Prediction of stand susceptibility to feeding damage by red squirrels in young lodgepole pine
This paper provides a forecast model to predict when and where significant feeding damage by red squirrels (Tamiasciurushudsonicus Erxleben) will occur in managed stands of lodgepole pine (Pinuscontorta Dougl. var. latifolia Engelm.). Information from 51 managed stands (average DBH > 6.0 cm) in the interior of British Columbia and from past squirrel population studies was used to formulate the model. Incidence of damage was significantly greater in stands originating from wildfire than from harvesting. Stands within the Montane Spruce biogeoclimatic zone had the highest levels of damage of the five zones sampled. There was no relationship between damage incidence and average stand diameter, area of managed stand, or site class. In a subset of stands where understory shrub data were available, shrub cover was positively related to incidence of damage. Major factors in the forecast model that influence stand susceptibility include (i) stand origin and proximity to mature timber; (ii) frequency of cone crops leading to squirrel population increases in juvenile pine; and (iii) shrub cover, which may provide security from predators. Additional potential factors include (iv) switching of predators from squirrels to snow-shoe hares (Lepusamericanus Erxleben) during peak years of the hare cycle and (v) fertilization. A decisionmaking profile is outlined for prespacing surveys to identify susceptible stands.