Top pruning Douglas-fir seedlings: morphology, physiology, and field performance
Seedlings from nine Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) seed sources were top pruned at six nurseries in the Pacific Northwest and then planted at field sites in their own seed zones and on one common site. Seedlings pruned tall (25 cm) and early in the growing season flushed again about 5 weeks after pruning and set bud 2 weeks later than unpruned controls. Nursery yield (number of shippable seedlings) was the same for tall–early pruned and control seedlings; however, pruning increased the number of seedlings with multiple leaders from 10 to 38%. Pruned seedlings were smaller than unpruned seedlings in height, stem diameter, bud length, and root and shoot weights at the time of field planting. First-year field survival and growth were the same for pruned and unpruned seedlings at all planting sites. Second-year field growth of pruned seedlings was generally greater than that of unpruned seedlings, and frequency of multiple leaders was reduced. However, pruned seedlings were still shorter than unpruned ones after 2 years. Field growth was greater when seedlings were pruned early rather than late (end of growing season) and tall rather than short (15 cm). Late–short pruning yielded shorter seedlings with smaller terminal buds and should be avoided. If pruning benefits nurseries by expediting grading, handling, and planting and, perhaps, enhancing growth later in the field, it should be continued as a cultural practice. However, longer-term measurements are needed for assessing the full effect of pruning on seedling growth.