Changes in litter and dead wood loads following tree death beneath subalpine conifer species in northern Colorado

2011 ◽  
Vol 41 (2) ◽  
pp. 331-340 ◽  
Author(s):  
Christof Bigler ◽  
Thomas T. Veblen
Keyword(s):  
Pinus Contorta ◽  
Tree Mortality ◽  
Dead Wood ◽  
Fire Behavior ◽  
Subalpine Forest ◽  
Abies Lasiocarpa ◽  
Climate Trends ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Northern Colorado

Litter and dead wood affect important processes in forest ecosystems such as nutrient and carbon cycling and are key influences on biodiversity and fire behavior. Increased tree mortality rates in western North America associated with climate trends and increased bark beetle activity highlight the need to better understand the dynamics of litter and dead wood following tree death. For eight old-growth stands in a subalpine forest landscape in northern Colorado (USA), we compared litter and dead wood loads beneath more than 200 dead and live Engelmann spruce (Picea engelmannii Parry ex Engelm.), subalpine fir (Abies lasiocarpa (Hook.) Nutt.), and lodgepole pine (Pinus contorta Douglas ex Loudon). The dynamics of litter and dead wood were analyzed using chronosequences of tree death dates over >100 years that we determined from tree rings. Immediately following tree death, high loads of litter accumulated, particularly for the biggest spruces, which accumulated 10 times more litter than live spruces (five times more for fir, two times more for pine). We estimated a higher decay rate of litter for spruce (half-life of four years) than for pine (15 years) and fir (19 years). The accumulation rates for dead wood following tree death were highly variable among trees, but maximum accumulation was attained during the first 50–60 years.

Dynamics of an old-growth, fire-initiated, subalpine forest in southern interior British Columbia: tree size, age, and spatial structure

2002 ◽  
Vol 32 (11) ◽  
pp. 1935-1946 ◽  
Author(s):  
Joseph A Antos ◽  
Roberta Parish
Keyword(s):  
Pinus Contorta ◽  
Subalpine Forest ◽  
Abies Lasiocarpa ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Seedling Bank ◽  
Dendrochronological Analysis ◽  
Canopy Position ◽  
Forest Regions ◽  
Almost All

We used dendrochronological analysis of over 2000 trees in four 50 × 50 m plots to reconstruct the history and dynamics of a 330-year-old, fire-initiated spruce-fir forest. All lodgepole pine (Pinus contorta Dougl. ex Loud.), half of the canopy Engelmann spruce (Picea engelmannii Parry ex Engelm.), but less than 10% of the canopy subalpine fir (Abies lasiocarpa (Hook.) Nutt.) dated from the first 50 years of stand development. Tree-ring patterns of individual surviving trees showed no evidence of disturbance during the first 200 years after stand initiation; subsequently, episodes of disturbance are indicated by periods of release in understory fir. Although many fir owe their canopy position to release after disturbance, few canopy fir in the current stand established in response to either the stand-initiating event or subsequent partial disturbances. A seedling bank of long-lived fir appears critical to the dynamics of this forest. In contrast, establishment of almost all canopy spruce can be related to disturbance. This stand, although fire initiated, was structured primarily by a combination of partial disturbances and autogenic processes. We suspect that most old, fire-initiated stands in many forest regions are similarly structured and emphasize that the contribution of partial disturbances and autogenic processes should be fully assessed when examining their dynamics or managing such forests.

Inonotus tomentosus and the dynamics of unmanaged and partial-cut wet sub-boreal spruce–fir forests

2007 ◽  
Vol 37 (12) ◽  
pp. 2663-2676 ◽  
Author(s):  
J. E. (Ted) Newbery ◽  
Kathy J. Lewis ◽  
Michael B. Walters
Keyword(s):  
Picea Glauca ◽  
Dead Wood ◽  
Basal Area ◽  
Tree Density ◽  
Small Scale ◽  
Abies Lasiocarpa ◽  
Partial Cutting ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Live Tree

For wet sub-boreal spruce–fir forests (white spruce ( Picea glauca (Moench) Voss) × Engelmann spruce ( Picea engelmannii Parry ex Engelm.) – subalpine fir ( Abies lasiocarpa (Hook.) Nutt.)) in east-central British Columbia, we asked (i) do compositional and structural dynamics differ for unmanaged (UN) and partial-cut (PC) (50% removal 45 years before measurement) forests and (ii) how does Inonotus tomentosus Fr. (Teng) affect these dynamics? Inonotus tomentosus infected stands had 17% less spruce basal area (P = 0.059) than uninfected stands, but PC did not exacerbate I. tomentosus effects. PC and UN had similar live tree density, but UN had lower dead tree density. In all stands, snag longevity was typically <32 years, and ~40 years was required for dead wood to reach decay stage 3 or greater. UN was characterized by variable severity disturbances averaging ~8% of the canopy per decade. Management implications include the following: (i) harvest systems designed to emulate small-scale disturbance could remove trees at 8% of the canopy per decade, varied spatiotemporally, (ii) emulating dead wood abundance with partial cutting may be difficult given the impacts of partial cutting on dead wood abundance, and (iii) forests with moderate levels of I. tomentosus should not respond differently to harvesting than uninfected forests and thus require no special management.

Space–time modelling of lightning-caused ignitions in the Blue Mountains, Oregon

2001 ◽  
Vol 31 (9) ◽  
pp. 1579-1593 ◽  
Author(s):  
Carlos Díaz-Avalos ◽  
David L Peterson ◽  
Ernesto Alvarado ◽  
Sue A Ferguson ◽  
Julian E Besag
Keyword(s):  
Pinus Contorta ◽  
Vegetation Type ◽  
Abies Lasiocarpa ◽  
Time Data ◽  
Subalpine Fir ◽  
Blue Mountains ◽  
Engelmann Spruce ◽  
Covariate Information ◽  
A Value ◽  
Starting Location

Generalized linear mixed models (GLMM) were used to study the effect of vegetation cover, elevation, slope, and precipitation on the probability of ignition in the Blue Mountains, Oregon, and to estimate the probability of ignition occurrence at different locations in space and in time. Data on starting location of lightning-caused ignitions in the Blue Mountains between April 1986 and September 1993 constituted the base for the analysis. The study area was divided into a pixel–time array. For each pixel–time location we associated a value of 1 if at least one ignition occurred and 0 otherwise. Covariate information for each pixel was obtained using a geographic information system. The GLMMs were fitted in a Bayesian framework. Higher ignition probabilities were associated with the following cover types: subalpine herbaceous, alpine tundra, lodgepole pine (Pinus contorta Dougl. ex Loud.), whitebark pine (Pinus albicaulis Engelm.), Engelmann spruce (Picea engelmannii Parry ex Engelm.), subalpine fir (Abies lasiocarpa (Hook.) Nutt.), and grand fir (Abies grandis (Dougl.) Lindl.). Within each vegetation type, higher ignition probabilities occurred at lower elevations. Additionally, ignition probabilities are lower in the northern and southern extremes of the Blue Mountains. The GLMM procedure used here is suitable for analysing ignition occurrence in other forested regions where probabilities of ignition are highly variable because of a spatially complex biophysical environment.

Black Bear Damage to Thinned Timber Stands in Northwest Montana

1989 ◽  
Vol 4 (1) ◽  
pp. 10-13 ◽  
Author(s):  
Andrew C. Mason ◽  
David L. Adams
Keyword(s):  
Pinus Contorta ◽  
Black Bear ◽  
Tsuga Heterophylla ◽  
Thuja Plicata ◽  
Abies Lasiocarpa ◽  
Pinus Monticola ◽  
Larix Occidentalis ◽  
Western Larch ◽  
Engelmann Spruce ◽  
Northwest Montana

Abstract Bear damage was at least five times higher in thinned blocks than in adjacent unthinned blocks of western larch (Larix occidentalis), lodgepole pine (Pinus contorta), and Engelmann spruce (Picea engelmannii) on the Kootenai National Forest in northwest Montana. Western larch suffered the greatest damage (63% of all trees damaged and 92% of the trees killed). Damaged larch ranged from 4 to 13 in. dbh; the 4 to 8-in. dbh class accounted for 85% of the damage. Douglas-fir (Pseudotsuga menziesii), western redcedar (Thuja plicata), subalpine fir (Abies lasiocarpa), western white pine (Pinus monticola), and western hemlock (Tsuga heterophylla) were not damaged. Stand projections showed up to a 17% reduction in board-foot yield from bear damage, after 50 years, compared with hypothetical undamaged stands. West. J. Appl. For. 4(1):10-13, January 1989.

scholarly journals Natural and Advance Regeneration of Engelmann Spruce and Subalpine Fir Compared 21 Years After Site Treatment

1980 ◽  
Vol 56 (2) ◽  
pp. 55-57 ◽  
Author(s):  
L. J. Herring ◽  
R. G. McMinn
Keyword(s):  
Natural Regeneration ◽  
Mineral Soil ◽  
Poor Performance ◽  
Abies Lasiocarpa ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Advance Regeneration ◽  
Residual Tree ◽  
Mean Current ◽  
Picea Engelmanni

The mean height of Engelmann spruce (Picea engelmanni Parry) advance growth 21 years after release by overstorey harvesting and residual tree felling, was eight times that of natural regeneration established following brush blade scarification. Subalpine fir (Abies lasiocarpa (Hook.) Nutt.) advance growth was nine times taller than natural regeneration established on scarified soil. Mean current annual height increment of Engelmann spruce and subalpine fir advance growth was 39 and 34 cm, respectively, compared with only 7 cm for natural regeneration on scarified soil. The performance gap does not appear to be narrowing. The poor performance of natural regeneration on mineral soil exposed by blade scarification is attributed to removal of organic and top mineral soil horizons beyond the immediate reach of seedlings. These soil layers remained available to the advance growth. Consideration should be given to preserving advance growth when scarification may be inappropriate.

Segregation of allozyme loci in megagametophytes of Engelmann spruce and subalpine fir

Genome ◽  
1988 ◽  
Vol 30 (2) ◽  
pp. 103-107 ◽  
Author(s):  
Kathleen L. Shea
Keyword(s):  
Natural Populations ◽  
Segregation Ratio ◽  
Colorado Front Range ◽  
Abies Lasiocarpa ◽  
Front Range ◽  
Subalpine Fir ◽  
Polymorphic Loci ◽  
Engelmann Spruce ◽  
Low Levels ◽  
Allozyme Loci

Segregation ratios and linkage of 10 allozyme loci were examined in haploid megagametophytes obtained from natural populations of Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) in the Colorado Front Range. For data pooled over trees, the 1:1 segregation ratio expected at Mendelian loci was obtained for five polymorphic loci in 32 Engelmann spruce trees and for seven polymorphic loci in 40 subalpine fir trees. The Gdh and Idh loci in spruce were very tightly linked: no recombinants were detected among 60 megagametophytes of trees heterozygous for both loci. In fir only the Aco and Pgm-1 loci were linked, with an estimated recombination rate of 0.317 ± 0.073. The low levels of among-tree heterogeneity and of segregation distortion found in these populations suggest that reliable estimates of both genetic variation and outcrossing rates can be obtained using allozyme data from these wind-pollinated species.Key words: segregation, linkage, allozymes, Engelmann spruce, subalpine fir.

scholarly journals Hypogeous Fungi Occurrence, Distribution and Mycorrhizal Hosts in Grand Teton National Park and John D. Rockefeller, Jr. Memorial Parkway

1990 ◽  
Vol 14 ◽  
pp. 13-14
Author(s):  
Steven Miller
Keyword(s):  
Populus Tremuloides ◽  
National Park ◽  
Pinus Contorta ◽  
Quaking Aspen ◽  
Abies Lasiocarpa ◽  
Hypogeous Fungi ◽  
Grand Teton National Park ◽  
Subalpine Fir ◽  
Field Season ◽  
Speckled Alder

The 1990 field season constituted the last of a three year study to survey the hypogeous fungi of Grand Teton National Park and the Greater Yellowstone ecosystem. The objectives were to: 1. collect and identify hypogeous fungi found in association with ectomycorrhizal tree hosts such as lodgepole pine (Pinus contorta), subalpine fir (Abies lasiocarpa), Douglas-frr (Pseudotsuga menziesii), quaking aspen (Populus tremuloides), and speckled alder (Alnus tenuifolia), and several species of willow (Salix sp.) throughout the area; and 2. to gain an initial understanding of the importance of these fungi as food for small mammals.

scholarly journals Captures and Recaptures of Small Mammals to Assess Responses to Fire in a Coniferous Forest in the Greater Yellowstone Area

1998 ◽  
Vol 22 ◽  
pp. 71-77
Author(s):  
N. Stanton ◽  
R. Seville ◽  
S. Buskirk ◽  
S. Miller ◽  
D. Spildie ◽  
...  
Keyword(s):  
Small Mammals ◽  
Pinus Contorta ◽  
Deer Mouse ◽  
Coniferous Forest ◽  
Common Species ◽  
Abies Lasiocarpa ◽  
Engelmann Spruce ◽  
Natural Fires ◽  
Greater Yellowstone Area ◽  
Greater Yellowstone

Natural fires are common in coniferous forests in the Rocky Mountains, and one of the largest fires in recent history occurred in the Greater Yellowstone Area (GYA) in 1988 when over a million acres of lodgepole pine (Pinus contorta), subalpine fir (Abies lasiocarpa) and Engelmann spruce (Picea engelmannii) burned. In the summers of 1989, 1990 and 1991 and again in 1997 and 1998, we trapped small mammals in two burned and two adjacent unburned forests in the Huckleberry Mountain fire in the Rockefeller Memorial Parkway, 0.5 km north of Grand Teton National Park (GTNP). Here we report on the captures and recaptures of the two most common species of small mammals, the deer mouse (Peromyscus maniculatus) and the southern red-backed vole (Clethrionomys gapperi); and analyze retrapping frequency for each species in the burned and unburned forest. Our intent was to test the hypothesis that the probability of recapture is the same for both species in burned and unburned habitats. These capture/recapture data will be used by other co-investigators in additional publications to report on estimated population sizes and microhabitat associations.

Microhabitat comparisons of transpiration and photosynthesis in three subalpine conifers

1988 ◽  
Vol 66 (5) ◽  
pp. 963-969 ◽  
Author(s):  
Gregory A. Carter ◽  
William K. Smith
Keyword(s):  
Water Use Efficiency ◽  
Water Use ◽  
Pinus Contorta ◽  
Photosynthetic Performance ◽  
Leaf Conductance ◽  
Forest Understory ◽  
Subalpine Forest ◽  
Abies Lasiocarpa ◽  
Use Efficiency ◽  
Central Rocky Mountains

Differences in water and photosynthetic relations were compared for three codominant conifers (Engelmann spruce (Picea engelmaniï), subalpine fir (Abies lasiocarpa), and lodgepole pine (Pinus contorta) at microhabitats within a subalpine forest (central Rocky Mountains, U.S.A.) that were considered representative of different successional stages. Diumal measurements of photosynthesis, leaf conductance, and transpiration were taken at microhabitats considered early-successional (open), intermediate (forest gap), and late-successional (forest understory) environments to evaluate possible influences of gas-exchange physiology in observed distributional and successional patterns. Pine had greater water-use efficiency (photosynthesis/transpiration) in early- versus late-successional environments, primarily as a result of a lower leaf conductance and transpiration. Photosynthetic performance was similar among all three species at each respective microhabitat and increased as the openness of the microhabitat increased. Greater water-use efficiency may significantly improve the growth of pine over spruce and fir on more open, drier sites at lower elevation. Higher transpiration in spruce and fir may limit these species to higher elevation sites, to understory sites at middle elevations, and to moister open sites at lower elevations (e.g., riparian sites).

Effect of gap size on litter decomposition and soil nitrate concentrations in a high-elevation spruce–fir forest

2003 ◽  
Vol 33 (11) ◽  
pp. 2210-2220 ◽  
Author(s):  
Cindy E Prescott ◽  
Graeme D Hope ◽  
Leandra L Blevins
Keyword(s):  
Forest Floor ◽  
Mineral Soil ◽  
High Elevation ◽  
N Availability ◽  
Abies Lasiocarpa ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Nitrate Losses ◽  
Nitrate Concentrations ◽  
High Elevation Forest

Possible mitigation of nitrate losses associated with clearcuts through harvesting smaller gaps was tested in a high-elevation forest of Engelmann spruce (Picea engelmannii Parry ex Engelm.) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.). We measured concentrations of ammonium and nitrate after 6-week buried bag incubations of forest floor and mineral soil samples in replicated plots of uncut forest and gaps of 10, 1.0, and 0.1 ha and single-tree removal for 7 years after harvest. Nitrate concentrations in forest floor and mineral soil were elevated 3–7 years after harvesting in gaps of 0.1 ha and larger. Removal of the same proportion of trees as single trees did not result in increased nitrate concentrations, suggesting that nitrate losses could be reduced by harvesting single trees rather than creating gaps. Greater N availability was not associated with faster rates of decomposition of litter and forest floor, which were similar in gaps of all sizes (0–10 ha). Reciprocal transplant of forest floor and soil from the 10-ha gaps and the uncut forests indicated that changes in the nature of the forest floor or soil following harvest had a greater influence on nitrate concentrations than the changes in environmental conditions in the gaps.

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