Flock size and formation in black-billed gulls
Black-billed gulls (Larus bulleri) depart form breeding colonies and foraging sites in nonrandom, clumped formations (flocks). Flocks leaving colonies were significantly smaller than those leaving foraging sites, in part owing to more "upflights" at foraging sites. When more birds left a colony per unit time, flocks were both larger and more frequent.A causal model was developed based on the assumptions that flock departures from a colony or foraging site result from the effects of social attractions superimposed on otherwise random departure intervals. This model successfully predicted the size distribution of flocks departing from four colonies and foraging sites. In conjunction with the concepts of site tenacity and habituation, the model also permits a causal explanation of differences in the size of flocks departing colonies compared with foraging sites, and of size differences in flocks arising from "upflights" as opposed to the more common "straggling" columnar formations. The model and results are consistent with the hypothesis that colonies function as assembly points that facilitate group foraging.