A new phylogeny and phylogenetic classification for the Canthyloscelidae (Diptera: Psychodomorpha)

2000 ◽  
Vol 78 (6) ◽  
pp. 1067-1077 ◽  
Author(s):  
Dalton de Souza Amorim

A new phylogeny and phylogenetic classification for the Canthyloscelidae (Diptera: Psychodomorpha) is presented. A phylogenetic analysis of the Scatopsoidea is performed. A sister-group relationship between the Canthyloscelidae and Scatopsidae is accepted and the monophyly of the Canthyloscelidae is corroborated, including the genera Exiliscelis, Synneuron, Hyperoscelis, and Canthyloscelis. An earlier phylogenetic analysis of the group is considered, in which Synneuron was accepted as the sister-group of the Scatopsidae and Exiliscelis was considered the sister-group of Synneuron + Scatopsidae. Some apomorphic similarities between the larvae of all genera of Canthyloscelidae, especially the reduction of the head capsule, are considered true synapomorphies. Exiliscelis is considered the sister-group of the rest of the family and is placed in a new subfamily, Exiliscelinae. In the Canthyloscelinae, Synneuron is the sister-group of Hyperoscelis + Canthyloscelis. A phylogenetic classification of the group is proposed. Prohyperoscelis rohdendorfi Kovalev, 1985, from the Middle Jurassic in Russia, is accepted as the sister-group of Canthyloscelis.

Zootaxa ◽  
2021 ◽  
Vol 4992 (1) ◽  
pp. 1-89
Author(s):  
ADRIAN ARDILA-CAMACHO ◽  
CALEB CALIFRE MARTINS ◽  
ULRIKE ASPÖCK ◽  
ATILANO CONTRERAS-RAMOS

Adult external morphology of the extant raptorial Mantispoidea (Insecta: Neuroptera: Mantispidae and Rhachiberothidae) is compared emphasizing the morphology of the subfamily Symphrasinae as a key group to understand the phylogenetic relationships among the members of the superfamily. Plega dactylota Rehn, 1939 is thoroughly characterized in order to exemplify the morphology of the Symphrasinae. Additionally, following a review of the literature and examination of comparative material of Dilaridae, Berothidae, Rhachiberothidae and all Mantispidae subfamilies, a new interpretation of the components of the raptorial apparatus (i.e., head, prothorax, grasping forelegs, as well as integumentary specializations) is presented. Also, wing venation for these groups is reinterpreted, and new homology hypotheses for wing venation are proposed based on tracheation and comparative analyses. Given the high morphological divergence on the genital sclerites within the Mantispoidea, plus the confusing previous usage of neutral terminology and terms referring to appendages across taxonomic and morphological studies, we attempt to standardize, simplify, and situate terminology in an evolutionary context under the “gonocoxite concept” (multi-coxopod hypothesis). The remarkable morphological similarity of the genital sclerites of Symphrasinae and Rhachiberothidae (sensu U. Aspöck & Mansell 1994) with the Nallachinae (Dilaridae) was taken as a starting point to understand the morphology of other Mantispidae subfamilies. Based on these morphological comparisons, we provide a revised phylogenetic analysis of Mantispoidea. This new phylogenetic analysis supports a sister group relationship between the family Rhachiberothidae, comprising Rhachiberothinae and Symphrasinae, and the family Mantispidae, including the subfamily Mantispinae and its sister taxa Drepanicinae and Calomantispinae, which may represent a single subfamily. Based on these analyses, raptorial condition probably evolved a single time in these insects and subsequently became diversified in the two sister clades of the raptorial Mantispoidea.  


2020 ◽  
Vol 96 (2) ◽  
pp. 455-498
Author(s):  
Kai Horst George

Uncovering the systematics of CopepodaHarpacticoida, the second-most abundant component of the meiobenthos after Nematoda, is of major importance for any further research dedicated especially to ecological and biogeographical approaches. Based on the evolution of the podogennontan first swimming leg, a new phylogenetic concept of the Ancorabolidae Sars and Cletodidae T. Scott sensu Por (Copepoda, Harpacticoida) is presented, using morphological characteristics. It confirms the polyphyletic status of the Ancorabolidae and its subfamily Ancorabolinae Sars and the paraphyletic status of the subfamily Laophontodinae Lang. Moreover, it clarifies the phylogenetic relationships of the so far assigned members of the family. An exhaustive phylogenetic analysis was undertaken using 150 morphological characters, resulting in the establishment of a now well-justified monophylum Ancorabolidae. In that context, the Ancorabolus-lineage sensu Conroy-Dalton and Huys is elevated to sub-family rank. Furthermore, the membership of Ancorabolina George in a rearranged monophylum Laophontodinae is confirmed. Conversely, the Ceratonotus-group sensu Conroy-Dalton is transferred from the hitherto Ancorabolinae to the Cletodidae. Within these, the Ceratonotus-group and its hypothesised sister-group Cletodes Brady are combined to form a monophyletic subfamily Cletodinae T. Scott, subfam. nov. Consequently, it was necessary to restructure the Ancorabolidae, Ancorabolinae and Laophontodinae and extend the Cletodidae to include the displacement and exclusion of certain taxa. Moreover, comparison of the Ancorabolidae, Cletodidae, Laophontoidea and other Podogennonta shows that the Ancorabolidae and Cletodidae form sister-groups in a monophylum Cletodoidea Bowman and Abele, which similarly has a sister-group-relationship with the Laophontoidea T. Scott. According to the present study, both taxa constitute a derived monophylum within the Podogennonta Lang.


Pathogens ◽  
2021 ◽  
Vol 10 (1) ◽  
pp. 41
Author(s):  
Marcos Godoy ◽  
Daniel A. Medina ◽  
Rudy Suarez ◽  
Sandro Valenzuela ◽  
Jaime Romero ◽  
...  

Piscine orthoreovirus (PRV) belongs to the family Reoviridae and has been described mainly in association with salmonid infections. The genome of PRV consists of about 23,600 bp, with 10 segments of double-stranded RNA, classified as small (S1 to S4), medium (M1, M2 and M3) and large (L1, L2 and L3); these range approximately from 1000 bp (segment S4) to 4000 bp (segment L1). How the genetic variation among PRV strains affects the virulence for salmonids is still poorly understood. The aim of this study was to describe the molecular phylogeny of PRV based on an extensive sequence analysis of the S1 and M2 segments of PRV available in the GenBank database to date (May 2020). The analysis was extended to include new PRV sequences for S1 and M2 segments. In addition, subgenotype classifications were assigned to previously published unclassified sequences. It was concluded that the phylogenetic trees are consistent with the original classification using the PRV genomic segment S1, which differentiates PRV into two major genotypes, I and II, and each of these into two subgenotypes, designated as Ia and Ib, and IIa and IIb, respectively. Moreover, some clusters of country- and host-specific PRV subgenotypes were observed in the subset of sequences used. This work strengthens the subgenotype classification of PRV based on the S1 segment and can be used to enhance research on the virulence of PRV.


2021 ◽  
Vol 20 (7) ◽  
pp. 911-927
Author(s):  
Lucia Muggia ◽  
Yu Quan ◽  
Cécile Gueidan ◽  
Abdullah M. S. Al-Hatmi ◽  
Martin Grube ◽  
...  

AbstractLichen thalli provide a long-lived and stable habitat for colonization by a wide range of microorganisms. Increased interest in these lichen-associated microbial communities has revealed an impressive diversity of fungi, including several novel lineages which still await formal taxonomic recognition. Among these, members of the Eurotiomycetes and Dothideomycetes usually occur asymptomatically in the lichen thalli, even if they share ancestry with fungi that may be parasitic on their host. Mycelia of the isolates are characterized by melanized cell walls and the fungi display exclusively asexual propagation. Their taxonomic placement requires, therefore, the use of DNA sequence data. Here, we consider recently published sequence data from lichen-associated fungi and characterize and formally describe two new, individually monophyletic lineages at family, genus, and species levels. The Pleostigmataceae fam. nov. and Melanina gen. nov. both comprise rock-inhabiting fungi that associate with epilithic, crust-forming lichens in subalpine habitats. The phylogenetic placement and the monophyly of Pleostigmataceae lack statistical support, but the family was resolved as sister to the order Verrucariales. This family comprises the species Pleostigma alpinum sp. nov., P. frigidum sp. nov., P. jungermannicola, and P. lichenophilum sp. nov. The placement of the genus Melanina is supported as a lineage within the Chaetothyriales. To date, this genus comprises the single species M. gunde-cimermaniae sp. nov. and forms a sister group to a large lineage including Herpotrichiellaceae, Chaetothyriaceae, Cyphellophoraceae, and Trichomeriaceae. The new phylogenetic analysis of the subclass Chaetothyiomycetidae provides new insight into genus and family level delimitation and classification of this ecologically diverse group of fungi.


2016 ◽  
Vol 48 (2) ◽  
pp. 89 ◽  
Author(s):  
Vasily V. Grebennikov

This paper reports discovery of the genus <em>Disphaerona</em> Jordan, 1902 in two widely separated regions in China: in three isolated highlands in Yunnan (the Cang Shan Range, Mt. Jizu and Mt. Haba) and in the Dabie Mountains of Hubei, at the border with Anhui. Till present the genus was known from China only from four specimens of<em> D.</em> <em>chinensis</em> (including its holotype) described from Sichuan in 1995. Sixteen specimens from each of the three newly detected localities in Yunnan were DNA barcoded and the data were made publicly available at dx.doi.org/10.5883/DS-DISPHAE. Temporal phylogenetic analysis revealed a topology suggesting monophyly and recent (0.2–0.4 Ma) origin of the populations on both Mt. Jizu and Mt. Haba. The population on the Cang Shan Range harbours representatives of both clades forming the ingroup, mtDNA of which diverged some 7.9 Ma. The habitus of the holotype of the only nominative Chinese <em>Disphaerona</em> species, <em>D. chinensis,</em> is illustrated, together with habitus and male and female genitalia of specimens newly discovered in both Yunnan and Hubei. All specimens of <em>Disphaerona</em> from Yunnan are seemingly conspecific and are taxonomically assigned to D. chinensis. Two similar males and one sympatric but dissimilar, smaller female from the Dabie Mountains do not have associated DNA barcode data. It remains unknown whether the three Dabie Mountains specimens are conspecific or even congeneric; pending this uncertainty they are tentatively treated as <em>Disphaerona</em> without a Linnaean name. <em>Disphaerona</em> population in the Dabie Mountains, if indeed a member of this hypothesised clade, forms the easternmost record of the genus. Monophyly and sister group of <em>Disphaerona</em> are briefly discussed and both are shown as unknown and unverifiable. A brief overview of the state of phylogenetic knowledge of Anthribidae is given, which is practically non-existing and with neither the family, nor any of its tribes and the vast majority of genera tested as monophyletic.


1997 ◽  
Vol 10 (6) ◽  
pp. 855 ◽  
Author(s):  
Mats H. G. Gustafsson ◽  
Kåre Bremer

The genus Carpodetus from New Zealand, New Guinea, and the Solomon Islands, traditionally has been included in the extremely heterogeneous Saxifragaceae sensu lato, but on account of morphological peculiarities it has sometimes been classified in its own family. On palynological grounds it has been suggested to belong near the Ericales. Parsimony analyses of matrices comprising rbcL sequences of 80 taxa sampled from the entire Asteridae and Rosidae provide support for a sister group relationship between Carpodetus and a clade comprising the closely related Australian genera Abrophyllum and Cuttsia, also formerly placed in Saxifragaceae sensu lato, but recently shown to belong within the order Asterales sensu lato. A morphological comparison between the three interrelated genera is provided. They have in common an indumentum of thick-walled unicellular hairs with warty cuticle, and are also uniform in wood anatomy as well as fruit and seed structure. It is proposed that the family Carpodetaceae be expanded to encompass Abrophyllum and Cuttsia.


Zootaxa ◽  
2005 ◽  
Vol 891 (1) ◽  
pp. 1 ◽  
Author(s):  
Magdalena Szarowska ◽  
Andrzej Falniowski ◽  
FRANK Riedel ◽  
Thomas Wilke

The phylogenetic position of the subfamily Pyrgulinae within the superfamily Rissooidea has been discussed very controversially. Different data sets not only led to different evolutionary scenarios but also to different systematic classifications of the taxon. The present study uses detailed anatomical data for two pyrgulinid taxa, the type species of the subfamily, Pyrgula annulata (Linnaeus, 1767), and the type species of the little known genus Dianella, D. thiesseana (Kobelt, 1878), as well as DNA sequencing data of three gene fragments from representatives of eight rissooidean families to A) infer the phylogenetic position of Pyrgulinae with emphasis on its relationships within the family Hydrobiidae, B) to study the degree of concordance between anatomyand DNAbased phylogenies and C) to trace the evolution of anatomical characters along a multi-gene molecular phylogeny to find the anatomical characters that might be informative for future cladistic analyses. Both anatomical and molecular data sets indicate either a very close or even sister-group relationship of Pyrgulinae and Hydrobiinae. However, there are major conflicts between the two data sets on and above the family level. Notably, Hydrobiidae is not monophyletic in the anatomical analysis. The reconstruction of anatomical character evolution indicates that many of the characters on which the European hydrobioid taxonomy is primarily based upon are problematic. The inability to clearly separate some hydrobiids from other distinct families based on those characters might explain why until only a few years ago, "Hydrobiidae" was a collecting box for numerous rissooidean taxa (mostly species with shells small and lacking any characteristic features). The present study not only stresses the need for comprehensive molecular studies of rissooidean taxa, it also demonstrates that much of the problems surrounding anatomical analyses in rissooidean taxa are due to the lack of comprehensive data for many representatives. In order to aid future comparativeanatomical studies and a better understanding of character evolution in the species-rich family Hydrobiidae, detailed anatomical descriptions for P. annulata and D. thiesseana are provided.Key words: Pyrgulinae, Pyrgula, Dianella, Hydrobiidae, phylogeny, DNA, anatomy, Greece


1998 ◽  
Vol 19 (4) ◽  
pp. 385-405 ◽  
Author(s):  
Van Wallach ◽  
Rainer Günther

AbstractThe internal anatomy of Xenophidion is described and compared with that of members of other snake families. A suite of primitive characters eliminates Xenophidion as a possible member of the Caenophidia; only two characters could conceivably suggest a relationship to the Caenophidia and both are examples of losses and thus of low phylogenetic value in assessing relationships. However, among lower snakes a sister group relationship is demonstrated with the Tropidophiidae of the Neotropical region. Besides possessing nearly identical viscera and topographical arrangement thereof, Xenophidion shares several characters with the Tropidophiidae. A new family is created to contain the genus, the Xenophidiidae. The Xenophidiidae share one synapomorphy with both the Tropidophiidae and Bolyeriidae. Therefore, it is proposed that these three families be united in the superfamily Tropidophioidea. A phylogenetic analysis of 52 characters results in the following preferred hypothesis of relationships: (Boinae, (((Bolyeria, Casarea), (Xenophidion, ((Exiliboa, Ungaliophis), (Trachyboa, Tropidophis)))), Acrochordus)).


2011 ◽  
Vol 279 (1731) ◽  
pp. 1093-1099 ◽  
Author(s):  
Maria Heikkilä ◽  
Lauri Kaila ◽  
Marko Mutanen ◽  
Carlos Peña ◽  
Niklas Wahlberg

Although the taxonomy of the ca 18 000 species of butterflies and skippers is well known, the family-level relationships are still debated. Here, we present, to our knowledge, the most comprehensive phylogenetic analysis of the superfamilies Papilionoidea, Hesperioidea and Hedyloidea to date based on morphological and molecular data. We reconstructed their phylogenetic relationships using parsimony and Bayesian approaches. We estimated times and rates of diversification along lineages in order to reconstruct their evolutionary history. Our results suggest that the butterflies, as traditionally understood, are paraphyletic, with Papilionidae being the sister-group to Hesperioidea, Hedyloidea and all other butterflies. Hence, the families in the current three superfamilies should be placed in a single superfamily Papilionoidea. In addition, we find that Hedylidae is sister to Hesperiidae, and this novel relationship is supported by two morphological characters. The families diverged in the Early Cretaceous but diversified after the Cretaceous–Palaeogene event. The diversification of butterflies is characterized by a slow speciation rate in the lineage leading to Baronia brevicornis , a period of stasis by the skippers after divergence and a burst of diversification in the lineages leading to Nymphalidae, Riodinidae and Lycaenidae.


Zootaxa ◽  
2007 ◽  
Vol 1535 (1) ◽  
pp. 1-92 ◽  
Author(s):  
DALTON DE SOUZA AMORIM ◽  
EIRIK RINDAL

A phylogenetic analysis of the Mycetophiliformia (= Sciaroidea) was performed to determine the relationships among its families and to place the following genera of uncertain position in the system: Heterotricha, Ohakunea, Colonomyia, Freemanomyia, Rhynchoheterotricha, Chiletricha, Afrotricha, Anisotricha, Kenyatricha, Nepaletricha, Sciarosoma, Sciaropota, Insulatricha, Cabamofa, Rogambara, and Starkomyia. Eratomyia n. gen. is described based on a new species from Ecuador. Colonomyia brasiliana sp.n. and Colonomyia freemani sp.n. are described respectively from southern Brazil and Chile. The male of Cabamofa mira Jaschhof is described for the first time. A total of 64 terminal taxa and 137 transformation series (with 202 characters) were included in the data matrix, with a number of new features from thoracic morphology. Willi Hennig’s 1973 system for the higher Bibionomorpha was adopted using the name Mycetophiliformia for the Sciaroidea. The Mycetophiliformia are monophyletic. The family Cecidomyiidae appears as the sister group of the remaining Mycetophiliformia, followed by the Sciaridae. In the preferred topology, the Rangomaramidae appear as the group sister of a clade consisting of (Ditomyiidae + Bolitophilidae + Diadocidiidae + Keroplatidae) and of (Lygistorrhinidae + Mycetophilidae). The topology within the Rangomaramidae is (Chiletrichinae subfam. n. (Heterotrichinae subfam. n. ((Rangomaraminae + Ohakuneinae subfam. n.))). The Chiletrichinae include the genera Kenyatricha, Rhynchoheterotricha, Insulatricha, Chiletricha, and Eratomyia n. gen. Heterotrichinae and Rangomaraminae are monotypic. The subfamily Ohakuneinae includes Ohakunea, Colonomyia, Cabamofa, and Rogambara. The positions of Freemanomyia, Loicia, Taxicnemis, Sciaropota, Starkomyia, Anisotricha, Nepaletricha, and Sciarosoma are considered. Afrotricha might belong to the Sciaridae. The similarities used by many authors to gather the Sciaridae and Mycetophilidae in a clade are shown to be a combination of plesiomorphies and homoplasies.


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