Classification of basal Cucujoidea (Coleoptera:Polyphaga): cladistic analysis, keys and review of new families

2005 ◽  
Vol 19 (1) ◽  
pp. 17 ◽  
Author(s):  
R. A. B. Leschen ◽  
J. F. Lawrence ◽  
S. A. Ślipiński
Keyword(s):  
New Taxa ◽  
Phylogenetic Relationships ◽  
Type Species ◽  
Cladistic Analysis ◽  
Data Matrix ◽  
Family Status ◽  
The Family

Phylogenetic relationships among the basal Cucujoidea were reconstructed by a cladistic analysis of a data matrix consisting of 37 exemplar taxa and 99 adult and larval characters. Eight most parsimonious cladograms provided evidence for the polyphyly of Phloeostichidae, the paraphyly of Cucujoidea (with respect to the placement of Trogossitidae), and the monophyly of Protocucujidae + Sphindidae, Biphyllidae + Erotylidae, Cryptophagidae, Cucujidae + Silvanidae, Propalticidae + Laemophloeidae, and the Nitidulidae groups (Nitidulidae, Smicripidae, and Brachypteridae). The following families are elevated from subfamily to family status: Agapythidae (one genus), Phloeostichidae (four genera; the subfamilies Phloeostichinae and Hymaeinae are supressed), Priasilphidae (three genera), Tasmosalpingidae (one genus), and Myraboliidae (one genus). These families are described in detail and adult and larval keys to all families of Cucujoidea are provided. The genus Bunyastichus, gen. nov. (type species: B. monteithi, sp. nov.) is described in the family Phloeostichidae and the family Priasilphidae is revised with the following new taxa: Chileosilpha, gen. nov. (type species: C. elguetai, sp. nov.), Priasilpha (P. angulata, sp. nov., P. aucklandica, sp. nov., P. bufonia, sp. nov., P. carinata, sp. nov., P. earlyi, sp. nov., and P. embersoni, sp. nov.), Priastichus (P. crowsoni, sp. nov. and P. megathorax, sp. nov.).

scholarly journals Phylogenetic relationships within the Pylochelidae (Decapoda: Anomura: Paguroidea): A cladistic analysis based on morphological characters

Zootaxa ◽  
2009 ◽  
Vol 2022 (1) ◽  
pp. 1-14 ◽  
Author(s):  
RAFAEL LEMAITRE ◽  
PATSY A. MCLAUGHLIN ◽  
ULF SORHANNUS
Keyword(s):  
Hermit Crab ◽  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
Data Matrix ◽  
Parsimony Analysis ◽  
Consensus Tree ◽  
Strict Consensus Tree ◽  
The Family

Phylogenetic relationships within the “symmetrical” hermit crab family Pylochelidae were analyzed for 41 of the 45 species and subspecies currently considered valid. In the analyses, 78 morphological characters comprised the data matrix and the outgroup consisted of Thalassina anomala, a member of the Thalassinidae, and Munida quadrispina, a member of the Galatheidae. A poorly resolved strict consensus tree was obtained from a heuristic parsimony analysis of unweighted and unordered characters, which showed the family Pylochelidae and the subfamilies Pylochelinae and Pomatochelinae to be monophyletic taxa – the latter two groups had the highest Bremer support values. Additionally, while the subgenus Pylocheles (Pylocheles) was strongly supported, the subgenera Xylocheles, and Bathycheles were not. More fully resolved trees were obtained when using implied weighting, which recognized the monotypic subfamilies Parapylochelinae, Cancellochelinae and Mixtopagurinae. The subfamily Trizochelinae was found to have four distinct clades and several ambiguously placed taxa.

Phylogenetic relationships and a new classification of the family Cyclopteridae (Perciformes: Cottoidei)

Zootaxa ◽  
2017 ◽  
Vol 4221 (1) ◽  
pp. 1 ◽  
Author(s):  
KANAMI OKU ◽  
HISASHI IMAMURA ◽  
MAMORU YABE
Keyword(s):  
Phylogenetic Analysis ◽  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
New Classification ◽  
The Family ◽  
Dorsal Process ◽  
Transformation Series

 Phylogenetic relationships of the family Cyclopteridae were reconstructed based on osteological and external characters.  The monophyly of the family was strongly supported by 47 commonly recognized synapomorphies, including six autapomorphies (plus one additional autapomorphy, presence of a dorsal process on the pelvis, recognized after the phylogenetic analysis) among the suborder Cottoidei.  As a result of the cladistic analysis, a single most parsimonious phylogeny was obtained, based on characters in 32 transformation series.  A new classification of Cyclopteridae based on reconstructed relationships, including three subfamilies [Liparopsinae, Cyclopterinae and Eumicrotreminae (newly established)] and four genera (Aptocyclus, Cyclopsis, Cyclopterus and Eumicrotremus), is proposed.    

scholarly journals A current generic classification of sap beetles (Coleoptera, Nitidulidae)

2008 ◽  
Vol 17 (1) ◽  
pp. 107-122
Author(s):  
A.G. Kirejtshuk
Keyword(s):  
New Taxa ◽  
Type Species ◽  
New Genera ◽  
Type Series ◽  
Sap Beetles ◽  
The Family ◽  
New Interpretation ◽  
New Synonymy ◽  
A Current

The list of generic and subgeneric taxa arranged into subfamilies and tribes proposed for sap beetles is given, which includes also new genera [Parapocadius gen. nov. (Nitidulinae: Nitidulini) and Interfaxia gen. nov. (Nitidulinae: Cyllodini)] and subgenera [Semocarpolus subgen. nov. Gaplocarpolus subgen. nov. Askocarpolus subgen. nov. (Carpophilinae: Carpophilus Stephens, 1829)]. The Meoncerus Sharp, 1891; Apsectochilus Reitter, 1874 and Lordyrops Reitter, 1875 are considered as quite distinct each from other above mentioned as from all other generic taxa. In the list there are given the complete synonymy, including new synonymy of generic and subgeneric names [Crepuraea Kirejtshuk, 1990 and Nyujwa Perkovsky, 1990 syn. nov.; Haptoncus Murray, 1864 and Haptoncurina Jelínek, 1977, syn. nov.; Ecnomaeus Erichson, 1843 and Somaphorus Murray, 1864, syn. nov.; Ecnomorphus Motschulsky, 1858; Tribrachys LeConte, 1861 syn. nov.; Stauroglossicus Murray, 1864, syn. nov. and Microxanthus Murray, 1864, syn. nov.; Pria Stephens, 1829 and Allopria Kirejtshuk, 1980, syn. nov.; Megauchenia Macleay, 1825 and Orvoenia Dajoz, 1980, syn. nov.; Tetrisus Murray, 1864 and Pseudoischaena Grouvelle, 1897, syn. nov.; Neopocadius Grouvelle, 1906 and Pseudostelidota Grouvelle, 1906, syn. nov.; Cychramus Kugelann, 1794 and Aethinopsis Grouvelle, 1908, syn. nov.; Mystrops Erichson, 1843, and Cryptoraea Retter, 1873, syn. nov.; Cyllodes Erichson, 1843 and Mecyllodes Sharp, 1891, syn. nov.; Grammorus Murray, 1868 and Colopteroides Watrous, 1982, syn. nov.; Cryptarcha Shuckard, 1839 and Priatelus Broun, 1881, syn. nov.]. For some taxa the rank is changed, namely, Lordyra Gemminger & Harold, 1868, stat. nov. is regarded as a subgenus of Lasiodactylus Perty, 1830-1834; Brounthina Kirejtshuk, 1997, stat. nov. as a subgenus Neopocadius Grouvelle, 1906 and Teichostethus Sharp, 1891, stat. nov. as a subgenus of Hebascus Erichson, 1843, while the taxa Coxollodes Kirejtshuk, 1987 (stat. nov.) and Onicotis Murray, 1864 (stat. nov.) are regarded as separate genera. The new taxa and new taxonomical proposals are supplied with corresponding data in the notes below the list. In these notes there are also proposed the new synonymy for the following species names: Carpophilus (Ecnomorphus) acutangulus Reitter, 1884 and C. (E.) cingulatus Reitter, 1884, syn. nov.; C. (E.) bakeweli Murray, 1864; C. (E.) planatus Murray, 1864, syn. nov. and C. (E.) aterrimus Macleay, 1864, syn. nov.; C. (E.) debilis Grouvelle, 1897 and C. (E.) opaculus Grouvelle, 1897, syn. nov.; C. (E.) luridipennis Macleay, 1873 and C. (E.) loriai Grouvelle, 1906, syn. nov.; C. (E.) murrayi Grouvelle, 1892 and C. (E.) hebetatus Grouvelle, 1908, syn. nov.; C. (E.) plagiatipennis (Motschulsky, 1858) and C. (E.) nigricans Grouvelle, 1897, syn. nov.; C. (E.) terminalis Murray, 1864 and C. (E.) gentilis Murray, 1864, syn. nov.; Lasiodactylus brunneus Perty, 1830; L. centralis Cline et Carlton, 2004, syn. nov.; L. falini Cline et Carlton, 2004, syn. nov. and L. kelleri Cline et Carlton, 2004, syn. nov.); Pallodes opacus Grouvelle, 1906 and P. loriai Grouvelle, 1906, syn. nov.; Cyllodes fauveli Grouvelle, 1903 and Pallodes vagepunctus Grouvelle, 1903, syn. nov.; Pallodes jucundus Reitter, 1873 and Mecyllodes nigropictus Sharp, 1891, syn. nov.; Pallodes birmanicus Grouvelle, 1892 and P. kalingus Kirejtshuk, 1987, syn. nov.; P. gestroi Grouvelle, 1906 and P. misellus Grouvelle, 1906, syn. nov.; P. ruficollis Reitter, 1873 and P. cyanescens Grouvelle, 1898, syn. nov.; Grammophorus caelatus Gerstäcker, 1864 and Colopterus striaticollis Murray, 1864, syn. nov. In connection with a preliminary revision of many type series of the family and the mentioned taxonomical changes for some species names are established new taxonomical interpretation, namely: Pleoronia nitida (Grouvelle, 1898), comb. nov. (Axyra : Axyrodes); Parapocadius immerizi (Grouvelle, 1899), comb. nov. (Pallodes); Camptodes ruficornis (Grouvelle, 1898), comb. nov. (Pallodes); Neopallodes aestimabilis (Grouvelle, 1906), comb. nov. (Pallodes); N. alluaudi (Grouvelle, 1899), comb. nov. (Pallodes); N. aterrimus (Grouvelle, 1906), comb. nov. (Pallodes); N. dorsalis (Grouvelle, 1896), comb. nov. (Pallodes); N. fairmairei (Grouvelle, 1906), comb. nov. (Pallodes); N. incertus (Grouvelle, 1906), comb. nov. (Pallodes); N. klugi (Grouvelle, 1896), comb. nov. (Pallodes); N. limbicollis (Reitter, 1880), comb. nov. (Pallodes); N. militaris (Grouvelle, 1906), comb. nov. (Pallodes); N. niger (Grouvelle, 1906), comb. nov. (Pallodes); N. nigrocyaneus Grouvelle, 1906), comb. nov. (Pallodes); N. nitidus (Grouvelle, 1906), comb. nov. (Pallodes); N. orthogonus (Grouvelle, 1906), comb. nov. (Pallodes); N. perrieri (Grouvelle, 1906), comb. nov. (Pallodes); N. scutellaris (Grouvelle, 1906), comb. nov. (Pallodes); N. sicardi (Grouvelle, 1906), comb. nov. (Pallodes); N. sikordi (Grouvelle, 1896), comb. nov. (Pallodes); N. variabilis (Grouvelle, 1896) , comb. nov. (Pallodes); Coxollodes cyrtusoides (Reitter, 1884), comb. nov. (Pallodes); C. amamiensis (Hisamatsu, 1956), comb. nov. (Pallodes); Coxollodes opacus (Grouvelle, 1906), comb. nov. (Pallodes); C. parvulus (Grouvelle, 1908), comb. nov. (Pallodes); C. reitteri Kirejtshuk, 1987, comb. nov. (Pallodes); Pallodes fauveli (Grouvelle, 1903), comb. nov. (Cyllodes); Cyllodes jucundus (Reitter, 1873), comb. nov. (Pallodes); Interfaxia fasciata (Sharp, 1891), comb. nov.; Onicotis auritus Murray, 1864 comb. nov.; Platyarcha biguttata (Motschulsky, 1858), comb. nov. (Carphophilus: Ecnomorphus); Cryptarcha optanda (Broun, 1881), comb. nov. (Priateles). Besides, because of the new interpretation for Pallodes laetus Grouvelle, 1898, which should be transfered to the genus Camptodes, C. grouvellei nom. nov. (non Camptodes laetus Kirsch, 1873) is proposed. For the generic names Perilopa Erichson, 1843 and Meoncerus Sharp, 1891 the type species are designated as well as for Pallodes laetus Grouvelle, 1898 (Camptodes grouvellei nom. nov.) the lectotype designation is made.

Phylogeny and revised classification of the tribe Elachipterini (Diptera: Chloropidae)

Zootaxa ◽  
2018 ◽  
Vol 4471 (1) ◽  
pp. 1
Author(s):  
JULIA J. MLYNAREK ◽  
TERRY A. WHEELER
Keyword(s):  
Phylogenetic Relationships ◽  
Type Species ◽  
Cladistic Analysis ◽  
Morphological Characters ◽  
New Combinations ◽  
Old World ◽  
Neotropical Species ◽  
Valid Genus ◽  
Outgroup Species

The phylogenetic relationships of the chloropid tribe Elachipterini were analysed. Sixty-eight exemplar species and seven outgroup species were included in a cladistic analysis based on 76 morphological characters of adult specimens in order to test existing, non-phylogenetic, classifications of the tribe. Nine genera are recognized in the Elachipterini: Allomedeia Mlynarek & Wheeler, Alombus Becker, Anatrichus Loew, Ceratobarys Coquillett, Disciphus Becker, Elachiptera Macquart, Goniaspis Duda, Melanochaeta Bezzi and Sepsidoscinis Hendel. Myrmecosepsis Kertesz is synonymised with Anatrichus, and Togeciphus Nishijima and Cyrtomomyia Becker are synonymised with Elachiptera. Ceratobarys is removed from synonymy with Elachiptera and all Neotropical species and two Nearctic species previously assigned to Elachiptera are transferred to Ceratobarys. Melanochaeta is a valid genus; the type species Melanochaeta capreolus clusters with other species of Melanochaeta and not Oscinella. New combinations include Anatrichus hystrix (Kertesz, 1914) (Myrmecosepsis); Anatrichus taprobane (Andersson, 1977) (Myrmecosepsis), Ceratobarys attenuata (Adams, 1908) (Elachiptera); Ceratobarys cultrata (Wheeler & Forrest, 2002) (Elachiptera); Ceratobarys flavida (Williston, 1896) (Elachiptera); Ceratobarys melinifrons (Mlynarek & Wheeler, 2008) (Elachiptera); Ceratobarys fucosa (Mlynarek & Wheeler, 2008) (Elachiptera); Ceratobarys queposana (Mlynarek & Wheeler, 2008) (Elachiptera); Ceratobarys rubida (Becker, 1912) (Elachiptera); Ceratobarys sacculicornis (Enderlein, 1911) (Elachiptera); Ceratobarys willistoni (Sabrosky, 1948)  (Elachiptera), Elachiptera ensifer (Sabrosky, 1951) (Cyrtomomyia); Elachiptera ericius (Kanmiya, 1983) (Togeciphus); Elachiptera katoi (Nishijima, 1955) (Togeciphus); Elachiptera maculinervis (Becker, 1910) (Cyrtomomyia); Elachiptera punctulata (Becker, 1912) (Cyrtomomyia); Elachiptera subelongata (Kanmiya, 1983) (Disciphus); Elachiptera truncatus (Liu & Yang, 2012) (Togeciphus); Elachiptera tuberculata (Adams, 1905) (Cyrtomomyia) and all the species that were placed in Lasiochaeta are returned to Melanochaeta. A key to genera of the tribe Elachipterini is provided and diagnoses are provided for all genera. The tribe is divided into two geographically distinct clades: the Anatrichus clade includes the Old World tropical genera Allomedeia, Alombus, Anatrichus, Disciphus and Sepsidoscinis; the Elachiptera clade includes the primarily Neotropical genera Goniaspis and Ceratobarys and the widespread, but primarily Holarctic, genera Elachiptera and Melanochaeta. 

Three new species, two new genera, and new family Biphragmosagittidae (Сhaetognatha) from Southwest Pacific Ocean

2011 ◽  
Vol 20 (1) ◽  
pp. 161-173
Author(s):  
A.P. Kassatkina
Keyword(s):  
New Species ◽  
Type Species ◽  
Morphological Characters ◽  
The Body ◽  
New Order ◽  
New Genera ◽  
New Family ◽  
The Family ◽  
Three New Species

Resuming published and own data, a revision of classification of Chaetognatha is presented. The family Sagittidae Claus & Grobben, 1905 is given a rank of subclass, Sagittiones, characterised, in particular, by the presence of two pairs of sac-like gelatinous structures or two pairs of fins. Besides the order Aphragmophora Tokioka, 1965, it contains the new order Biphragmosagittiformes ord. nov., which is a unique group of Chaetognatha with an unusual combination of morphological characters: the transverse muscles present in both the trunk and the tail sections of the body; the seminal vesicles simple, without internal complex compartments; the presence of two pairs of lateral fins. The only family assigned to the new order, Biphragmosagittidae fam. nov., contains two genera. Diagnoses of the two new genera, Biphragmosagitta gen. nov. (type species B. tarasovi sp. nov. and B. angusticephala sp. nov.) and Biphragmofastigata gen. nov. (type species B. fastigata sp. nov.), detailed descriptions and pictures of the three new species are presented.

A comparison of methodological approaches to the subfamilial classification of the Naididae (Oligochaeta)

1987 ◽  
Vol 65 (3) ◽  
pp. 691-707 ◽  
Author(s):  
A. F. L. Nemec ◽  
R. O. Brinkhurst
Keyword(s):  
Data Matrix ◽  
Jaccard Coefficient ◽  
Data Set ◽  
Parsimony Method ◽  
Fit Statistics ◽  
Reconstruction Methods ◽  
Linkage Methods ◽  
The Family ◽  
First Time

A data matrix of 23 generic or subgeneric taxa versus 24 characters and a shorter matrix of 15 characters were analyzed by means of ordination, cluster analyses, parsimony, and compatibility methods (the last two of which are phylogenetic tree reconstruction methods) and the results were compared inter alia and with traditional methods. Various measures of fit for evaluating the parsimony methods were employed. There were few compatible characters in the data set, and much homoplasy, but most analyses separated a group based on Stylaria from the rest of the family, which could then be separated into four groups, recognized here for the first time as tribes (Naidini, Derini, Pristinini, and Chaetogastrini). There was less consistency of results within these groups. Modern methods produced results that do not conflict with traditional groupings. The Jaccard coefficient minimizes the significance of symplesiomorphy and complete linkage avoids chaining effects and corresponds to actual similarities, unlike single or average linkage methods, respectively. Ordination complements cluster analysis. The Wagner parsimony method was superior to the less flexible Camin–Sokal approach and produced better measure of fit statistics. All of the aforementioned methods contain areas susceptible to subjective decisions but, nevertheless, they lead to a complete disclosure of both the methods used and the assumptions made, and facilitate objective hypothesis testing rather than the presentation of conflicting phylogenies based on the different, undisclosed premises of manual approaches.

Philippinozercon, a new genus of Heterozerconidae (Parasitiformes: Mesostigmata), with description of all active instars

Zootaxa ◽  
2018 ◽  
Vol 4540 (1) ◽  
pp. 7
Author(s):  
BEVERLY S. GERDEMAN ◽  
RUFINO C. GARCIA ◽  
ANDREW HERCZAK ◽  
HANS KLOMPEN
Keyword(s):  
Type Species ◽  
New Genus ◽  
The Philippines ◽  
Junior Synonym ◽  
Temperate Species ◽  
Oriental Region ◽  
The Family

The generic classification of millipede associated Heterozerconidae in the Oriental region is revised. The genus Allozercon Vitzthum is re-diagnosed and Asioheterozercon Fain is designated as an subjective junior synonym of Allozercon. Philippinozercon gen. nov., with the type species P. makilingensis sp. nov., is described for all instars. This genus may be endemic for the Philippines, but is quite widespread in that country. All immature instars are described, making this the second species of Heterozerconidae known for all instars. The morphology of the immatures is compared with that of immatures of the temperate species Narceoheterozercon ohioensis and unnamed species from Brazil and Thailand. All immatures were collected from millipede frass and litter, never from millipedes. Adults are associated with millipedes in the family Trigoniulidae (Spirobolida). 

Phylogeny and systematics of the Trapeziidae Miers, 1886 (Crustacea: Brachyura), with the description of a new family

Zootaxa ◽  
2004 ◽  
Vol 643 (1) ◽  
pp. 1 ◽  
Author(s):  
PETER CASTRO ◽  
PETER K.L. NG ◽  
SHANE T. AHYONG
Keyword(s):  
Cladistic Analysis ◽  
Type Material ◽  
Family Status ◽  
Reef Corals ◽  
New Family ◽  
The Family ◽  
Repeated Use ◽  
Name Changes ◽  
Fossil Genus

A revision of the family Trapeziidae Miers, 1886, has shown that it consists of three clades, one of which is elevated to family status, Tetraliidae fam. nov., for the genera Tetralia Dana, 1851, and Tetraloides Galil, 1986. The genera Trapezia Latreille, 1828, Calocarcinus Calman, 1909, Hexagonalia Galil, 1986, Philippicarcinus Garth & Kim, 1983, Quadrella Dana, 1851, and Sphenomerides Rathbun, 1897, remain in the Trapeziidae; Domecia Eydoux & Souleyet, 1842, Jonesius Sankarankutty, 1962, Maldivia Borradaile, 1902, Palmyria Galil & Takeda, 1986, and the fossil genus Eomaldivia M ller & Collins, 1991, in Domeciidae Ortmann, 1893. Cladistic analysis shows that Trapeziidae sensu Miers, 1886, consists of three clades that show convergence as a result of similar habits as symbionts of reef corals and other cnidarians. A list of all recognised genera and species in the three families and their primary synonyms is provided. Keys are also included for four families of Brachyura symbiotic with reef corals, and for the genera and species of Domeciidae, Tetraliidae, and Trapeziidae. Some rare colour figures are reproduced. Three name changes have resulted within the Tetraliidae: Cancer glaberrimus Herbst, 1790, for Tetralia fulva Ser ne, 1984, and Cancer mutus Linnaeus, 1758, for Tetralia armata Dana, 1852, and Tetralia vanninii Galil & Clark, 1988. Nomenclatural problems associated with the repeated use of "forma typica" for various species of Trapezia and Tetralia are resolved. To stabilise the nomenclature of a number of well-known species, neotypes are designated for 13 species of Trapeziidae for which type material is not extant: Trapezia cymodoce (Herbst, 1801), and its three synonyms (Trapezia dentifrons Latreille, 1828, Trapezia dentata var. subintegra Dana, 1852, Trapezia cymodoce var. ornatus Chen, 1933); Trapezia bidentata (Forsk l, 1775), and one of its synonyms (Trapezia ferruginea Latreille, 1828); Trapezia digitalis Latreille, 1828, and one of its synonyms (Trapezia nigrofusca Stimpson, 1858); Trapezia septata Dana, 1852, and one of its synonyms (Trapezia reticulata Stimpson, 1858); Trapezia areolata Dana, 1852; Trapezia bella Dana, 1852; and Trapezia speciosa Dana, 1852. Neotypes are also designated for seven species of Tetraliidae: Tetralia glaberrima (Herbst, 1790), and three synonyms (Trapezia integra Latreille, 1828, Trapezia serratifrons Jacquinot, 1846, Tetralia laevissima Stimpson, 1858); Tetralia muta (Linnaeus, 1758), and one of its synonyms (Tetralia armata Dana, 1852); and Tetraloides nigrifrons (Dana, 1852).

Phylogenetic relationships of the African genera Alestes and Brycinus (Teleostei, Characiformes, Alestidae)

2002 ◽  
Vol 80 (11) ◽  
pp. 1887-1899 ◽  
Author(s):  
Alison M Murray ◽  
Kathlyn M Stewart
Keyword(s):  
Phylogenetic Relationships ◽  
Cladistic Analysis ◽  
Sister Group ◽  
Monophyletic Group ◽  
The Family ◽  
The Subject

The family Alestidae (also referred to as the African Characidae) comprises the African dwarf forms ("Petersiini") and the genera Alestes, Brycinus, Bryconaethiops, and Hydrocynus. Although several authors have presented characters to support the monophyly of the family, a cladistic analysis of the group has not been published. Furthermore, the interrelationships of the constituent groups are the subject of some controversy. A cladistic analysis of the Alestidae is presented, including characters to support the monophyly of the family. The results of this study indicate that several species should be removed from the genus Brycinus, that Hydrocynus is the sister group of Alestes s.str. (containing only five species), and that the dwarf alestids ("Petersiini") do not form a monophyletic group.

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