Microbial-, fusulinid limestones with large gastropods and calcareous algae: an unusual facies from the Early Permian Khao Khad Formation of Central Thailand

The Early Permian (Kungurian) Khao Khad Formation of Central Thailand consists mostly of carbonates deposited on the western margin of the Indochina Terrane. This formation has yielded unusual microbial-fusulinid limestones with large gastropods which contribute most to the rock volume. With a height of more than 6 cm, the gastropods are amongst the largest Early Permian gastropods ever reported. Gastropods as major rock formers are rare in the Palaeozoic. This, and other recently reported invertebrate faunas from Thailand show that gastropods may dominate Permian fossil assemblages not only in diversity, but also regarding abundance and in some cases also regarding biomass. Besides gastropods, fusulinids, various calcareous algae, intraclasts and thick microbial-cyanobacterial (Girvanella and Archaeolithoporella) coatings and reticular microbial patches as well as thick inter- and intragranular radial fibrous cement crusts are present. The gastropods represent at least four species and belong probably to undescribed taxa. The fusulinid genus Pseudofusulina and Misellina (M.) termieri are reported from the Khao Khad Formation for the first time and indicate a Bolorian age. Calcareous algae are dominated by dasycladaceans followed by gymnocodiaceans and solenoporaceans. The studied limestone almost completely lacks metazoan reef builders such as corals and sponges. Likewise, brachiopods and bivalves are absent in the studied samples and echinoderms are very scarce. The carbonate is interpreted as product of shallow water, back-reef lagoonal platform community with a high productivity providing the large gastropods with sufficient food. However, conditions were too eutrophic for sessile filter feeders including metazoan reef builders.

A Maastrichtian microbial reef and associated limestones in the Roca Formation of Patagonia (Neuquén Province, Argentina)

We describe a small microbial reef and associated limestones occurring in a Maastrichtian transgressive succession of mixed carbonate-siliciclastic lithologies at Sierra Huantraico near Chos Malal (Neuquén, Argentina). Strontium isotope data suggest that the reef is of earliest Maastrichtian age. The small reef (0.8 m thick, 2 m wide) is mostly composed of peloidal bindstone, dense stromatolite-cement crusts and thrombolite. Except for some ostracods, no metazoan fossils were found in the reef structure, although the majority of peloids are fecal pellets, probably of larger crustaceans. Small foraminifers with calcite tests and probable green algae have also been noted. Sedimentological data and fossils within and immediately above the reef suggest that the reef was formed in a transgressive systems tract under freshwater to brackish-water conditions. Limestones above the reef are serpulid-bryozoan packstones and intraclast-ooid grainstones. These limestones yield a mixture of typical non-tropical (common serpulids and bryozoans) and typical tropical aspects (common dasycladaceans and ooids). This mosaic is explained by salinity fluctuations, which in our case dominate over temperature in determining the grain associations. Wir beschreiben ein kleines mikrobielles Riff, das in der Sierra Hunatraico (Neuquén, Argentinien) in einer transgressiven, gemischt siliziklastisch-kalkigen Abfolge gefunden wurde. Nach Strontiumisotopen-Datierung ist das Riff in das unterste Maastrichtium zu stellen. Das kleine Riff (0,8 m Mächtigkeit, 2 m Breite) besteht überwiegend aus peloidalem Bindstone, dichten Stromatolith-Zement-Krusten und Thrombolith. Mit Ausnahme von Ostrakoden konnten keine Metazoen in der Riffstruktur nachgewiesen werden, obwohl die Mehrzahl der Peloide als Kotpillen zu interpretieren sind, die vermutlich auf größere Krebse zurückgehen. Kleine Foraminiferen und mögliche Grünalgen sind die einzigen zusätzlich nachweisbaren Eukaryoten. Die Fossilien im Riff und in den überlagernden Kalken sprechen für ein Riffwachstum unter transgressiven aber hyposalinen Bedingungen. Die Kalke über dem Riff tragen ein gemischtes paläoklimatisches Signal, das sowohl typisch nicht-tropische als auch typisch tropische Komponenten beinhaltet. Dieses Mosaik ist möglicherweise durch die starken Salinitätsschwankungen erklärbar und erfordert ein Überdenken der bisherigen Modelle zur klimatischen Steuerung der Karbonatsedimentation. doi:10.1002/mmng.200600007