Time budgets and synchrony of activities in muskoxen: the influence of sex, age, and season

1997 ◽  
Vol 75 (10) ◽  
pp. 1628-1635 ◽  
Author(s):  
Steeve D. Côté ◽  
James A. Schaefer ◽  
François Messier
Keyword(s):  
Group Size ◽  
Time Budget ◽  
High Arctic ◽  
Late Winter ◽  
Time Budgets ◽  
Adult Males ◽  
Ovibos Moschatus ◽  
Group Type ◽  
June July ◽  
Victoria Island

Synchrony of activities and time budgets of age–sex classes of muskoxen (Ovibos moschatus) was studied on Victoria Island in the Canadian High Arctic during late winter (April – late May), spring (late May – early June) and summer (mid-June – July). As revealed by the kappa (κ) coefficient of agreement, herd members exhibited significant synchrony in 79 of 82 groups. The degree of synchrony was not related to group size in mixed herds but decreased with group size in bachelor herds. Diversity in age–sex classes, group type (mixed versus bachelor), and season did not affect synchrony. However, synchrony was less among adult males than among other age–sex classes. Muskoxen were more synchronised at the start of activity bouts than at the end. As revealed by principal components analysis, calves tended to spend more time lying and standing and less time feeding than other age–sex classes. In addition, males in bachelor groups tended to feed more and rest less than males in mixed herds. Duration of active and lying bouts was not affected by age–sex class. Length of active bouts did not change with season, but lying bouts decreased in length significantly from late winter to spring and from spring to summer. We suggest that synchrony of activities maintains group cohesion but constrains the time budget of some group members, especially adult males.

Seasonal Habitat Use by Nubian Ibex (Capra Nubiana) Evaluated with Behavioral Indicators

2011 ◽  
Vol 57 (3) ◽  
pp. 223-246 ◽  
Author(s):  
Solomon A. Tadesse ◽  
Burt P. Kotler
Keyword(s):  
Group Size ◽  
Age Structure ◽  
Time Allocation ◽  
Environmental Variables ◽  
Time Budget ◽  
Habitat Type ◽  
Time Budgets ◽  
Behavioral Models ◽  
Risk Of Predation ◽  
Focal Observation

Nubian ibex (Capra nubiana) prefer steep terrain in their landscape to reduce risks of predation and human nuisance disturbances. They also use vigilance and time allocation to manage risk of predation. We studied time budgets and habitat selection of Nubian ibex to: (1) identify the habitat variables to which Nubian ibex were behaviorally responsive; (2) investigate how time budget responses of Nubian ibex were related to season, slope condition, group size, and sex-age structure; and (3) develop behavioral-based models that account for the variations in the behaviors of Nubian ibex across the landscape and seasons.To quantify time budgets, we took regular field observations on focal individuals of Nubian ibex classified according to their habitat, group size, sex, and age. For each focal observation, we quantified environmental variables that were thought to influence the behavioral responses of ibex. Then, we developed behavioral models by correlating the proportion of behaviors measured in focal animal observations to the influential environmental variables. The behaviors of Nubian ibex significantly varied with sex and age structure, season, habitat type, and slope conditions. Adult females are more vigilant than adult male ibex, especially in the spring. This correlates with breeding and nursing activities. Based on the characteristics of the habitat, ibex behave to minimize risks of predation and human nuisance disturbances while maximizing their food intake.

Time-budget variations in relation to density-dependent social interactions in female and yearling male fallow deer during the rut

1997 ◽  
Vol 75 (6) ◽  
pp. 971-977 ◽  
Author(s):  
Christophe Pélabon ◽  
Petr E. Komers
Keyword(s):  
Population Density ◽  
Social Interactions ◽  
Time Budget ◽  
Resource Depletion ◽  
Fallow Deer ◽  
Operational Sex Ratio ◽  
Time Budgets ◽  
Adult Males ◽  
Density Dependent ◽  
Male Density

We investigated time-budget variations among female and yearling (socially immature 1.5-year-old) male fallow deer (Dama dama) during the rutting season in six captive populations in central Sweden. The effects of population density, male density, and operational sex ratio on time budgets were analyzed. Although no differences were apparent before the rut, populations differed in the time budgets of females and yearling males during the rut. During this period, the time spent by females in feeding and resting activities of yearling males decreased with increasing population density, whereas feeding and resting activities of yearling males decreased with increasing male density. Behaviour of adult males during the rut and interactions of adult males with other animals appeared to be the principal factors affecting the time budgets of both females and yearling males. Our results show that density-dependent effects may occur through social interactions alone, regardless of the effects on resource depletion. The decrease in the proportion of time spent in restoring (lying and grazing) behaviours represents an energy cost for both females and young males. Given the timing of the rut just before the winter, this cost may negatively effect the condition of individuals.

Effects of habitat, group-size, sex-age class and seasonal variation on the behavioural responses of the mountain nyala (Tragelaphus buxtoni) in Munessa, Ethiopia

2013 ◽  
Vol 30 (1) ◽  
pp. 33-43 ◽  
Author(s):  
Solomon A. Tadesse ◽  
Burt P. Kotler
Keyword(s):  
Group Size ◽  
Time Budget ◽  
Activity Patterns ◽  
Habitat Type ◽  
Dry Season ◽  
Adult Males ◽  
Wet Season ◽  
Behavioural Responses ◽  
Age Class ◽  
Habitat Variables

Abstract:Activity patterns of animals are generally influenced by many factors. We hypothesized that the behavioural responses (i.e. activity time-budget allocated to vigilance, feeding and moving) of mountain nyala (Tragelaphus buxtoni) should vary with habitat type, season, group-size and sex-age class. We randomly established a total of 12 permanent walking transects with the aid of a GPS device across three major habitat types used by the mountain nyala (i.e. four transects in each habitat). Following each transect, we conducted focal-animal observations to quantify the time-budget allocated to vigilance, feeding and moving. A total of 119 and 116 focal-animals were assessed in the wet and dry season respectively. Moreover, along each transect, seven habitat variables were collected in systematically laid 109 circular plots each with a 5-m radius (i.e. 31, 41 and 37 plots in the cleared vegetation, plantation and natural forest respectively) in the wet and dry season. We developed behavioural models by correlating the time-budget (i.e. proportion of time vigilance, feeding and moving) of the focal-animals in accordance with habitat variables, group-size and sex-age class. In the wet season, mountain nyala devoted most of their time to vigilance, but they allocated the largest proportion of their time to moving in the dry season. Vigilance differed among the three habitats and was highest in the cleared vegetation during the dry season. Contrary to expectations, adult males were more vigilant than both adult females and sub-adults during the dry season. The behavioural models based on time-budget help to predict how the mountain nyala perceive their environment and trade-off between food acquisition and safety in the wet and dry season. The study also improves our understanding of the adaptive behavioural ecology of the endangered mountain nyala.

Group size modulates time budget and foraging efficiency in captive Skylarks, Alauda arvensis

2011 ◽  
Vol 153 (2) ◽  
pp. 485-490 ◽  
Author(s):  
Thibaut Powolny ◽  
Cyril Eraud ◽  
Vincent Bretagnolle
Keyword(s):  
Group Size ◽  
Time Budget ◽  
Foraging Efficiency ◽  
Alauda Arvensis

Time-Budgets of Camargue Horses Ii. Time-Budgets of Adult Horses and Weaned Sub-Adults

Behaviour ◽  
1980 ◽  
Vol 72 (1-2) ◽  
pp. 26-48 ◽  
Author(s):  
Patrick Duncan
Keyword(s):  
Social Structure ◽  
Social System ◽  
General Trend ◽  
Small Population ◽  
Time Budgets ◽  
Adult Males ◽  
Adult Females ◽  
Young Horses

AbstractTime-budgets of adult and weaned sub-adult horses were studied in a small population of Camargue horses living in semi-liberty. The categories of activities used were: Standing resting, Lying flat, Lying up, Standing alert, Walking, Trotting, Galloping, Rolling and Foraging. The main differences in time-budgets were related to age and to sex : young horses spent more time lying (sleeping), males spent more time standing alert and in rapid movements (trot, gallop), while usually foraging less than did the adult females. During the three years of the study the population increased from 20 to 54 horses and there were considerable changes in social structure as the number of adult males increased. Associated with these developments there were some changes between years in the time-budgets: the most striking of which was a general trend for all horses to spend less time lying. Nonetheless the time-budgets showed a considerable constancy across years and age/sex-classes, especially with regard to time spent foraging. This conclusion may provide a clue as to why horses have an unusual social system based on long term relationships between a male and the females of his harem.

scholarly journals Characteristics of Arctic tides at CANDAC-PEARL (80° N, 86° W) and Svalbard (78° N, 16° E) for 2006–2009: radar observations and comparisons with the model CMAM-DAS

2011 ◽  
Vol 29 (10) ◽  
pp. 1939-1954 ◽  
Author(s):  
A. H. Manson ◽  
C. E. Meek ◽  
X. Xu ◽  
T. Aso ◽  
J. R. Drummond ◽  
...  
Keyword(s):  
Middle Atmosphere ◽  
Radar Data ◽  
High Arctic ◽  
Wind Component ◽  
Meteor Radar ◽  
The North ◽  
Wave Numbers ◽  
Central Canada ◽  
June July ◽  
Abstract Operation

Abstract. Operation of a Meteor Radar (MWR) at Eureka, Ellesmere Island (80° N, 86° W) began in February 2006: this is the location of the Polar Environmental and Atmospheric Research Laboratory (PEARL), operated by the "Canadian Network for the Detection of Atmospheric Change" (CANDAC). The first 36 months of tidal wind data (82–97 km) are here combined with contemporaneous tides from the Meteor Radar (MWR) at Adventdalen, Svalbard (78° N, 16° E), to provide the first significant evidence for interannual variability (IAV) of the High Arctic's diurnal and semidiurnal migrating (MT) and non-migrating tides (NMT). The three-year monthly means for both diurnal (DT) and semi-diurnal (SDT) winds demonstrate significantly different amplitudes and phases at Eureka and Svalbard. Typically the summer-maximizing DT is much larger (~24 m s−1 at 97 km) at Eureka, while the Svalbard tide (5–24 m s−1 at 97 km)) is almost linear (north-south) rather than circular. Interannual variations are smallest in the summer and autumn months. The High Arctic SDT has maxima centred on August/September, followed in size by the winter features; and is much larger at Svalbard (24 m s−1 at 97 km, versus 14–18 m s−1 in central Canada). Depending on the location, the IAV are largest in spring/winter (Eureka) and summer/autumn (Svalbard). Fitting of wave-numbers for the migrating and non-migrating tides (MT, NMT) determines dominant tides for each month and height. Existence of NMT is consistent with nonlinear interactions between migrating tides and (quasi) stationary planetary wave (SPW) S=1 (SPW1). For the diurnal oscillation, NMT s=0 for the east-west (EW) wind component dominates (largest tide) in the late autumn and winter (November–February); and s=+2 is frequently seen in the north-south (NS) wind component for the same months. The semi-diurnal oscillation's NMT s=+1 dominates from March to June/July. There are patches of s=+3 and +1, in the late fall-winter. These wave numbers are also consistent with SPW1-MT interactions. Comparisons for 2007 of the observed DT and SDT at 78–80° N, with those within the Canadian Middle Atmosphere Model Data Assimilation System CMAM-DAS, are a major feature of this paper. The diurnal tides for the two locations have important similarities as observed and modeled, with seasonal maxima in the mesosphere from April to October, and similar phases with long/evanescent wavelengths. However, differences are also significant: observed Eureka amplitudes are generally larger than the model; and at Svalbard the modeled tide is classically circular, rather than anomalous. For the semi-diurnal tide, the amplitudes and phases differ markedly between Eureka and Svalbard for both MWR-radar data and CMAM-DAS data. The seasonal variations from observed and modeled archives also differ at each location. Tidal NMT-amplitudes and wave-numbers for the model differ substantially from observations.

scholarly journals Intra-annual and intra-seasonal flow dynamics of a High Arctic polythermal valley glacier

2003 ◽  
Vol 37 ◽  
pp. 181-188 ◽  
Author(s):  
Robert G. Bingham ◽  
Peter W. Nienow ◽  
Martin J. Sharp
Keyword(s):  
Drainage System ◽  
High Arctic ◽  
Ablation Zone ◽  
Accumulation Zone ◽  
Seasonal Flow ◽  
Excess Surface ◽  
Velocity Response ◽  
Spatio Temporal ◽  
June July ◽  
Arctic Glacier

AbstractMeasurements of surface dynamics on polythermal John Evans Glacier, Nunavut, Canada, over two winter periods and every 7–10 days throughout two melt seasons (June–July 2000, 2001) provide new insight into spatio-temporal patterns of High Arctic glacier dynamics. In the lower ablation zone, mean annual surface velocities are 10–21 m a–1, but peak velocities up to 50% higher are attained during late June/early July. In the upper ablation zone and lower accumulation zone, mean annual surface velocities are typically 10–18 m a–1, and peak velocities up to 40% higher occur during late July. In the upper accumulation zone, mean annual surface velocities are 2–9 m a–1, and motion in mid- to late July exceeds this by up to 10%. Rapid drainage of ponded supraglacial water in the upper ablation zone to an initially distributed subglacial drainage system in mid-June may force excess surface motion in the warm-based lower glacier. The data indicate that the duration of the velocity response may be related to the rate of channelization of the basal drainage, and the velocity response may be transmitted up-glacier by longitudinal coupling. An increase in surface velocities in the middle glacier in late July occurs in conjunction with the opening of two further moulins in the accumulation zone.

scholarly journals Technical Note: A mobile sea-going mesocosm system – new opportunities for ocean change research

2013 ◽  
Vol 10 (3) ◽  
pp. 1835-1847 ◽  
Author(s):  
U. Riebesell ◽  
J. Czerny ◽  
K. von Bröckel ◽  
T. Boxhammer ◽  
J. Büdenbender ◽  
...  
Keyword(s):  
Ocean Acidification ◽  
Environmental Changes ◽  
Biogeochemical Cycling ◽  
High Arctic ◽  
Technical Note ◽  
Plankton Community ◽  
Small Scale ◽  
Multiple Test ◽  
The Baltic ◽  
June July

Abstract. One of the great challenges in ocean change research is to understand and forecast the effects of environmental changes on pelagic communities and the associated impacts on biogeochemical cycling. Mesocosms, experimental enclosures designed to approximate natural conditions, and in which environmental factors can be manipulated and closely monitored, provide a powerful tool to close the gap between small-scale laboratory experiments and observational and correlative approaches applied in field surveys. Existing pelagic mesocosm systems are stationary and/or restricted to well-protected waters. To allow mesocosm experimentation in a range of hydrographic conditions and in areas considered most sensitive to ocean change, we developed a mobile sea-going mesocosm facility, the Kiel Off-Shore Mesocosms for Future Ocean Simulations (KOSMOS). The KOSMOS platform, which can be transported and deployed by mid-sized research vessels, is designed for operation in moored and free-floating mode under low to moderate wave conditions (up to 2.5 m wave heights). It encloses a water column 2 m in diameter and 15 to 25 m deep (∼50–75 m3 in volume) without disrupting the vertical structure or disturbing the enclosed plankton community. Several new developments in mesocosm design and operation were implemented to (i) minimize differences in starting conditions between mesocosms, (ii) allow for extended experimental duration, (iii) precisely determine the mesocosm volume, (iv) determine air–sea gas exchange, and (v) perform mass balance calculations. After multiple test runs in the Baltic Sea, which resulted in continuous improvement of the design and handling, the KOSMOS platform successfully completed its first full-scale experiment in the high Arctic off Svalbard (78°56.2′ N, 11°53.6′ E) in June/July 2010. The study, which was conducted in the framework of the European Project on Ocean Acidification (EPOCA), focused on the effects of ocean acidification on a natural plankton community and its impacts on biogeochemical cycling and air–sea exchange of climate-relevant gases. This manuscript describes the mesocosm hardware, its deployment and handling, CO2 manipulation, sampling and cleaning, including some further modifications conducted based on the experiences gained during this study.

scholarly journals Seasonal activity pattern of Swamp deer (Rucervus duvaucelii duvaucelii) in Dudhwa National Park, Uttar Pradesh, India

2014 ◽  
Vol 6 (1) ◽  
pp. 246-253 ◽  
Author(s):  
Kaleem Ahmed ◽  
Jamal A. Khan
Keyword(s):  
Activity Pattern ◽  
National Park ◽  
Time Budget ◽  
Activity Patterns ◽  
Winter Season ◽  
Uttar Pradesh ◽  
Seasonal Activity ◽  
Adult Males ◽  
Continuous Feeding ◽  
Swamp Deer

The data on activity pattern and time budget of Swamp deer (Rucervus duvauceli duvauceli) were collected through instantaneous scan sampling from Dudhwa National Park, Uttar Pradesh, India Diurnal activity pattern of Swamp deer showed marked reduction in resting in winter as compared to summer. In winter resting in adult males showed polymodal pattern with peaks occurring at different hours of the day and continued throughout the day without any break. Feeding of adult females in winter and summer seasons showed a polymodal pattern with peaks occurring at different hours of the day without break. In yearling males feeding and resting was observed to occur throughout the day during winter season with peaks occurring between 11:00 to 12:00 hours. Yearling females showed continuous feeding throughout the day with peaks in different times in both the seasons. The seasonal distribution of activity patterns of the fawns showed that feeding was slightly more in summer as compared to winter. In time budget, of the expenditure on different activities, resting accounted for 63.77% and feeding 24.70%. In both the seasons resting was the major portion of their activity. Analysis showed that in all age and sex categories of Swamp deer in the Dudhwa NP, resting dominated on all other activities. The observation on endangered Swamp deer indicates synchronization in activity only in the feeding in the morning and evening hours and resting throughout the day.

scholarly journals Technical Note: A mobile sea-going mesocosm system – new opportunities for ocean change research

2012 ◽  
Vol 9 (9) ◽  
pp. 12985-13017 ◽  
Author(s):  
U. Riebesell ◽  
J. Czerny ◽  
K. von Bröckel ◽  
T. Boxhammer ◽  
J. Büdenbender ◽  
...  
Keyword(s):  
Ocean Acidification ◽  
Environmental Changes ◽  
Biogeochemical Cycling ◽  
Single Species ◽  
High Arctic ◽  
Technical Note ◽  
Plankton Community ◽  
Multiple Test ◽  
The Baltic ◽  
June July

Abstract. One of the great challenges in ocean change research is to understand and forecast the effects of environmental changes on pelagic communities and the associated impacts on biogeochemical cycling. Mesocosms, experimental enclosures designed to approximate natural conditions, and in which environmental factors can be manipulated and closely monitored, provide a powerful tool to close the gap between single species laboratory experiments and observational and correlative approaches applied in field surveys. Existing pelagic mesocosm systems are stationary and/or restricted to well-protected waters. To allow mesocosm experimentation in a range of hydrographic conditions and in areas considered most sensitive to ocean change, we developed a mobile, sea-going mesocosm facility, the Kiel Off-Shore Mesocosms for Future Ocean Simulations (KOSMOS). The KOSMOS platform, which can be transported and deployed by mid-sized research vessels, is designed for operation in moored and free-floating mode under low to moderate wave conditions (up to 2.5 m wave heights). It encloses a water column 2 m in diameter and 15 to 25 m deep (~50–75 m3 in volume) without disrupting the vertical structure or disturbing the enclosed plankton community. Several new developments in mesocosm design and operation were implemented to (i) minimize differences in starting conditions between mesocosms, (ii) allow for extended experimental duration, (iii) precisely determine the mesocosm volume, (iv) determine air–sea gas exchange, and (v) perform mass balance calculations. After multiple test runs in the Baltic Sea, which resulted in continuous improvement of the design and handling, the KOSMOS platform successfully completed its first full-scale experiment in the high Arctic off Svalbard (78° 56.2′ N, 11° 53.6′ E) in June/July 2010. The study, which was conducted in the framework of the European Project on Ocean Acidification (EPOCA), focused on the effects of ocean acidification on a natural plankton community and its impacts on biogeochemical cycling and air/sea exchange of climate relevant gases. This manuscript describes the mesocosm hardware, its deployment and handling, CO2 manipulation, sampling and cleaning, including some further modifications conducted based on the experiences gained during this study.

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