Inonotus tomentosus and the dynamics of unmanaged and partial-cut wet sub-boreal spruce–fir forests

2007 ◽  
Vol 37 (12) ◽  
pp. 2663-2676 ◽  
Author(s):  
J. E. (Ted) Newbery ◽  
Kathy J. Lewis ◽  
Michael B. Walters
Keyword(s):  
Picea Glauca ◽  
Dead Wood ◽  
Basal Area ◽  
Tree Density ◽  
Small Scale ◽  
Abies Lasiocarpa ◽  
Partial Cutting ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Live Tree

For wet sub-boreal spruce–fir forests (white spruce ( Picea glauca (Moench) Voss) × Engelmann spruce ( Picea engelmannii Parry ex Engelm.) – subalpine fir ( Abies lasiocarpa (Hook.) Nutt.)) in east-central British Columbia, we asked (i) do compositional and structural dynamics differ for unmanaged (UN) and partial-cut (PC) (50% removal 45 years before measurement) forests and (ii) how does Inonotus tomentosus Fr. (Teng) affect these dynamics? Inonotus tomentosus infected stands had 17% less spruce basal area (P = 0.059) than uninfected stands, but PC did not exacerbate I. tomentosus effects. PC and UN had similar live tree density, but UN had lower dead tree density. In all stands, snag longevity was typically <32 years, and ~40 years was required for dead wood to reach decay stage 3 or greater. UN was characterized by variable severity disturbances averaging ~8% of the canopy per decade. Management implications include the following: (i) harvest systems designed to emulate small-scale disturbance could remove trees at 8% of the canopy per decade, varied spatiotemporally, (ii) emulating dead wood abundance with partial cutting may be difficult given the impacts of partial cutting on dead wood abundance, and (iii) forests with moderate levels of I. tomentosus should not respond differently to harvesting than uninfected forests and thus require no special management.

Windthrow following four harvest treatments in an Engelmann spruce - subalpine fir forest in southern interior British Columbia, Canada

1999 ◽  
Vol 29 (10) ◽  
pp. 1547-1556 ◽  
Author(s):  
David J Huggard ◽  
Walt Klenner ◽  
Alan Vyse
Keyword(s):  
Large Scale ◽  
Basal Area ◽  
Random Trees ◽  
Abies Lasiocarpa ◽  
Ecological Processes ◽  
Individual Tree ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Lower Height

We used transect surveys at a large-scale experimental site at Sicamous Creek, B.C., to measure the effects of five treatments on windthrow: 10-ha clearcuts, arrays of 1-ha patch cuts, arrays of 0.1-ha patch cuts, individual-tree selection cuts, and uncut controls. We also examined edge effects and conditions predisposing trees to windthrow. Windthrow of subalpine fir (Abies lasiocarpa (Hook.) Nutt.) in the 2.7 years following harvesting increased from 0.6% of basal area per year in uncut forest to 0.8-1.8% per year in harvested treatments, with highest rates in individual tree selection units and lowest rates in 0.1-ha patch-cut arrays. Engelmann spruce (Picea engelmannii Parry ex Engelm.) showed similar patterns of windthrow but lower rates (0.2-0.7% of basal area per year in harvested treatments). Windthrow was concentrated near north and east edges of 1-ha and 10-ha openings but was dispersed throughout the more uniform treatments. Windthrown trees did not differ from random trees in diameter but had lower height/diameter ratios, probably reflecting the greater windthrow observed in subxeric sites on complex, elevated topography. The rates and distribution of windthrow in different harvest treatments have implications for ecological processes, salvage, long-term windthrow potential, and mitigation possibilities.

Changes in litter and dead wood loads following tree death beneath subalpine conifer species in northern Colorado

2011 ◽  
Vol 41 (2) ◽  
pp. 331-340 ◽  
Author(s):  
Christof Bigler ◽  
Thomas T. Veblen
Keyword(s):  
Pinus Contorta ◽  
Tree Mortality ◽  
Dead Wood ◽  
Fire Behavior ◽  
Subalpine Forest ◽  
Abies Lasiocarpa ◽  
Climate Trends ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Northern Colorado

Litter and dead wood affect important processes in forest ecosystems such as nutrient and carbon cycling and are key influences on biodiversity and fire behavior. Increased tree mortality rates in western North America associated with climate trends and increased bark beetle activity highlight the need to better understand the dynamics of litter and dead wood following tree death. For eight old-growth stands in a subalpine forest landscape in northern Colorado (USA), we compared litter and dead wood loads beneath more than 200 dead and live Engelmann spruce (Picea engelmannii Parry ex Engelm.), subalpine fir (Abies lasiocarpa (Hook.) Nutt.), and lodgepole pine (Pinus contorta Douglas ex Loudon). The dynamics of litter and dead wood were analyzed using chronosequences of tree death dates over >100 years that we determined from tree rings. Immediately following tree death, high loads of litter accumulated, particularly for the biggest spruces, which accumulated 10 times more litter than live spruces (five times more for fir, two times more for pine). We estimated a higher decay rate of litter for spruce (half-life of four years) than for pine (15 years) and fir (19 years). The accumulation rates for dead wood following tree death were highly variable among trees, but maximum accumulation was attained during the first 50–60 years.

Short-term impacts of partial cutting on lichen retention and canopy microclimate in an Engelmann spruce – subalpine fir forest in north-central British Columbia

2003 ◽  
Vol 33 (5) ◽  
pp. 830-841 ◽  
Author(s):  
Darwyn Coxson ◽  
Susan Stevenson ◽  
Jocelyn Campbell
Keyword(s):  
British Columbia ◽  
Sampling Techniques ◽  
Abies Lasiocarpa ◽  
Short Term ◽  
Partial Cutting ◽  
North Central ◽  
Subalpine Fir ◽  
Single Tree ◽  
Engelmann Spruce ◽  
Single Tree Selection

The retention of canopy lichens (Alectoria, Bryoria, and foliose) in group selection (GS) and single-tree selection (STS) partial cuts (30% removal) was studied within a subalpine spruce–fir forest (Picea englemannii Parry ex Engelm. – Abies lasiocarpa (Hook.) Nutt.) in north-central British Columbia. Baseline canopy lichen loading (preharvest and immediate postharvest) was compared with lichen loading 2 years after harvesting, using both ground- and canopy-based sampling techniques. Additionally, canopy microclimate was compared between GS and adjacent stand areas with no harvest treatment (NT). Partial cutting did not have a significant effect on lichen loading in residual trees, although a trend of declining lichen abundance was observed in each of the GS, STS, and NT areas. Partitioning of lichen abundance data showed significant postharvest declines for upper canopy STS Bryoria and smaller, but significant, changes in foliose lichen abundance in the NT and STS areas. No significant changes in canopy microclimate were observed between treatments, with the exception of south-facing aspects in GS trees, which showed reduced duration of lichen hydration. Taken together, these factors suggest that although the placement of Bryoria in residual trees may shift towards lower canopy exposures, partial-cut harvest treatments can maintain short-term lichen loading and associated ecosystem values (e.g., caribou forage) in subalpine spruce–fir stands.

scholarly journals Rocky pine forests in the High Coast Region in Sweden: structure, dynamics and history

2020 ◽  
Vol 38 ◽  
pp. 101-130
Author(s):  
Jennie Sandström ◽  
Mattias Edman ◽  
Bengt Gunnar Jonsson
Keyword(s):  
Human Impact ◽  
Fire History ◽  
Dead Wood ◽  
Forest Type ◽  
Basal Area ◽  
Pine Forests ◽  
Small Scale ◽  
Natural Dynamics ◽  
Almost All ◽  
High Degree

Almost all forests in Sweden are managed and only a small fraction are considered natural. One exception is low productive forests where, due to their limited economical value, natural dynamics still dominate. One example is the Scots pine (Pinus sylvestris L.) forests occurring on rocky and nutrient-poor hilltops. Although these forests represent a regionally common forest type with a high degree of naturalness, their dynamics, structure and history are poorly known. We investigated the structure, human impact and fire history in eight rocky pine forests in the High Coast Area in eastern Sweden, initially identified as good representatives of this forest type. This was done by sampling and measuring tree sizes, -ages, fire-scarred trees, as well as dead wood volumes and quality along three transects at each site. The structure was diverse with a sparse layer of trees (basal area 9 m2 and 640 trees larger than 10 cm ha-1) in various sizes and ages; 13 trees ha-1 were more than 300 years old. Dead wood (DW), snags and logs in all stages of decay, was present and although the actual DW (pine) volume (4.4 m3 ha-1) and number of units (53 ha-1) was low, the DW share of total wood volume was 18% on average. Dead wood can be present for several centuries after death; we found examples of both snags and logs that had been dead more than 300 years. Frequent fires have occurred, with an average cycle of 40 years between fires. Most fires occurred between 1500-1900 and many of them (13) during the 1600s. However, fires were probably small since most fire years were only represented at one site and often only in one or a few samples. The rocky pine forests in the High Coast Area are representative of undisturbed forests with low human impact, exhibiting old-growth characteristics and are valuable habitats for organisms connected to sun-exposed DW. Management of protected rocky pine forests may well include small-scale restoration fires and the limited DW volumes should be protected.

scholarly journals Natural and Advance Regeneration of Engelmann Spruce and Subalpine Fir Compared 21 Years After Site Treatment

1980 ◽  
Vol 56 (2) ◽  
pp. 55-57 ◽  
Author(s):  
L. J. Herring ◽  
R. G. McMinn
Keyword(s):  
Natural Regeneration ◽  
Mineral Soil ◽  
Poor Performance ◽  
Abies Lasiocarpa ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Advance Regeneration ◽  
Residual Tree ◽  
Mean Current ◽  
Picea Engelmanni

The mean height of Engelmann spruce (Picea engelmanni Parry) advance growth 21 years after release by overstorey harvesting and residual tree felling, was eight times that of natural regeneration established following brush blade scarification. Subalpine fir (Abies lasiocarpa (Hook.) Nutt.) advance growth was nine times taller than natural regeneration established on scarified soil. Mean current annual height increment of Engelmann spruce and subalpine fir advance growth was 39 and 34 cm, respectively, compared with only 7 cm for natural regeneration on scarified soil. The performance gap does not appear to be narrowing. The poor performance of natural regeneration on mineral soil exposed by blade scarification is attributed to removal of organic and top mineral soil horizons beyond the immediate reach of seedlings. These soil layers remained available to the advance growth. Consideration should be given to preserving advance growth when scarification may be inappropriate.

Temporal development of decaying log habitats in wet spruce–fir stands in east-central British Columbia

2005 ◽  
Vol 35 (12) ◽  
pp. 2841-2850 ◽  
Author(s):  
S Craig DeLong ◽  
Lori D Daniels ◽  
Ben Heemskerk ◽  
Ken Olaf Storaunet
Keyword(s):  
British Columbia ◽  
Picea Glauca ◽  
Model Development ◽  
Wildlife Habitat ◽  
Abies Lasiocarpa ◽  
Time Since Death ◽  
Morphological Attributes ◽  
East Central ◽  
Subalpine Fir ◽  
Advanced Stages

Time since death and time since fall were estimated for hybrid spruce (Picea glauca (Moench) Voss × Picea engelmannii Parry ex Engelm.) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.) logs to quantify temporal changes in log decay and habitat quality in east-central British Columbia. We sampled 136 logs (72 spruce and 64 fir) for species, size, and morphological attributes and used dendroecological techniques to estimate year of death (n = 97) and fall (n =  22). Time since death and time since fall of spruce and fir were similar in decay classes 1 and 2; fir was older than spruce in more advanced stages of decay. Discriminant analysis based on time since fall correctly classified logs into four decay classes for 67% and 80% of spruce and fir, respectively. Function as wildlife habitat changed significantly as logs decayed. Logs served as elevated runways for approximately 15 years and then increased in value as habitat for invertebrates and insectivores as wood softened and vegetation established. Concealed spaces increased as the wood decomposed but decreased when logs collapsed. We conclude that decay classes represent biologically and statistically significant stages of log decomposition that are relevant to wildlife habitat and therefore provide a useful construct for model development and field interpretation..

Segregation of allozyme loci in megagametophytes of Engelmann spruce and subalpine fir

Genome ◽  
1988 ◽  
Vol 30 (2) ◽  
pp. 103-107 ◽  
Author(s):  
Kathleen L. Shea
Keyword(s):  
Natural Populations ◽  
Segregation Ratio ◽  
Colorado Front Range ◽  
Abies Lasiocarpa ◽  
Front Range ◽  
Subalpine Fir ◽  
Polymorphic Loci ◽  
Engelmann Spruce ◽  
Low Levels ◽  
Allozyme Loci

Segregation ratios and linkage of 10 allozyme loci were examined in haploid megagametophytes obtained from natural populations of Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) in the Colorado Front Range. For data pooled over trees, the 1:1 segregation ratio expected at Mendelian loci was obtained for five polymorphic loci in 32 Engelmann spruce trees and for seven polymorphic loci in 40 subalpine fir trees. The Gdh and Idh loci in spruce were very tightly linked: no recombinants were detected among 60 megagametophytes of trees heterozygous for both loci. In fir only the Aco and Pgm-1 loci were linked, with an estimated recombination rate of 0.317 ± 0.073. The low levels of among-tree heterogeneity and of segregation distortion found in these populations suggest that reliable estimates of both genetic variation and outcrossing rates can be obtained using allozyme data from these wind-pollinated species.Key words: segregation, linkage, allozymes, Engelmann spruce, subalpine fir.

Stem- and stand-level growth and mortality following partial cutting in eastern boreal poplar – white spruce stands

2019 ◽  
Vol 49 (5) ◽  
pp. 463-470 ◽  
Author(s):  
Suzanne Brais ◽  
Brian D. Harvey ◽  
Arun K. Bose
Keyword(s):  
Growth Rates ◽  
Picea Glauca ◽  
Basal Area ◽  
Natural Disturbance ◽  
White Spruce ◽  
Primary Objective ◽  
Post Treatment ◽  
Tree Level ◽  
Growth Responses ◽  
Partial Cutting

Variable retention (VR) and partial cutting are both considered important silvicultural tools of natural disturbance or ecosystem based forest management approaches. Partial harvesting differs from VR in that post-treatment growth responses and stand regeneration are the primary objective rather than the maintenance of biodiversity. This partial cutting study is undertaken in mixed poplar (Populus spp.) – white spruce (Picea glauca (Moench) Voss) stands in the eastern Canadian boreal mixedwood forest. It compares, at the tree level, absolute growth rates (AGR) and relative growth rates (RGR) of basal area (BA) and stem survival; and at the stand level, it also compares absolute BA growth, mortality, and sapling density 10 years following treatment. The completely randomized experiment was established with four intensities of partial cutting (0, 50%, 65%, and 100% of poplar BA). All partial cutting intensities had a significant and similar positive effect on AGR of residual spruce stems. Complete poplar removal resulted not only in the highest increase in RGR of suppressed and intermediate spruce stems, but also in higher spruce mortality. Removal of 50% of the initial poplar stand BA provided the best trade-off between positive residual stem growth of spruce and poplar and limited post-treatment mortality.

Effect of gap size on litter decomposition and soil nitrate concentrations in a high-elevation spruce–fir forest

2003 ◽  
Vol 33 (11) ◽  
pp. 2210-2220 ◽  
Author(s):  
Cindy E Prescott ◽  
Graeme D Hope ◽  
Leandra L Blevins
Keyword(s):  
Forest Floor ◽  
Mineral Soil ◽  
High Elevation ◽  
N Availability ◽  
Abies Lasiocarpa ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Nitrate Losses ◽  
Nitrate Concentrations ◽  
High Elevation Forest

Possible mitigation of nitrate losses associated with clearcuts through harvesting smaller gaps was tested in a high-elevation forest of Engelmann spruce (Picea engelmannii Parry ex Engelm.) and subalpine fir (Abies lasiocarpa (Hook.) Nutt.). We measured concentrations of ammonium and nitrate after 6-week buried bag incubations of forest floor and mineral soil samples in replicated plots of uncut forest and gaps of 10, 1.0, and 0.1 ha and single-tree removal for 7 years after harvest. Nitrate concentrations in forest floor and mineral soil were elevated 3–7 years after harvesting in gaps of 0.1 ha and larger. Removal of the same proportion of trees as single trees did not result in increased nitrate concentrations, suggesting that nitrate losses could be reduced by harvesting single trees rather than creating gaps. Greater N availability was not associated with faster rates of decomposition of litter and forest floor, which were similar in gaps of all sizes (0–10 ha). Reciprocal transplant of forest floor and soil from the 10-ha gaps and the uncut forests indicated that changes in the nature of the forest floor or soil following harvest had a greater influence on nitrate concentrations than the changes in environmental conditions in the gaps.

Genetic variation between and within populations of Engelmann spruce and subalpine fir

Genome ◽  
1990 ◽  
Vol 33 (1) ◽  
pp. 1-8 ◽  
Author(s):  
Kathleen L. Shea
Keyword(s):  
Genetic Variation ◽  
Genetic Structure ◽  
Genetic Differentiation ◽  
Colorado Front Range ◽  
Abies Lasiocarpa ◽  
Front Range ◽  
Dispersal Patterns ◽  
Subalpine Fir ◽  
Engelmann Spruce ◽  
Inbreeding Coefficients

The genetic structure of Engelmann spruce (Picea engelmannii) and subalpine fir (Abies lasiocarpa) in two adjacent sites in the Colorado Front Range was examined using allozyme data from 21 loci in spruce and 18 loci in fir. The genetic diversity measures of alleles per locus, percent loci polymorphic, and mean heterozygosity did not differ significantly between or within species. However, the observed heterozygosity (0.126 spruce, 0.081 fir) values suggest that Engelmann spruce is more genetically variable than subalpine fir. Mean inbreeding coefficients were twice as high in fir as in spruce (FIS = 0.154 spruce, 0.341 fir). There were significant differences in allele frequencies in both species between and within sites, and among age-classes, with spatial differences greater than temporal differences. F-statistics showed greater genetic differentiation within (FST = 2.3–2.6% spruce, 2.0–2.8% fir) than between sites (FST = 1.7% spruce, 1.3% fir). Both species had higher FST and genetic distance values, yet lower FIS values, for spatial subdivisions within rather than between sites. This pattern suggests that the size of a spatial subdivision, where maximum gene flow is 54 m, approximates the size of the largest panmictic unit. In contrast with studies on other conifers, these results suggest that factors such as differential selection pressures, seed dispersal patterns, and clumped spatial distribution have resulted in genetic differentiation at the microhabitat level in Engelmann spruce and subalpine fir.Key words: genetic variation, genetic structure, allozymes, Engelmann spruce, subalpine fir, microgeographic differentiation.

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