 1952 and Zirophorus Dalman, 1821 have not been confirmed, as they were found to be poly- or paraphyletic, or are here removed from Piestus, and therefore subgenera are not used. The main taxonomic changes are as follows. Lissopiestus, syn. nov. is proposed as new synonym of Eleusis Laporte, 1835 and its species, E. interrupta (Erichson, 1840), comb. rest., is transferred again to that genus. Antropiestus, syn. nov. and Eccoptopiestus, syn. nov. are proposed as new synonyms of Hypotelus Erichson, 1839 and their species, H. laevis (Solsky, 1872), comb. nov. and H. andinus (Bernhauer, 1917), comb. nov., are transferred to Hypotelus. Fourteen new synonymies are proposed (valid species listed first): P. lacordairei Laporte, 1835 = Z. furcatus Sharp, 1887, syn. nov.; P. capricornis Laporte, 1835 = P. frontalis Sharp, 1876, syn. nov.; P. pennicornis Fauvel, 1864 = P. plagiatus Fauvel, 1864, syn. nov.; P. rectus Sharp, 1876, syn. nov.; P. pygialis Fauvel, 1902, syn. nov.; P. surinamensis Bernhauer, 1928, syn. nov.; P. minutus Erichson, 1840 = P. nigrator Fauvel, 1902, syn. nov.; P. sulcatus Gravenhorst, 1806 = P. sanctaecatharinae Bernhauer, 1906, syn. nov.; P. condei Wendeler, 1955, syn. nov.; P. gounellei Fauvel, 1902 = P. wasmanni Fauvel, 1902, syn. nov.; P. mexicanus Laporte, 1835 = P. alternans Sharp, 1887, syn. nov.; P. aper Sharp, 1876 = P. schadei Scheerpeltz, 1952, syn. nov.; P. angularis Fauvel, 1864 = P. crassicornis Sharp, 1887, syn. nov.; H. andinus (Bernhauer, 1917) = P. strigipennis Bernhauer, 1921, syn. nov. One species is revalidated: P. fronticornis (Dalman, 1821), stat. rev., and one synonym is restored: P. penicillatus (Dalman, 1821) = P. erythropus Erichson, 1840, syn. rest. Neotypes are designated for P. lacordairei Laporte, 1835 and Oxytelus bicornis Olivier, 1811, and lectotypes are designated for P. puncticollis Fauvel, 1902, P. capricornis variety muticus Fauvel, 1902, P. zischkai Scheerpeltz, 1951, P. pennicornis Fauvel, 1864, P. plagiatus Fauvel, 1864, P. pygmaeus Laporte, 1835, P. niger Fauvel 1864, P. minutus Erichson, 1840, P. nigratror Fauvel, 1902, P. sulcatus Gravenhorst, 1806, P. sanctaecatharinae Bernhauer, 1906, P. sulcipennis Scheerpeltz, 1952, P. aper Sharp, 1876, P. schadei Scheerpeltz, 1952 and P. andinus Bernhauer, 1917.


2004 ◽  
Vol 175 (5) ◽  
pp. 507-512 ◽  
Author(s):  
Isabelle Rouget ◽  
Pascal Neigeet ◽  
Jean-Louis Dommergues

Abstract Two main types of data are available to resolve phylogenies using fossils data: (1) stratigraphic ordering of taxa, and (2) morphological characters. In most phylogenetic studies dealing with ammonites, authors have given priority to the stratigraphic distribution of taxa. This practice is classically justified by the fact that the ammonite fossil record is frequently outstandingly good. In practice, the level of integration of stratigraphic and morphologic information in a single analysis depends on the confidence that authors have in the quality of data. Besides, many evolutionary concepts, which could differ over time and between authors (e.g. anagenesis, cladogenesis, iterative evolution), are added to these data to help infer phylogenetic relationships. As a result, phylogenetic hypotheses are based on eclectic methods which depend on the relative weight given to stratigraphic and morphologic information as well as on evolutionary concepts used. The validity of relationships proposed by previous authors is not dealt with in this paper. Instead, our goal is to draw attention to problems that these eclectic methods may cause, that is to say: (1) ammonites systematics is poorly formalised and (2) phylogenetic hypotheses as they are classically constructed are not rigorously testable. During the last 10 years, cladistic analysis has been applied to ammonites but is still unpopular among ammonitologists. However, studies have consistently shown that cladistics is not as unsuited a tool for ammonites phylogenetic reconstruction as is widely believed. Moreover, classical works open new questions about ammonite phylogeny and in particular, help to reappraise our view on the definition of morphological characters and their phylogenetic significance.


1995 ◽  
Vol 26 (4) ◽  
pp. 393-424 ◽  
Author(s):  
Analia A. Lanteri

AbstractThe genus Ericydeus Pascoe (Polydrosinae: Naupactini) distributed throughout the United States of America up to Argentina is revised and a cladistic analysis including 16 species is conducted. Three new species are described: E. bahiensis, E. argentinensis and E. cupreolus. Ericydeus humeralis Hustache is synonymized under E. nigropunctatus (Chevrolat); and E. modestus viridans (Boheman) and E. modestus duodecimpunctatus (Dalla-Torre, Emden & Emden) are elevated to species rank. Other valid species are: E. hancocki (Kirby), E. schoenherri (Perty), E. sedecimpunctatus (Linnaeus), E. yucatanus (Champion), E. roseiventris (Champion), E. quadripunctatus (Champion), E. modestus (Gyllenhal), E. forreri (Champion), E. lautus (LeConte) and E. placidus (Horn). In the cladogram obtained the species from South America gather in a clade (E. argentinensis, E. sedecimpunctatus, E. nigropunctatus, E. schoenherri, E. hancocki, and E. bahiensis) and the species from Central and North America form a separate clade (E. yucatanus, E. roseiventris - E. quadripunctatus, E. cupreolus, E. viridans - E. modestus, E. duodecimpunctatus, E. forreri, E. lautus - E. placidus). The character evolution follows a southern-northern direction. The paper includes a redescription of the genus, redescriptions or descriptions of its 16 species, a dichotomous key, habitus photographs, drawings of diagnostic structures, distribution maps, a cladogram, and a discussion of the phylogeny and distribution of the genus.


1995 ◽  
Vol 43 (6) ◽  
pp. 557 ◽  
Author(s):  
MS Springer ◽  
LJ Hollar ◽  
JAW Kirsch

Andersen's 1912 monograph on megachiropterans remains the definitive work on the systematics of this group. Andersen argued that the Macroglossinae, containing the eonycterine and notopterine sections, are a monophyletic sister-group to other fruitbats (i.e. Andersen's Rousettus, Cynopterus and Epomophorus sections). Two recent molecular studies (DNA hybridisation and restriction mapping of ribosomal cistrons), as well as an analysis of female reproductive characters, challenge the monophyly of the Macroglossinae and several of Andersen's other conclusions such as the phylogenetic position of Nyctimene. We performed a cladistic analysis on 36 morphological characters, including 33 that were gleaned from Andersen, to determine whether phylogenetic hypotheses based on modem phylogenetic methods are in agreement with Andersen's original conclusions and to compare morphological and molecular phylogenetic hypotheses. Minimum-length trees based on parsimony are largely consistent with Andersen and support (1) a monophyletic Macroglossinae, within which the eonycterine section is paraphyletic with respect to a monophyletic notopterine section, (2) a monophyletic Cynopterus section, excepting the exclusion of Myonycteris, (3) a monophyletic Epomophorus section, excepting the exclusion of Plerotes, and (4) a paraphyletic Rousettus section, with several of the Rousettus-like forms branching off near the base of the tree. Bootstrapping analyses on a reduced data-set that included taxa shared in common with the DNA hybridisation study did not provide strong support (greater than or equal to 95%) for any clades but did provide moderate support (greater than or equal to 70) for several clades, including a monophyletic Macroglossinae. These findings are in marked contrast to the DNA hybridisation phylogeny. A high index of between-data-set incongruence is further evidence for the clash between DNA hybridisation and morphology. A phylogenetic framework was constructed on the basis of morphological data and DNA hybridisation data using a criterion of moderate support and shows little resolution, whereas employing a criterion of strong support produced a framework resolving several additional nodes. One implication of this framework is that characteristic macroglossine features such as a long tongue with a thick carpet of filiform papillae have evolved independently on several occasions (or evolved once and were lost several times). Rates of character evolution for the morphological characters employed in our analysis were calculated using divergence times estimated from DNA hybridisation data. Rates have apparently been fastest in the interior branches, and slower along the external branches, which suggests an early adaptive radiation in the history of fruitbats.


2006 ◽  
Vol 84 (2) ◽  
pp. 225-241 ◽  
Author(s):  
M Manuel

The most recent advances concerning the phylogeny and evolution of calcareous sponges (Calcarea or Calcispongia) are reviewed here, in the light of the history of taxonomy of the group and conceptions about its evolution, starting from Haeckel's works at the end of the 19th century. Calcisponge phylogeny has recently started to be addressed using modern tools of phylogenetic reconstruction: cladistic analysis of morphological characters and molecular phylogeny (so far using 18S and 28S rDNA sequences). The monophyly of calcareous sponges is strongly supported in these works, as is their subdivision into two clades, Calcinea (whose proposed synapomorphy is the basal position of nuclei in choanocytes, with no relation to the flagella) and Calcaronea (whose possible synapomorphy is the formation of the amphiblastula larva through the original process of eversion of the stomoblastula). While the molecular phylogeny of Calcinea is still in its infancy because of insufficient taxonomic sampling, several lines are emerging for the phylogeny of Calcaronea, and these are in strong disagreement with the classification issued from the "traditional" morphological approach. Phylogenetic hypotheses also permit the reconstruction of morphological character evolution, which appears complex and subject to a high level of homoplasy.


ZooKeys ◽  
2021 ◽  
Vol 1025 ◽  
pp. 91-137
Author(s):  
Marília Pessoa-Silva ◽  
Marcos Ryotaro Hara ◽  
Ricardo Pinto-da-Rocha

Species of the genus Sadocus Sørensen, 1886 are conspicuous gonyleptids that occur in Chile and Argentina. Here, the genus is revised for the first time and the cladistic analysis based on morphological characters does not corroborate its monophyly unless a phylogenetically unrelated species is excluded (explained further on). A new classification is proposed for the seven species left in the genus and considered valid, of the 13 nominal species previously recognized. Two out of the seven valid species are considered as species inquirendae: Sadocus allermayeri (Mello-Leitão, 1945) [= Carampangue allermayeri Mello-Leitão, 1945] and Sadocus nigronotatus (Mello-Leitão, 1943) [= Carampangue nigronotatum Mello-Leitão, 1943]. The following synonymies are proposed: Sadocus bicornis (Gervais, 1849) [original combination = Gonyleptes bicornis Gervais, 1849] is a junior synonym of Sadocus asperatus (Gervais, 1847) [= Gonyleptes asperatus Gervais, 1847]; Sadocus conspicillatus Roewer, 1913, Sadocus exceptionalis (Mello-Leitão, 1946) [= Araucanoleptes exceptionalis Mello-Leitão, 1946] and Sadocus guttatus Sørensen, 1902 are junior synonyms of the valid name Sadocus polyacanthus (Gervais, 1847) [= Gonyleptes polyacanthus Gervais, 1847]; and Sadocus calcar (Roewer, 1913) [= Lycomedes calcar Roewer, 1913] is a junior synonym of the valid name Gonyleptes horridus Kirby, 1819. Sadocus brasiliensis Soares & Soares, 1949 is not congeneric with Argentinean/Chilean species of the genus according to the cladistic analysis and is here synonymized with Discocyrtus catharinensis (Mello-Leitão, 1923 [= Sadocus catharinensis Mello-Leitão, 1923]).


Zootaxa ◽  
2021 ◽  
Vol 5027 (1) ◽  
pp. 61-86
Author(s):  
EVERTON E. NAZARÉ-SILVA ◽  
FERNANDO A.B. SILVA

The South American species of Pseudocanthon Bates, 1887 (Coleoptera: Scarabaeidae: Scarabaeinae: Deltochilini) are reviewed. In this work, 10 valid species and two subspecies are recognized for the genus. Among these species, five are recorded in South America, including three new species: P. perplexus (LeConte, 1847), P. xanthurus (Blanchard, 1847), P. vazdemelloi new species, P. pantanensis new species, and P. chaquensis new species. A lectotype is designated for P. xanthurus (Blanchard, 1847). In this revision, each species is analyzed as follows: a detailed literature review, an identification key for the South American species, a diagnosis, descriptions, illustrations of key morphological characters, list of examined material, and geographic distribution.  


Zootaxa ◽  
2017 ◽  
Vol 4232 (4) ◽  
pp. 535 ◽  
Author(s):  
DANIEL BURCKHARDT ◽  
DALVA L. QUEIROZ

The Neotropical psyllid genus Tainarys Brèthes, 1920 is revised to include 14 extant and one fossil species from Dominican amber. Eight species are described as new, viz. Tainarys aroeira sp. nov., T. atra sp. nov., T. hapla sp. nov., T. myracrodrui sp. nov., T. nigricornis sp. nov., T. didyma sp. nov. and T. orientalis sp. nov. from Brazil, the last two also from Uruguay, as well as T. lozadai sp. nov. from Peru. The fifth instar immatures are described for nine species. †Vicinilura Klimaszewski, 1996, erected for the fossil †V. reposta Klimaszewski, 1996 and previously synonymised with Leurolophus Tuthill, 1942, is synonymised here (syn. nov.) with Tainarys and †V. reposta is transferred to become †Tainarys reposta (Klimaszewski), comb. nov. The descriptions are supplemented by illustrations and keys for the identification of adults and immatures. Phylogenetic relationships between species are investigated with a cladistic analysis using 22 adult and six immature morphological characters. The analysis resulted in a single most parsimonious, fully resolved tree. The fossil species is nested within the genus rather than being the sister taxon of the remainder of species. The extant species are restricted to the subtropical and temperate parts of South America. Three pairs of sister clades display an east‒west South American and one a midwest‒southern Brazilian geographical vicariance. Host plants are confirmed for nine and likely for another four species. They are Astronium, Haplorhus, Myracrodruon, Schinopsis and Schinus (Anacardiaceae). All Tainarys species appear to be oligophagous inducing irregular leaf curls on their hosts. 


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