Deep-Sea Trace Fossils In The West Crocker Formation, Sabah (Malaysia), And Their Palaeoenvironmental Significance

In the “flysch” series of the West Crocker Formation (Eocene–Oligocene), Kota Kinabalu, Sabah, trace fossils are fairly common although not ubiquitous. The trace fossils commonly occur as hypichnial semi- or full-reliefs on the sole of thin turbiditic sandstone beds (mainly Bouma Tc division) in the thinly bedded heterolithic sandstone-mudstone facies interpreted as submarine fan lobe deposits. Their presence in mainly the thinly bedded facies of the fan system suggests preferential production and preservation in the fine-grained “distal” parts of the Crocker submarine fan system. Trace fossil assemblages characteristic of the Nereites ichnofacies indicate sedimentary environments mainly in bathyal to abyssal water depths (>2000 m). This ichnofacies is dominated by horizontal grazing, farming and feeding traces, ranging from solitary to branching tubular burrows (Ophiomorpha, Palaeophycus and Planolites) to meandering trails and tunnels (Nereites, Cosmorhaphe, Helminthopsis), as well as the spiriform burrows Spirophycus. Graphoglyptids are the most diagnostic of the Nereites ichnofacies, produced by sediment grazers and farmers (agrichnia) and often displaying intricate networks of mainly horizontal tunnels preserved as hypichnial semi-reliefs. They include the delicate spiral traces of Spirorhaphe, as well as the enigmatic hexagonal network burrow Paleodictyon. Other ichnogenera include Planolites, Thalassinoides and Ophiomorpha which are facies-crossing and not environment specific. Detailed observations of the trace fossil assemblages and the degree of bioturbation enabled different sub-ichnofacies of the Nereites ichnofacies to be distinguished. Ophiomorpha is more common in sandy “proximal” facies and tend to penetrate deeply into pre-existing turbidite beds, its presence suggests a well-oxygenated newly deposited turbidite substrate, probably in the axial region of the fan lobes. Hence, channel axis and proximal fan deposits tend to be dominated by the Ophiomorpha rudis sub-ichnofacies. The Paleodictyon sub-ichnofacies is more typical of the lower energy lobe/fan fringe subenvironments. Proximal but off-axis areas are characterized by a mixture of the Ophiomorpha rudis and Paleodictyon sub-ichnofacies.

The Late Miocene Rifian corridor as a natural laboratory to explore a case of ichnofacies distribution in ancient gateways

Oceanic gateways have modulated ocean circulation and have influenced climatic variations throughout the Earth´s history. During the late Miocene (7.8–7.35 Ma), the Atlantic Ocean and the Mediterranean Sea were connected through the Rifian Corridor (Morocco). This gateway is one of the few examples of deep ancient seaways with a semi-continuous sedimentary record. Deposits comprise turbidites intercalated between deep-sea mudstone (i.e., hemipelagites and drift deposits), channelized sandstone contourite facies, and shallow marine sandstone. Herein an ichnological analysis was conducted in these upper Miocene sediments to improve characterisation of palaeoenvironmental conditions. In addition, ichnofacies were analysed to elucidate how bottom currents control ichnofacies distribution and can modify their attributes. Turbidite deposits are typified by vertical trace fossils (i.e., Ophiomorpha), conforming the Ophiomorpha rudis ichnosubfacies. Contouritic sandstone exhibits high density and low diversity trace-fossil assemblage, with predominant Macaronichnus and Scolicia, resembling a proximal expression of the Cruziana ichnofacies. Shallow marine environments are dominated by vertical trace fossils (e.g., Conichnus, Ophiomorpha, Skolithos), allowing an assignation to the Skolithos ichnofacies. This study reveals for the first time a variability in ichnofacies attributes and distribution at the Rifian Corridor, associated with turbidites, contourite and shallow marine sediments. Hydrodynamic energy reveals as the major factor controlling trace maker communities in the studied seaway. Highly energetic conditions typical of shallower settings are present in deeper-water environments (i.e., slope), contributing to ichnodiversity impoverishment in ichnofacies.

Exploring Ichnofacies Distribution in Ancient Seaways: The Late Miocene Rifian Corridor as a Natural Laboratory

Oceanic gateways have modulated ocean circulation and have influenced climatic variations throughout the Earth´s history. During the Late Miocene (7.8 - 7.35 Ma), the Atlantic Ocean and the Mediterranean Sea were connected through the Rifian Corridor (Morocco). This gateway is one of the few examples of deep ancient seaways with a semi-continuous sedimentary record. Deposits comprise turbidites intercalated between deep-sea fine grain sediments (i.e., hemipelagites and drift deposits), channelized sandstone contourite facies, and shallow marine sandstones. Herein an ichnological analysis was conducted in these upper Miocene sediments to improve characterisation of palaeoenvironmental conditions. In addition, ichnofacies were analysed to elucidate how bottom currents control ichnofacies distribution and can modified their attributes. Turbidite deposits are typified by vertical trace fossils (i.e., Ophiomorpha), conforming the Ophiomorpha rudis ichnosubfacies. Contouritic sandstones exhibit high density and low diverse trace fossil assemblage, with predominant Macaronichnus and Scolicia, resembling a proximal expression of Cruziana ichnofacies. Shallow marine environments are dominated by vertical trace fossils (e.g., Conichnus, Ophiomorpha and Skolithos), allowing an assignation to Skolithos ichnofacies. This study reveals energy to be a major controlling factor determining ichnofacies attributes and distribution in ancient deep seaways. Within these seaways, highly energetic conditions typical of shallower settings are present in deeper environments (i.e., slope), contributing to ichnodiversity impoverishment in ichnofacies.

Ichnology of Upper Cretaceous deep-sea thick-bedded flysch sandstones: Lower Istebna Beds, Silesian Unit (Outer Carpathians, southern Poland)

Ichnology of Upper Cretaceous deep-sea thick-bedded flysch sandstones: Lower Istebna Beds, Silesian Unit (Outer Carpathians, southern Poland) The Ophiomorpha rudis ichnosubfacies of the Nereites ichnofacies was recognized in thick- and very thick-bedded sandstones of the Lower Istebna Beds (Campanian-Maastrichtian), which were deposited mainly in deep-sea clastic ramps and aprons. It contains mainly Ophiomorpha rudis (produced by deeply burrowing decapod crustaceans) and rarely Zoophycos isp. and Chondrites isp. The impoverished Paleodictyon ichnosubfacies of the Nereites ichnofacies is present in the medium- and thin-bedded packages of flysch sandwiched between the thick- and very thick-bedded sandstones. They contain Chondrites isp., Phycosiphon incertum, Planolites isp., Arthrophycus strictus, Thalassinoides isp., Ophiomorpha annulata, O. rudis, Scolicia strozzii and Helminthorhaphe flexuosa. The relatively low diversity of this assemblage is influenced by limited areas covered by muddy substrate, which favours deep-sea tracemakers, and partly by a lowered oxygenation in the sediment.