Carboniferous fusuline Foraminifera: taxonomy, regional biostratigraphy, and palaeobiogeographic faunal development

This paper proposes a synthesis of the taxonomy, phylogeny, palaeogeographic distribution, regional biostratigraphy, and palaeobiogeographic faunal development of Carboniferous fusuline foraminifers. They appeared in the latest Tournaisian and comprised a small-sized, morphologically conservative taxonomic group during the Mississippian. Fusulines became larger and prevailed in Pennsylvanian foraminiferal assemblages. Carboniferous fusulines consist of Ozawainellidae, Staffellidae, Schubertellidae, Fusulinidae, and Schwagerinidae, in which 95 genera are considered as valid taxonomically. Upsizing their shells throughout the Pennsylvanian is likely related to symbiosis with photosynthetic microorganisms, which was accelerated by the acquisition of a keriothecal wall in Late Pennsylvanian schwagerinids. Regional fusuline succession data from 40 provinces provide a refined biostratigraphy, enabling zonation and correlation with substage- or higher-resolution precision in the Pennsylvanian. Their spatio-temporal faunal characteristics show that fusulines had a cosmopolitan palaeobiogeographic signature in Mississippian time, suggesting unrestricted faunal exchange through the palaeoequatorial Rheic Ocean. After the formation of Pangea, Pennsylvanian fusulines started to show provincialism, and their distributions defined the Ural-Arctic Region in the Boreal Realm, Palaeotethys, Panthalassa, and North American Craton regions in the Palaeoequatorial Realm, and Western Gondwana and Eastern Peri-Gondwana regions in the Gondwana Realm. The Western Palaeotethys and East European Platform Subregions maintained higher generic diversity throughout the Pennsylvanian.

Coevolution of global brachiopod palaeobiogeography and tectonopalaeogeography during the Carboniferous

The global brachiopod palaeobiogeography of the Mississippian is divided into three realms, six regions, and eight provinces, while that of the Pennsylvanian is divided into three realms, six regions, and nine provinces. On this basis, we examined coevolutionary relationships between brachiopod palaeobiogeography and tectonopalaeogeography using a comparative approach spanning the Carboniferous. The appearance of the Boreal Realm in the Mississippian was closely related to movements of the northern plates into middle–high latitudes. From the Mississippian to the Pennsylvanian, the palaeobiogeography of Australia transitioned from the Tethys Realm to the Gondwana Realm, which is related to the southward movement of eastern Gondwana from middle to high southern latitudes. The transition of the Yukon–Pechora area from the Tethys Realm to the Boreal Realm was associated with the northward movement of Laurussia, whose northern margin entered middle–high northern latitudes then. The formation of the six palaeobiogeographic regions of Mississippian and Pennsylvanian brachiopods was directly related to “continental barriers”, which resulted in the geographical isolation of each region. The barriers resulted from the configurations of Siberia, Gondwana, and Laurussia, which supported the Boreal, Tethys, and Gondwana realms, respectively. During the late Late Devonian–Early Mississippian, the Rheic seaway closed and North America (from Laurussia) joined with South America and Africa (from Gondwana), such that the function of “continental barriers” was strengthened and the differentiation of eastern and western regions of the Tethys Realm became more distinct. In the Barents Ocean tectonic domain during the Pennsylvanian, the brachiopods on the northern margin of the Barents Ocean formed the Verkhoyansk–Taymyr Province, while those on the southern margin formed the Yukon–Pechora Province. The Mongolia–Okhotsk Province was formed by brachiopods of the Mongolia–Okhotsk Ocean tectonic domain. The Northern Margin of the Palaeo-Tethys Ocean Province and the Southern Margin of the Palaeo-Tethys Ocean Province were formed, respectively, by brachiopods on the northern and southern margins of the Palaeo-Tethys Ocean tectonic domain. South China and Southeast Asia were dissociated from the major continental blocks mentioned above, and formed the South China Province.

Climate changes across Oceanic Anoxic Event 1a: new data from the Boreal realm

<div><span>Our understanding of the climatic evolution during the early Cretaceous in general, and the Oceanic Anoxic Event 1a (OAE-1a) in peculiar, is widely derived from the investigation of Tethyan localities, leaving large uncertainties about their significance on a global scale. In this study, we have performed high-resolution clay-mineral assemblage analyses in the upper Hauterivian to lower Aptian of the </span><span>North Jens-1 core, located in the Danish central graben that was part of the Boreal realm. High amount of detrital Kaolinite is observed throughout the core, indicating the presence of a local, kaolinite-rich source. A long-term decline in Kaolinite content is occurring from the lower Hauterivian to the lowermost, pre-OAE-1a Aptian, followed by a sharp rise within the OAE-1a. This trend is similar to the one observed in the Tethyan realm, indicating a supra-regional climatic evolution from humid conditions in the late Hauterivian toward drier conditions in the latest Barremian – earliest Aptian, followed by renewed humid conditions during the unfolding of OAE-1a. Precise timing of climate change during the OAE-1a differs however in between the Tethyan and Boreal realm. Shift toward humid conditions coincides hence with the onset of OAE-1a (segment C3) in the Tethyan realm, followed by a return to drier conditions in the second half of OAE-1a. In the Boreal realm, the onset of OAE-1a is characterized by a relatively dry climate, followed by the installation of humid climate during the mid-OAE-1a (segments C4-C5) that persists through the remainder of the early Aptian. Across latitudinal belts, there is thus a non-linear change in precipitation pattern during the unfolding of OAE-1a. Similar observations have been made for other OAEs, suggesting more complex weathering feedback mechanism during hyperthermal events than generally assumed.</span></div>

Middle and Late Jurassic climatic, oceanographic and environmental trends along the Viking Corridor

<p><strong>Keywords: Late Jurassic; palaeoclimate; Greenland; carbon cycling; Viking Corridor; belemnite stable isotopes</strong></p><p>The “polar amplification” effect, whereby the poles experience greater changes in temperature compared to the low latitudes for a given global average temperature change, makes high-latitude isotope records ideally suited to investigate fluctuations in palaeoclimate. The present study investigates palaeoclimatic and oceanographic changes along the Viking Corridor – the narrow seaway that connected the Tethys to the Arctic Boreal Realm during the Middle and Late Jurassic.</p><p>Stable-isotope data obtained from belemnites from East Greenland, originating from along the western margin of the Viking Corridor, show a M. Bathonian warming trend, which may indicate the reopening of the corridor after North Sea doming. We also discuss various controls on the carbon-isotope record that may dampen or amplify global signals. Changes in local depositional settings caused partial overprinting of the δ<sup>13</sup>C record during the Late Jurassic VOICE event.</p>

New insight on the paleoclimatic evolution in the Boreal Realm at the onset of the Early Cretaceous chalk sea

<p>During the Albian, the open marine carbonate production underwent a profound revolution with the onset of the dominance of planktonic production in the total carbonate budget. This led to the deposition of vast amounts of chalk across the world’s Oceans as a result of the accumulation of large amounts of nannoplankton. The worldwide Upper Cretaceous white chalks are however not the first true chalks (i.e. deposits dominated by calcareous nannofossils) to be recorded in Earth’s History. Already during the Barremian, chalks were deposited in the North Sea Central Graben. These chalks did not extend until the Albian, since a ‘nannoconid crisis’ occurred at the onset of the early Aptian OAE-1a, with the deposition of an organic-rich marlstone layer named the Fischschiefer. To better understand if climatic changes have governed the occurrence of the Barremian true chalks and the switch to organic-rich marlstones during OAE-1a, we have reconstructed the evolution of climate in the North Sea Basin based on clay mineral assemblages. Clay mineral composition and distribution are proven indicators of paleo­climate and evolution of a basin as the formation of clay minerals in soils depends on the climate under which it develops. Hence, based on high-resolution clay mineral data from various cores from the North Sea, a paleoclimatic reconstruction of the late Hauterivian to early Aptian stratigraphic interval is proposed. Based on a long-term decrease of kaolinite content, a trend toward aridification is observed during the late Barremian, concordant with the development of the first true chalks. A sharp increase in kaolinite content is recorded at the onset of OAE-1a, with its highest peak occurring towards the end of the event. This suggest that a significant increase in humidity accompanies the unfolding of OAE-1a in the North Sea Basin. Further investigation is needed to confirm the hypothesis that paleoclimatic changes in the Boreal Realm are responsible for the onset of chalk deposition and the change in clay mineral assemblages.</p>

Biostratigraphic constraints of the Early Toarcian Oceanic Anoxic Event: new data from calcareous nannofossil investigations of Boreal and Tethyan sections

Calcareous nannofossil biostratigraphy was carried out in Upper Pliensbachian – Lower Toarcian sediments, which cover the Toarcian Oceanic Anoxic Event (T-OAE) interval. In particular, semiquantitative analyses were performed on a total of 156 samples in the composite Sogno Core (Lombardy Basin, Southern Alps) representing a pelagic Tethyan section. Quantitative investigations were applied to additional 168 samples across the Amaltheenton Fm. and Posidonienschiefer Fm., from two cores of the Boreal Realm (Lower Saxony Basin, northern Germany). Primary and secondary events of the Tethyan and Boreal zonations were recognized, allowing the identifications of the NJT5, NJT6 nannofossil Zones for the Sogno Core and the NJ5, NJ6, NJ7 Zones for the German sections, respectively. The sequence of nannofossil biohorizons is generally consistent with data available for various areas at lower and higher latitudes, confirming their reproducibility and reliability for intra and inter-regional correlations. Geochemistry evidences the presence of the negative C isotopic excursion across the “Fish Level” black shale interval expression of the T-OAE in the Sogno Core. The same anomaly is recorded in the German successions at the base of the Posidonia Shale witnessing the passage from well oxygenated to predominantly anoxic conditions. Our results show that the T-OAE C isotopic excursion recorded in the Sogno Core is excellently constrained by the first occurrence (FO) of Carinolithus superbus at the onset and the last occurrence (LO) of Mitrolithus jansae at the end. A significant decrease in abundance and size of Schizosphaerella punctulata (the “S. punctulata crisis”) and an abundance drop of M. jansae further characterise the T-OAE perturbation. Only S. punctulata shows a recovery at the end of the T-OAE, while M. jansae barely survived the palaeoenviromental stress and disappeared soon after its termination. The extreme rareness of S. punctulata and the absence of M. jansae in the Boreal Realm prevent the recognition of the “S. punctulata crisis” and the M. jansae decline. Our study reveals the LO of Biscutum finchii together with the FO of C. superbus as an additional event approximating the onset of the C isotopic excursion exclusively in the German successions. Further events, such as the LOs of Biscutum grandis, Crepidolithus granulatus and Parhabdolithus liasicus are detected within the C isotopic anomaly exclusively in the German sections. Nannofossil biostratigraphy permits the effective dating and correlating of Early Jurassic major palaeoceanographic events and particularly of the T-OAE which are of a great importance to derive a definitive model for the Posidonia Shale deposition.

Biostratigraphic constraints of the Early Toarcian Oceanic Anoxic Event: new data from calcareous nannofossil investigations of Boreal and Tethyan sections

Calcareous nannofossil biostratigraphy was carried out in Upper Pliensbachian – Lower Toarcian sediments, which cover the Toarcian Oceanic Anoxic Event (T-OAE) interval. In particular, semiquantitative analyses were performed on a total of 156 samples in the composite Sogno Core (Lombardy Basin, Southern Alps) representing a pelagic Tethyan section. Quantitative investigations were applied to additional 168 samples across the Amaltheenton Fm. and Posidonienschiefer Fm., from two cores of the Boreal Realm (Lower Saxony Basin, northern Germany). Primary and secondary events of the Tethyan and Boreal zonations were recognized, allowing the identifications of the NJT5, NJT6 nannofossil Zones for the Sogno Core and the NJ5, NJ6, NJ7 Zones for the German sections, respectively. The sequence of nannofossil biohorizons is generally consistent with data available for various areas at lower and higher latitudes, confirming their reproducibility and reliability for intra and inter-regional correlations. Geochemistry evidences the presence of the negative C isotopic excursion across the “Fish Level” black shale interval expression of the T-OAE in the Sogno Core. The same anomaly is recorded in the German successions at the base of the Posidonia Shale witnessing the passage from well oxygenated to predominantly anoxic conditions. Our results show that the T-OAE C isotopic excursion recorded in the Sogno Core is excellently constrained by the first occurrence (FO) of Carinolithus superbus at the onset and the last occurrence (LO) of Mitrolithus jansae at the end. A significant decrease in abundance and size of Schizosphaerella punctulata (the “S. punctulata crisis”) and an abundance drop of M. jansae further characterise the T-OAE perturbation. Only S. punctulata shows a recovery at the end of the T-OAE, while M. jansae barely survived the palaeoenviromental stress and disappeared soon after its termination. The extreme rareness of S. punctulata and the absence of M. jansae in the Boreal Realm prevent the recognition of the “S. punctulata crisis” and the M. jansae decline. Our study reveals the LO of Biscutum finchii together with the FO of C. superbus as an additional event approximating the onset of the C isotopic excursion exclusively in the German successions. Further events, such as the LOs of Biscutum grandis, Crepidolithus granulatus and Parhabdolithus liasicus are detected within the C isotopic anomaly exclusively in the German sections. Nannofossil biostratigraphy permits the effective dating and correlating of Early Jurassic major palaeoceanographic events and particularly of the T-OAE which are of a great importance to derive a definitive model for the Posidonia Shale deposition.