The Phycosiphon Ichnofacies and the Rosselia Ichnofacies: Two new ichnofacies for marine deltaic environments

2020 ◽  
Vol 90 (8) ◽  
pp. 855-886 ◽  
Author(s):  
James A. MacEachern ◽  
Kerrie L. Bann

ABSTRACT Seilacherian Ichnofacies have been established, to date, for characterizing relatively stable depositional settings. Environments characterized by temporally and spatially varying physico-chemical stresses, however, have languished and been described ichnologically in the context of their “departures” from the archetypal expressions of otherwise ambient environments. Correspondingly, discrimination between shoreface and marine deltaic deposits have been addressed mainly by identifying variations in the individual trace-fossil suites without an over-arching ichnological model. Based on trace-fossil suites reported from globally distributed strata throughout the Phanerozoic, the case can now be made for erecting two new temporally and geographically recurring Seilacherian Ichnofacies for marine deltaic successions—the Phycosiphon Ichnofacies for muddy prodelta environments and the Rosselia Ichnofacies for sandy delta-front settings. The Phycosiphon Ichnofacies is characterized by variable bioturbation intensities (BI 0–5), although many sandstone and mudstone beds may have very low bioturbation intensities (BI 0–1), pointing to their rapid, event-style deposition. Most biogenic structures record grazing or deposit-feeding behaviors, with subordinate horizontal dwellings that reflect deposit feeding and/or carnivory. Meiofaunal cryptic bioturbation is locally present in tempestites. Episodic deposition is accompanied by common escape structures and locally, sediment-swimming structures. The Phycosiphon Ichnofacies typically shows beds characterized by diverse, fully marine trace-fossil suites intercalated with discrete beds dominated by low-diversity, facies-crossing traces. Such bed-scale juxtaposition points to short-term but recurring alternations between stable marine conditions and physico-chemically stressed conditions. The Rosselia Ichnofacies is also characterized by variable (BI 0–5; though typically BI 0–3) bioturbation intensities and sporadically distributed burrows. Most ichnogenera record deposit-feeding behaviors, many associated with vertically and horizontally oriented permanent dwelling structures. Dwelling structures commonly display re-equilibration and escape structures are typically abundant, both characteristic of elevated sedimentation rates and sporadic deposition. Most spreitenated structures are markedly retrusive, also attesting to elevated rates of sedimentation. Cryptic bioturbation is only locally abundant, particularly associated with erosionally amalgamated tempestites lacking mudstone drapes. While many of the trace fossils constitute facies-crossing elements, fully marine (ambient) ichnogenera also occur in some suites. Associated mudstone interbeds display low BI values, sediment-swimming structures, and top-down bioturbation, consistent with their rapid deposition as fluid mud. The two ichnofacies may pass gradationally into one another vertically, associated with lobe progradation or autogenic lobe abandonment. Further, the two ichnofacies may pass along depositional strike into their non-deltaic counterparts—the Phycosiphon Ichnofacies grading into the Cruziana Ichnofacies in distal positions, and the Rosselia Ichnofacies passing into the Skolithos Ichnofacies in shallow-water locales. The erection of these new ichnofacies will enhance the reliable identification of deltaic successions, particularly in wave-dominated settings, and their differentiation from classical strandplain shoreface deposits. As well, the two ichnofacies better explain animal–substrate relations in the context of the detailed sedimentological characteristics of delta deposits, refining the overall facies characterization of shallow-marine environments.

Author(s):  
Luis A. Buatois ◽  
Gabriela M. Mangano ◽  
Timothy R. Carr

Integration of facies and trace-fossil evidence tests and refines depositional models constructed solely on the basis of physical sedimentology. In recent years, the petroleum industry has increasingly used trace-fossil analysis of cores as an aid in reservoir characterization. In particular, ichnologic data have been instrumental in the recognition of estuarine deposits and their distinction from open-marine facies (e.g., MacEachern and Pemberton, 1994). Previous ichnologic analyses of cores, however, have concentrated on post-Paleozoic reservoirs (e.g., Bockelie, 1991; Pemberton, 1992; Taylor and Gawthorpe, 1993; Howell et al., 1996; Martin and Pollard, 1996; MacEachern and Pemberton, 1997). The present study represents one of the first attempts to apply trace-fossil analysis to cores from Paleozoic reservoirs. The Lower Pennsylvanian Morrow Sandstone contains oil and gas reservoirs in a wide variety of shallow and marginal-marine depositional environments. Delta-front, shoreface, and estuarine valley-fill reservoir sandstones are encased in offshore and estuarine mudstones (Sonnenberg, 1985; Krystinik and Blakeney, 1990; Sonnenberg et al., 1990; Wheeler et al., 1990). An integrated stratigraphic, sedimentologic, and ichnologic study provides a more accurate characterization of reservoir facies and geometry. This study allows distinction between marine-shoreface and estuarine valley-fill sandstones from four cores of the lower Morrow in southwestern Kansas. Core analysis subsequently was integrated with well-log information. Previous studies have emphasized the presence of estuarine valley-fills in the upper Morrow (Wheeler et al., 1990). Our integrated approach extends the estuarine valley interpretation into the lower Morrow. Within the midcontinent, trace fossils are useful in distinguishing different facies in estuarine incised valleys and marine shorefaces. Detailed study of biogenic structures provides high-resolution information to solve problems in facies, stratigraphic, and reservoir modeling. In some cases, they represent the only evidence available to develop a reasonable picture of depositional conditions and to estimate reservoir heterogeneity. The present study provides a detailed analysis of the sedimentary facies, documents the associated trace fossils, and illustrates how trace fossils are used to refine environmental interpretations of the lower Morrow sandstone reservoirs.


1993 ◽  
Vol 67 (6) ◽  
pp. 1070-1074 ◽  
Author(s):  
Ron K. Pickerill ◽  
Stephen K. Donovan ◽  
Harold L. Dixon

Rosette-shaped problematica are relatively common structures in the Phanerozoic rock record. Historically, they have been accorded a variety of names and documented from various shallow to deep marine environments. Unfortunately, the detailed interpretation of many such structures as biogenic (trace fossils, medusoids, or other body fossils; see, for example, Häntzschel, 1970, 1975) or nonbiogenic (for example, Pickerill and Harris, 1979) in origin still remains to be resolved. However, a detailed analysis of one such structure by Fürsich and Bromley (1985), namely Dactyloidites Hall, 1886, convincingly demonstrated its biogenic origin. The distinctive morphology of Dactyloidites and its synonyms was interpreted by Fürsich and Bromley (1985) to result from successive probings of an essentially stationary deposit-feeding, worm-like organism, possibly possessing a proboscis, to produce a rosetted, vertical spreiten with a centrally located, vertical or subvertical shaft.


The structural genes for three forms of Rubisco have been isolated from bacteria and introduced into various plasmids. Apart from details of the sequences which have been obtained from these constructs, they are now being exploited for mutagenesis to determine the identity and specific function of the individual amino acid residues that compose the active site. These methods have been applied to a plasmid that contains the structural gene for the simplest form of Rubisco from Rhodospirillum rubrum to obtain mutant enzymes with altered activity. The construct pRR2119 is also expressed to very high levels in Escherichia coli and enough recombinant protein of both the wild-type and m utant enzymes can be obtained for detailed physico-chemical studies. Other vectors have now been constructed, containing the genes of prokaryotic Rubisco that assemble into an active form I enzyme. The levels of expression are acceptable and the product is similar to the authentic enzyme. These constructs are now being used for mutagenesis in vitro to attempt to alter the relative rates of the oxygenase and carboxylase activities.


2010 ◽  
Vol 84 (4) ◽  
pp. 730-738 ◽  
Author(s):  
Noelia B. Carmona ◽  
María Gabriela Mángano ◽  
Luis A. Buatois ◽  
Juan José Ponce

Lower Miocene tide-influenced deltaic deposits from the Chenque Formation, Patagonia, Argentina, contain abundant and well-preserved biogenic structures attributed to locomotion of deposit-feeder protobranch bivalves. These trace fossils, assigned to the ichnogenus Protovirgularia, consist of delicate, inclined-to-horizontal, chevronate structures, mostly symmetrical with respect to a median axis. Identification of Protovirgularia at sandstone sole beds (hypichnion) is quite straightforward. Endichnial, exichnial and epichnial preservation in heterolithic facies, however, provides a wide variety of forms that depart from the archetypal Protovirgularia and challenges ichnotaxonomic classification. Specimens in prodelta and delta-front facies display morphologic features controlled by substrate fluidity, toponomy, and sedimentation rate. Most specimens show sharp, closely spaced chevrons and occur along sandstone/mudstone interfaces of the proximal prodelta and distal delta-front deposits. These forms reflect how tracemakers experienced significant friction while advancing through the sediment, which resulted in relatively smaller increments of movements. In contrast, variants of Protovirgularia formed in muddier beds, such as in prodeltaic facies, show irregular, poorly defined and unevenly spaced chevrons, and are locally asymmetric in relation to the axis, reflecting softer, water-rich, and plastic substrates. This sediment offered relatively low friction but poor anchorage for the foot. These occurrences of Protovirgularia in tide-influenced, marginal-marine deposits suggests that protobranchs were tolerant of fluctuations in salinity, sedimentation rates, turbidity, and oxygen depletion, displaying opportunistic strategies in stressed nearshore environments. Our evaluation of taphonomic controls and appropriate identification of Protovirgularia can provide valuable information for expanding our knowledge of the ethology and paleoecology of protobranch bivalves.


Open Medicine ◽  
2006 ◽  
Vol 1 (4) ◽  
pp. 370-378 ◽  
Author(s):  
Magdaléna Májeková ◽  
Vladimír Šnirc ◽  
Svorad Štolc ◽  
Pavol Májek ◽  
Želmíra Bezáková ◽  
...  

Abstractα1 — adrenolytic activities of pyridoindole derivatives recently synthesized in the Institute of Experimental Pharmacology, Slovak Academy of Sciences, were measured. A characteristic set of derivatives (five active, one with a threshold activity and two inactive) was chosen and an elementary structure-activity study was performed. The structure and energy properties were estimated by quantum-chemical semiempirical AM1 and molecular mechanics methods. The ionization constants pKα of the individual derivatives were calculated by program Pallas or estimated by potenciometric titration. The α 1 blocking activities were measured by rat thoracic aorta model. The experimental model used was not α 1 — adrenoreceptor subtypes specific, however, the α 1D subtype could be considered to be a predominant type in a rat aorta. The obtained physico-chemical parameters were then compared with the blocking activities of the derivatives and following determining characters for α 1 — adrenolytic activities were determined: 1) the polarity of the substituted phenol ring represented by a map of molecular electrostatic potential and 2) the hexahydro-pyridine nitrogen pKα constant, which represents the ability of the compound to be protonated by physiologic pH.


2016 ◽  
Vol 8 (3-4) ◽  
pp. 485-492
Author(s):  
E. Franzini ◽  
F. Driss ◽  
Ph. Darcet ◽  
Fr. Driss ◽  
F. Daoud ◽  
...  

The measurement of the combining weights of hydrogen and oxygen has been the subject of so many researches of a high order of excellence that any fresh investigation of this fundamental constant must be submitted with considerable diffidence. Nevertheless, it must be noted that the results obtained by various observers differ appreciably. According to Clarke ( 1 ), the values obtained by Morley and Noyes, by reason of the accuracy of their methods and the close concordance of the individual determinations, outweigh the results of all other investigators. The atomic weight of oxygen being 16, that of hydrogen, according to Morley( 2 ), is 1˙00762, and according to Noyes ( 3 ), 1˙00787. (Clarke, on Noyes’ data, prefers the value 1˙00783.) It is, further, a significant fact that the arithmetic mean of all determinations discussed by Clarke, lies between these two values, which differ by 1 part in 4000. Both values are based on the gravimetric synthesis of water and are independent of a knowledge of the densities of the gases. A physico-chemical method of determining the relative molecular weights depends on the knowledge of the ratio of the densities, together with that of the combining volumes.


2021 ◽  
Vol 7 (8) ◽  
pp. 119
Author(s):  
Miloš Baljozović ◽  
Xunshan Liu ◽  
Olha Popova ◽  
Jan Girovsky ◽  
Jan Nowakowski ◽  
...  

Single layer low-dimensional materials are presently of emerging interest, including in the context of magnetism. In the present report, on-surface supramolecular architecturing was further developed and employed to create surface supported two-dimensional binary spin arrays on atomically clean non-magnetic Au(111). By chemical programming of the modules, different checkerboards were produced combining phthalocyanines containing metals of different oxidation and spin states, diamagnetic zinc, and a metal-free ‘spacer’. In an in-depth, spectro-microscopy and theoretical account, we correlate the structure and the magnetic properties of these tunable systems and discuss the emergence of 2D Kondo magnetism from the spin-bearing components and via the physico-chemical bonding to the underlying substrate. The contributions of the individual elements, as well as the role of the electronic surface state in the bottom substrate, are discussed, also looking towards further in-depth investigations.


Author(s):  
Winarti Winarti ◽  
Arman Harahap

Macrozoobenthos used as a bioindicator of the waters, this happens because of the nature of the benthos which has mpergerakan low so it got a lot of influence from the environment. This study aims to determine the condition of the river Kundur with a view the community structure of macrozoobenthos which include: species composition, abundance, diversity, uniformity and dominance as well as the physical and chemical parameters of water as a paremeter supporters. Based on the results of observations, macrozoobenthos were found during the study in the river Kundur consists of 3 phylum, 4 classes, 17 family with 11 species. The composition of the class of macrozoobenthos in the whole of the observation station consists of a Gastropod (37%), Insecta (23%), Malacostraca (20%), Polychaeta (10%) dan Malacostraca (20%). The value of an abundance ranged from 325,7 - 3309,6 ind/m3. With the value of the total abundance at Station I which is equal to 3690,1 ind/m3, Station II 3723,1 ind/m3, Station III 3692,3 ind/m3 can be categorized diversity are light polluted with the value of the index of uniformity (E) that describes the spread of the individual tends to be uniform or relatively the same. While the condition of the physical-chemical parameters and the substrate bottom of the river Kundur in general has a range of relatively homogeneous, spread evenly throughout the observation stations and still be able to support life macrozoobenthos. To the results of measurements of the parameters of physico-chemical water and substrate in the river kundur in general has a range of relatively homogeneous in the whole observation stations and still be able to support the life of the macrozoobenthos.


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