Corrigendum to: Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera : Staphylinidae : Piestinae)

2013 ◽  
Vol 27 (1) ◽  
pp. 129
Author(s):  
Edilson Caron ◽  
Cibele S. Ribeiro-Costa ◽  
Alfred F. Newton
Keyword(s):  
Cladistic Analysis ◽  
Taxonomic Revision ◽  
Sister Group ◽  
Morphological Characters ◽  
Valid Species ◽  
Phylogenetic Position ◽  
Neotropical Forests ◽  
Rove Beetles ◽  
New Synonyms ◽  
Branch Support

Rove beetles of the genus Piestus Gravenhorst, 1806 are commonly captured under the bark of or inside decaying logs from Neotropical forests. Piestus belongs to the subfamily Piestinae, historically an ill-defined dumping-ground for Staphylinidae defined by plesiomorphic characters, but which has gradually been restricted in concept and currently includes only six additional extant genera worldwide. Piestinae in this restricted sense has been considered a probably monophyletic subfamily, but its status and phylogenetic position, as a possible sister-group of Osoriinae within the recently proposed Oxyteline group of staphylinid subfamilies, are uncertain and need confirmation. The main aim of the present study was to provide a morphological cladistic analysis and complete taxonomic revision of Piestus, which, as the type and most speciose genus of Piestinae, is critical for future phylogenetic studies involving the subfamily. In our study, the monophyly of Piestus is established and phylogenetic relationships among its species are proposed based on 70 adult morphological characters. Piestus is supported by 11 synapomorphies and high branch support. All species of Piestus are revised and the genus is redefined. The genus contains 43 species, including 13 species described here for the first time. The previously proposed subgenera Antropiestus Bernhauer, 1917, Eccoptopiestus Scheerpeltz, 1952, Elytropiestus Scheerpeltz, 1952, Lissopiestus Scheerpeltz, 1952, Piestus s. str., Trachypiestus Scheerpeltz, 1952 and Zirophorus Dalman, 1821 have not been confirmed, as they were found to be poly- or paraphyletic, or are here removed from Piestus, and therefore subgenera are not used. The main taxonomic changes are as follows. Lissopiestus, syn. nov. is proposed as new synonym of Eleusis Laporte, 1835 and its species, E. interrupta (Erichson, 1840), comb. rest., is transferred again to that genus. Antropiestus, syn. nov. and Eccoptopiestus, syn. nov. are proposed as new synonyms of Hypotelus Erichson, 1839 and their species, H. laevis (Solsky, 1872), comb. nov. and H. andinus (Bernhauer, 1917), comb. nov., are transferred to Hypotelus. Fourteen new synonymies are proposed (valid species listed first): P. lacordairei Laporte, 1835 = Z. furcatus Sharp, 1887, syn. nov.; P. capricornis Laporte, 1835 = P. frontalis Sharp, 1876, syn. nov.; P. pennicornis Fauvel, 1864 = P. plagiatus Fauvel, 1864, syn. nov.; P. rectus Sharp, 1876, syn. nov.; P. pygialis Fauvel, 1902, syn. nov.; P. surinamensis Bernhauer, 1928, syn. nov.; P. minutus Erichson, 1840 = P. nigrator Fauvel, 1902, syn. nov.; P. sulcatus Gravenhorst, 1806 = P. sanctaecatharinae Bernhauer, 1906, syn. nov.; P. condei Wendeler, 1955, syn. nov.; P. gounellei Fauvel, 1902 = P. wasmanni Fauvel, 1902, syn. nov.; P. mexicanus Laporte, 1835 = P. alternans Sharp, 1887, syn. nov.; P. aper Sharp, 1876 = P. schadei Scheerpeltz, 1952, syn. nov.; P. angularis Fauvel, 1864 = P. crassicornis Sharp, 1887, syn. nov.; H. andinus (Bernhauer, 1917) = P. strigipennis Bernhauer, 1921, syn. nov. One species is revalidated: P. fronticornis (Dalman, 1821), stat. rev., and one synonym is restored: P. penicillatus (Dalman, 1821) = P. erythropus Erichson, 1840, syn. rest. Neotypes are designated for P. lacordairei Laporte, 1835 and Oxytelus bicornis Olivier, 1811, and lectotypes are designated for P. puncticollis Fauvel, 1902, P. capricornis variety muticus Fauvel, 1902, P. zischkai Scheerpeltz, 1951, P. pennicornis Fauvel, 1864, P. plagiatus Fauvel, 1864, P. pygmaeus Laporte, 1835, P. niger Fauvel 1864, P. minutus Erichson, 1840, P. nigratror Fauvel, 1902, P. sulcatus Gravenhorst, 1806, P. sanctaecatharinae Bernhauer, 1906, P. sulcipennis Scheerpeltz, 1952, P. aper Sharp, 1876, P. schadei Scheerpeltz, 1952 and P. andinus Bernhauer, 1917.

Cladistic analysis and revision of Piestus Gravenhorst with remarks on related genera (Coleoptera : Staphylinidae : Piestinae)

2011 ◽  
Vol 25 (6) ◽  
pp. 490 ◽  
Author(s):  
Edilson Caron ◽  
Cibele S. Ribeiro-Costa ◽  
Alfred F. Newton
Keyword(s):  
Cladistic Analysis ◽  
Taxonomic Revision ◽  
Sister Group ◽  
Morphological Characters ◽  
Valid Species ◽  
Phylogenetic Position ◽  
Neotropical Forests ◽  
Rove Beetles ◽  
New Synonyms ◽  
Branch Support

Rove beetles of the genus Piestus Gravenhorst, 1806 are commonly captured under the bark of or inside decaying logs from Neotropical forests. Piestus belongs to the subfamily Piestinae, historically an ill-defined dumping-ground for Staphylinidae defined by plesiomorphic characters, but which has gradually been restricted in concept and currently includes only six additional extant genera worldwide. Piestinae in this restricted sense has been considered a probably monophyletic subfamily, but its status and phylogenetic position, as a possible sister-group of Osoriinae within the recently proposed Oxyteline group of staphylinid subfamilies, are uncertain and need confirmation. The main aim of the present study was to provide a morphological cladistic analysis and complete taxonomic revision of Piestus, which, as the type and most speciose genus of Piestinae, is critical for future phylogenetic studies involving the subfamily. In our study, the monophyly of Piestus is established and phylogenetic relationships among its species are proposed based on 70 adult morphological characters. Piestus is supported by 11 synapomorphies and high branch support. All species of Piestus are revised and the genus is redefined. The genus contains 43 species, including 13 species described here for the first time. The previously proposed subgenera Antropiestus Bernhauer, 1917, Eccoptopiestus Scheerpeltz, 1952, Elytropiestus Scheerpeltz, 1952, Lissopiestus Scheerpeltz, 1952, Piestus s. str., Trachypiestus Scheerpeltz, 1952 and Zirophorus Dalman, 1821 have not been confirmed, as they were found to be poly- or paraphyletic, or are here removed from Piestus, and therefore subgenera are not used. The main taxonomic changes are as follows. Lissopiestus, syn. nov. is proposed as new synonym of Eleusis Laporte, 1835 and its species, E. interrupta (Erichson, 1840), comb. rest., is transferred again to that genus. Antropiestus, syn. nov. and Eccoptopiestus, syn. nov. are proposed as new synonyms of Hypotelus Erichson, 1839 and their species, H. laevis (Solsky, 1872), comb. nov. and H. andinus (Bernhauer, 1917), comb. nov., are transferred to Hypotelus. Fourteen new synonymies are proposed (valid species listed first): P. lacordairei Laporte, 1835 = Z. furcatus Sharp, 1887, syn. nov.; P. capricornis Laporte, 1835 = P. frontalis Sharp, 1876, syn. nov.; P. pennicornis Fauvel, 1864 = P. plagiatus Fauvel, 1864, syn. nov.; P. rectus Sharp, 1876, syn. nov.; P. pygialis Fauvel, 1902, syn. nov.; P. surinamensis Bernhauer, 1928, syn. nov.; P. minutus Erichson, 1840 = P. nigrator Fauvel, 1902, syn. nov.; P. sulcatus Gravenhorst, 1806 = P. sanctaecatharinae Bernhauer, 1906, syn. nov.; P. condei Wendeler, 1955, syn. nov.; P. gounellei Fauvel, 1902 = P. wasmanni Fauvel, 1902, syn. nov.; P. mexicanus Laporte, 1835 = P. alternans Sharp, 1887, syn. nov.; P. aper Sharp, 1876 = P. schadei Scheerpeltz, 1952, syn. nov.; P. angularis Fauvel, 1864 = P. crassicornis Sharp, 1887, syn. nov.; H. andinus (Bernhauer, 1917) = P. strigipennis Bernhauer, 1921, syn. nov. One species is revalidated: P. fronticornis (Dalman, 1821), stat. rev., and one synonym is restored: P. penicillatus (Dalman, 1821) = P. erythropus Erichson, 1840, syn. rest. Neotypes are designated for P. lacordairei Laporte, 1835 and Oxytelus bicornis Olivier, 1811, and lectotypes are designated for P. puncticollis Fauvel, 1902, P. capricornis variety muticus Fauvel, 1902, P. zischkai Scheerpeltz, 1951, P. pennicornis Fauvel, 1864, P. plagiatus Fauvel, 1864, P. pygmaeus Laporte, 1835, P. niger Fauvel 1864, P. minutus Erichson, 1840, P. nigratror Fauvel, 1902, P. sulcatus Gravenhorst, 1806, P. sanctaecatharinae Bernhauer, 1906, P. sulcipennis Scheerpeltz, 1952, P. aper Sharp, 1876, P. schadei Scheerpeltz, 1952 and P. andinus Bernhauer, 1917.

scholarly journals Taxonomic revision and phylogenetic relationships of Dasyloricaria Isbrücker & Nijssen, 1979 (Siluriformes: Loricariidae), with description of a new species

2016 ◽  
Vol 14 (1) ◽  
Author(s):  
Alejandro Londoño-Burbano ◽  
Roberto E. Reis
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
Phylogenetic Relationships ◽  
Taxonomic Revision ◽  
Sister Group ◽  
Valid Species ◽  
Maracaibo Basin ◽  
New Synonyms ◽  
Branchial Arches ◽  
A New Species

ABSTRACT A taxonomic revision and phylogenetic analysis were completed for Dasyloricaria . The genus includes three valid species: D . filamentosa and D . latiura previously included in the genus, and a new species described herein. Dasyloricaria have a restricted trans-Andean distribution, with D . filamentosa occurring at the lower and middle Magdalena, lower Cauca, and Sinu in Colombia, and lago Maracaibo basin in Colombia and Venezuela; D . latiura in the Atrato and the Tuyra basins in Colombia and Panama, respectively; and the new species in the upper and middle Magdalena basin in Colombia. New synonyms for D . filamentosa and D . latiura are proposed, and a lectotype is designated for the latter. Dasyloricaria is herein recognized as monophyletic, with D . filamentosa as the sister group of D . latiura , and the new speciesas sister to that clade. Spatuloricaria is hypothesized to be the sister group of Dasyloricaria based on synapomorphies of the neurocranium, branchial arches and external morphology features. The subtribe Rineloricariina was partially corroborated through the phylogenetic analysis. An identification key for the species of Dasyloricaria is provided.

Cladistic analysis and taxonomic review of the millipede tribe Arthrosolaenomeridini Hoffman, 1976 (Polydesmida: Chelodesmidae)

Zootaxa ◽  
2021 ◽  
Vol 4970 (2) ◽  
pp. 201-256
Author(s):  
RODRIGO SALVADOR BOUZAN ◽  
LUIZ FELIPE M. INIESTA ◽  
ANTONIO DOMINGOS BRESCOVIT
Keyword(s):  
Cladistic Analysis ◽  
Taxonomic Revision ◽  
Sister Group ◽  
Morphological Characters ◽  
New Genera ◽  
Distribution Maps ◽  
West Region ◽  
The Matrix ◽  
First Time ◽  
Parsimony Criterion

Chelodesmidae is one of the most diverse families belonging to the Polydesmida, comprising almost 800 described species distributed in 172 genera, 20 tribes and two subfamilies. Among the tribes within the Neotropical subfamily Chelodesminae, Arthrosolaenomeridini represents a very interesting group occurring in the Central-West region of Brazil. Currently the tribe contains three genera: Arthrosolaenomeris Schubart, 1943, Gangugia Schubart, 1947, and Angelodesmus Schubart, 1962. In this study, we conducted a cladistic analysis including 18 species of Arthrosolaenomeridini and 7 species of other associated Chelodesminae tribes as an outgroup. The matrix is composed of 64 morphological characters from which 39 are herein proposed for the first time. The data were analyzed under the parsimony criterion using TNT 1.5. Based on the results, Arthrosolaenomeridini was recovered as monophyletic with Macrocoxodesmini + Telonychopodini as sister group. Arthrosolaenomeris Schubart, 1943 and Gangugia Schubart, 1947 were recovered as monophyletics. The genus Angelodesmus Schubart, 1962 was recovered as paraphyletic and redefined as monotypic. The tribe is now composed of the genera Arthrosolaenomeris (6 spp.), Gangugia (6 spp.), Angelodesmus (monotypic), Abiliodesmus n. gen. (4 spp.) and Folcloredesmus n. gen. (monotypic). In addition, we conduct a taxonomic revision of the tribe Arthrosolaenomeridini providing detailed redescriptions, drawings, and distribution maps for all species and genera, including the two new genera and the ten new species proposed: Arthrosolaenomeris saci n. sp.; A. curupira n. sp.; A. caipora n. sp.; A. iara n. sp.; Gangugia boitata n. sp.; G. cuca n. sp.; G. boto n. sp.; G. mula n. sp.; Abiliodesmus mapinguari n. sp. and Folcloredesmus thomasi n. sp. Furthermore, we include an identification key to all the genera and species. 

A giant within Carpocorini (Hemiptera: Heteroptera: Pentatomidae): Description of Adustonotus graziae sp. nov.

Zootaxa ◽  
2021 ◽  
Vol 4958 (1) ◽  
pp. 489-502
Author(s):  
FILIPE MICHELS BIANCHI
Keyword(s):  
Morphological Characters ◽  
The Other ◽  
Total Evidence ◽  
Valid Species ◽  
Phylogenetic Position ◽  
New Taxon ◽  
Large Size ◽  
Internal Genitalia ◽  
Evidence Analysis ◽  
Heteroptera Pentatomidae

The Carpocorini are distributed worldwide, and it is one of the most speciose tribes within the Pentatomidae with 127 genera and more than 500 valid species. Recently, Adustonotus Bianchi was described to contain eight species formerly placed within Euschistus Dallas. Among them, Adustonotus grandis (Rolston) and Adustonotus latus (Dallas) are remarkable for their large size. Herein, the phylogenetic position of a new taxon is inferred by a total evidence analysis based on 85 morphological characters and four molecular markers. Adustonotus graziae sp. nov. is described, and is recovered in a polytomic lineage, including A. grandis and A. latus. These species share a solid combination of features that enable them to be separated from the other Adustonotus species (e.g., large size, the humeral angles spatulate and exceptionally produced, and the capsula seminalis shortened). Illustrations of external and internal genitalia, and a distributional map are provided. 

A Taxonomic Revision of the Woody South African Genus Notobubon (Apiaceae: Apioideae)

2009 ◽  
Vol 34 (1) ◽  
pp. 220-242 ◽  
Author(s):  
Anthony R. Magee ◽  
Ben-Erik van Wyk ◽  
Patricia M. Tilney
Keyword(s):  
South African ◽  
Fruit Size ◽  
Cladistic Analysis ◽  
Taxonomic Revision ◽  
Morphological Characters ◽  
Cape Floristic Region ◽  
Evergreen Species ◽  
Inflorescence Structure ◽  
Peduncle Length ◽  
African Genus

A comprehensive taxonomic revision of the genus Notobubon (Apiaceae) is presented. Twelve woody evergreen species are recognised, all (with the exception of N. laevigatum) endemic to the Cape Floristic Region of South Africa. The taxonomy of these prominent, though poorly collected, species has until now been problematic. They are distinguished from one another by their habit (size and branching pattern), the overall shape, size, and colour of the ultimate leaflet segments, the inflorescence structure (peduncle length, number, and length of rays in the primary umbel), the fruit morphology (fruit size, presence or absence of wings), and the fruit anatomy (symmetry of the mericarps, presence or absence of additional rib vittae, size of commissural vittae). Species relationships are assessed in the form of a cladistic analysis of 26 morphological characters, resulting in a well-resolved phylogenetic hypothesis. A comprehensive key to the species, their correct nomenclature, and typification, together with descriptions and known geographical distribution for all the species are presented and illustrated.

scholarly journals Review of the Species of New World Erythroneurini (Hemiptera: Cicadellidae: Typhlocybinae)

2008 ◽  
Vol 38 (1-6) ◽  
pp. 129-175
Author(s):  
Christopher H. Dietrich ◽  
Dmitry A. Dmitriev
Keyword(s):  
Phylogenetic Analysis ◽  
New Species ◽  
South America ◽  
New World ◽  
Morphological Characters ◽  
Amazon Region ◽  
New Synonyms ◽  
Branch Support ◽  
North And South America ◽  
North And South

The 35 previously described species (including 4 junior synonyms) of the New World erythroneurine leafhopper genus Zyginama are reviewed and descriptions are provided for 43 new species, 2 from U.S.A., 26 from Mexico, 1 from Panama, and 14 from the Amazon region of South America. The following new synonyms are recognized: Z. aucta (McAtee) equals Erythroneura bilocularis Van Duzee, syn. n., and E. inclita Beamer, syn. n.; Z. nicholi (Beamer) equals E. canyonensis Beamer, syn. n., and E. ales Beamer, syn. n. Separate keys are provided for identification of males from North and South America, all known species are illustrated, and data on their distributions are summarized. Phylogenetic analysis of 40 morphological characters of the 71 species for which males are known yielded a reasonably well resolved estimate of relationships among the included species, but branch support was low overall.

The phylogenetic position of the tuatara, Sphenodon (Sphenodontida, Amniota), as indicated by cladistic analysis of the ultrastructure of spermatozoa

1992 ◽  
Vol 335 (1274) ◽  
pp. 207-219 ◽  
Keyword(s):  
Cladistic Analysis ◽  
Sister Group ◽  
Sister Taxon ◽  
Phylogenetic Position ◽  
Sister Group Relationship ◽  
Usual Concept ◽  
Cladistic Analyses ◽  
Relationship Of

Sphenodon has traditionally been regarded as a little changed survivor of the Permo-Triassic thecodont or eosuchian ‘stem reptiles’ but has alternatively been placed in the Lepidosauria as the plesiomorphic or even apomorphic sister-taxon of the squamates. A cladistic analysis of 16 characters from spermatozoal ultrastructure of Sphenodon and other amniotes unequivocally confirms its exceedingly primitive status. The analysis suggests that monotremes are the sister-group of birds; squamates form the sister-group of a bird + monotreme clade while the three sister-groups successively below the bird + monotreme + squa- mate assemblage are the caiman, the tuatara and the outgroup (turtles). The monotreme + bird couplet, supports the concept of the Haemothermia, but can only be regarded heuristically. The usual concept of mammals as a synapsid-derived outgroup of all other extant amniotes is not substantiated spermatologically. All cladistic analyses made, and a separate consideration of apomorphies, indicate that Sphenodon is spermatologically the most primitive amniote, excepting the Chelonia. It is advanced (apomorphic) for the amniotes in only two of the 16 spermatozoal characters considered. A close, sister-group relationship of Sphenodon with squamates is not endorsed.

A new species of Zoniopoda Stål (Orthoptera: Acridoidea: Romaleidae) from Argentina and its phylogenetic position within the genus

Zootaxa ◽  
2011 ◽  
Vol 2913 (1) ◽  
pp. 27 ◽  
Author(s):  
MARTINA E. POCCO ◽  
GONZALO D. RUBIO ◽  
M. MARTA CIGLIANO
Keyword(s):  
New Species ◽  
Cladistic Analysis ◽  
Type Specimen ◽  
Morphological Characters ◽  
Color Pattern ◽  
Species Group ◽  
The Body ◽  
Phylogenetic Position ◽  
Median Carina ◽  
A New Species

A new species of the romaleid grasshopper genus Zoniopoda Stål (Romaleidae: Romaleini) is described and illustrated from the Sierras Chicas of Córdoba Province, central Argentina. A cladistic analysis based on morphological characters indicates that the genus Zoniopoda constitutes a monophyletic group and that Zoniopoda serrana n. sp. must be assigned to Iheringi species group based on synapomorphies of the pronotum and body color. The new species is similar to Z. similis Bruner from Paraguay, Bolivia and Brazil, from which it can be distinguished by the color pattern of the body, shape of the pronotal dorso-median carina and characters of the male terminalia and epiphallus. This paper has been formatted with embedded links to images of the type specimen, maps based on georeferenced specimen data for the genus and an updated key to the species of Zoniopoda available on the Orthoptera Species File (OSF) online (http://orthoptera.speciesfile.org).

New lectotypifications and new synonyms in Festuca (Poaceae, Pooideae, Loliinae) from the Central Andes

Phytotaxa ◽  
2016 ◽  
Vol 247 (4) ◽  
pp. 247 ◽  
Author(s):  
JUAN C. OSPINA
Keyword(s):  
Morphological Characters ◽  
Central Andes ◽  
Andes Mountains ◽  
Valid Species ◽  
Second Step ◽  
New Synonyms ◽  
The Andes ◽  
New Synonym ◽  
Differential Characters ◽  
Anatomical Characters

New lectotypifications and new synonyms are provided as a result of a systematic study of the genus Festuca from the Central Andes. In the Andes mountains of Argentina, Bolivia, and Chile, several species of Festuca are dominant components of the grasslands and high Andean steppes. Festuca circinata and F. dissitiflora occur in this area and were described based on morphological characters. Some authors have considered F. circinata as a valid species while others have treated it as a synonym of F. dissitiflora, together with F. dissitiflora var. loricata , F. dissitiflora var. villipalea, and F. erecta var. aristulata. In this study, a review of the taxonomy and nomenclature of F. castilloniana, F. circinata, and F. potosiana was performed, and their morphological and anatomical characters were compared with F. dissitiflora. Based on morphoanatomical and epidermal characters, this study contributes to the delimitation of F. circinata and F. dissitiflora. Nomenclatural notes and new anatomical and epidermal descriptions are included for these species. Differential characters between F. circinata and F. dissitiflora are discussed and F. circinata is restored as a valid species for Argentina. Festuca castilloniana and F. potosiana are synonymised under F. dissitiflora. Festuca dissitiflora var. loricata, F. dissitiflora var. villipalea, and F. erecta var. aristulata are excluded from the synonymy of F. dissitiflora and it is suggested that these names should be subordinate under F. fiebrigii. Festuca stuckertii is proposed as a new synonym of F. uninodis. Lectotypes are designated for the names F. argentinensis, F. dissitiflora, F. nemoralis and F. parodii, and second-step lectotypes are designated for the names F. circinata and F. ampliflora.

Taxonomic revision and preliminary phylogenetic analysis of the subgenus Megalostomis (Scaphigenia) Lacordaire (Coleoptera: Chrysomelidae)

2007 ◽  
Vol 38 (3) ◽  
pp. 335-359 ◽  
Author(s):  
Sergio Roig-Juñent ◽  
Martha Domínguez ◽  
Federico Agrain
Keyword(s):  
Phylogenetic Analysis ◽  
Arid Regions ◽  
Cladistic Analysis ◽  
Taxonomic Revision ◽  
Morphological Characters ◽  
Distribution Data ◽  
Species Status ◽  
Systematic Revision ◽  
Male And Female ◽  
Preliminary Phylogenetic Analysis

AbstractThe subgenus M. (Scaphigenia) Lacordaire includes six species distributed in arid regions of South America. A systematic revision of this subgenus is presented, including re-descriptions and an update of distribution data of the six species. A preliminary cladistic analysis is performed to test the relationships among the species of the subgenus and if the morphological characters used are suitable. A key is presented to separate the seven subgenera of Megalostomis Chevrolat as well as a key for the species of the subgenus M. (Scaphigenia). Male aedeagus internal sac of the nine studied taxa are described and illustrated. This constitutes the first internal sac descriptions for members of this subtribe and could help clarify the homology of such structures within Cryptocephalinae. M. (S) consimilis Achard is reassigned to the species status on the base of genitalic characters. The cladistic analysis was done using forty-one characters from adult external morphology and male and female genitalia. Two species of two different subgenera of Megalostomis: M. (Megalostomis), and M. (Heterostomis) Lacordaire, and one representative of the genus Themesia Lacordaire were selected as out groups. Results were obtained by implicit enumeration using parsimony software. Three equally parsimonious trees were obtained of 45 steps, Ri=0.952, and Ci= 0.941. Support of each group was evaluated by Jacknifing, Bootstrap and Bremer values. Relationships between species are discussed.

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