Response of small mammals to alternative stand structures in the mixed-conifer forest of northeastern CaliforniaThis article is one of a selection of papers from the Special Forum on Ecological Studies in Interior Ponderosa Pine — First Findings from Blacks Mountain Interdisciplinary Research.

2008 ◽  
Vol 38 (5) ◽  
pp. 943-955 ◽  
Author(s):  
Chris C. Maguire ◽  
Douglas A. Maguire ◽  
Tom E. Manning ◽  
Sean M. Garber ◽  
Martin W. Ritchie

A common, but largely untested, strategy for maintaining forest biodiversity is to enhance stand structural complexity. A silvicultural experiment was implemented from 1996 to 1998 at Blacks Mountain Experimental Forest, California, to test the efficacy of two levels of structural diversity (high versus low) and two levels of prescribed underburning (burn versus no burn) for maintaining or restoring biodiversity. Small mammals were trapped and tagged in experimental units for 2 noncontiguous weeks in fall 2003 and 2004. Total number of captures and number of captured individuals varied by year (P < 0.002). No treatment effects were detected for all species lumped together or for the three most frequent species analyzed separately ( Tamias amoenus J.A. Allen, 1890, Peromyscus maniculatus (Wagner, 1845), and Spermophilus lateralis (Say, 1823)), with the exception that T. amoenus was captured more often in burned units in 2004 (P = 0.004 for year × burn interaction). Mixed-effects regression models indicated that the number of captures and captured individuals of T. amoenus and P. maniculatus decreased with increasing residual basal area of overstory trees, but opposite results were obtained for S. lateralis. After accounting for residual stand density differences, T. amoenus was captured more frequently in units of low structural diversity and S. lateralis in units of high structural diversity.

2016 ◽  
Vol 46 (5) ◽  
pp. 745-752 ◽  
Author(s):  
Carrie R. Levine ◽  
Flora Krivak-Tetley ◽  
Natalie S. van Doorn ◽  
Jolie-Anne S. Ansley ◽  
John J. Battles

In the western United States, forests are experiencing novel environmental conditions related to a changing climate and a suppression of the historical fire regime. Mixed-conifer forests, considered resilient to disturbance due to their heterogeneity in structure and composition, appear to be shifting to a more homogeneous state, but the timescale of these shifts is not well understood. Our objective was to assess the effects of climate and fire suppression on stand dynamics and demographic rates of an old-growth mixed-conifer forest in the Sierra Nevada. We used a Bayesian hierarchical analysis to quantify species and community rates of recruitment, growth, and mortality. Despite a warming climate, we found that stand density, basal area, and carbon have increased over 56 years. Fir recruitment and growth significantly exceeded the community-level median rates, whereas pine recruitment and growth was significantly lower than the community-level median rates. Shifts in species composition from a well-mixed stand to a more dense fir-dominated stand appear to be driven by low growth and recruitment rates of pines relative to firs. In forests such as these with consistent and relatively low mortality rates, we recommend that restoration and management activities be focused on promoting pine recruitment and growth.


2008 ◽  
Vol 38 (5) ◽  
pp. 909-918 ◽  
Author(s):  
Jianwei Zhang ◽  
Martin W. Ritchie ◽  
William W. Oliver

A large-scale interior ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws.) study was conducted at the Blacks Mountain Experimental Forest in northeastern California. The primary purpose of the study was to determine the influence of structural diversity on the dynamics of interior pine forests at the landscape scale. High structural diversity (HiD) and low structural diversity (LoD) treatments were created with mechanical thinning on 12 main plots. Each plot was then split in half with one-half treated with prescribed fire. During the 5 year period after the treatments, the LoD treatments showed slightly higher periodic annual increments for basal area (BA) and significantly higher diameter increments than did the HiD treatments, although HiD carried twice as much BA as LoD did immediately after the treatments. Prescribed fire did not affect growth, but killed and (or) weakened some trees. No interaction between treatments was found for any variable. Stand density was reduced from the stands before treatments, but species composition did not change. Old dominant trees still grew and large snags were stable during the 5 year period. Treatments had minor impacts on shrub cover and numbers. These results suggest that ponderosa pine forest can be silviculturally treated to improve stand growth and health without sacrificing understory shrub diversity.


2010 ◽  
Vol 19 (1) ◽  
pp. iii ◽  
Author(s):  
Miguel G. Cruz ◽  
Martin E. Alexander ◽  
Ronald H. Wakimoto

Application of crown fire behavior models in fire management decision-making have been limited by the difficulty of quantitatively describing fuel complexes, specifically characteristics of the canopy fuel stratum. To estimate canopy fuel stratum characteristics of four broad fuel types found in the western United States and adjacent areas of Canada, namely Douglas-fir, ponderosa pine, mixed conifer, and lodgepole pine forest stands, data from the USDA Forest Service's Forest Inventory and Analysis (FIA) database were analysed and linked with tree-level foliage dry weight equations. Models to predict canopy base height (CBH), canopy fuel load (CFL) and canopy bulk density (CBD) were developed through linear regression analysis and using common stand descriptors (e.g. stand density, basal area, stand height) as explanatory variables. The models developed were fuel type specific and coefficients of determination ranged from 0.90 to 0.95 for CFL, between 0.84 and 0.92 for CBD and from 0.64 to 0.88 for CBH. Although not formally evaluated, the models seem to give a reasonable characterization of the canopy fuel stratum for use in fire management applications.


Forests ◽  
2019 ◽  
Vol 10 (12) ◽  
pp. 1113 ◽  
Author(s):  
Juhan Park ◽  
Hyun Seok Kim ◽  
Hyun Kook Jo ◽  
II Bin Jung

Research Highlights: Using a long-term dataset on temperate forests in South Korea, we established the interrelationships between tree species and structural diversity and forest productivity and stability, and identified a strong, positive effect of structural diversity, rather than tree species diversity, on productivity and stability. Background and Objectives: Globally, species diversity is positively related with forest productivity. However, temperate forests often show a negative or neutral relationship. In those forests, structural diversity, instead of tree species diversity, could control the forest function. Materials and Methods: This study tested the effects of tree species and structural diversity on temperate forest productivity. The basal area increment and relative changes in stand density were used as proxies for forest productivity and stability, respectively. Results: Here we show that structural diversity, but not species diversity, had a significant, positive effect on productivity, whereas species diversity had a negative effect, despite a positive effect on diversity. Structural diversity also promoted fewer changes in stand density between two periods, whereas species diversity showed no such relation. Structurally diverse forests might use resources efficiently through increased canopy complexity due to canopy plasticity. Conclusions: These results indicate reported species diversity effects could be related to structural diversity. They also highlight the importance of managing structurally diverse forests to improve productivity and stability in stand density, which may promote sustainability of forests.


2006 ◽  
Vol 84 (3) ◽  
pp. 404-411 ◽  
Author(s):  
A J Edelman ◽  
J L Koprowski

We compared home ranges of introduced Abert's squirrels (Sciurus aberti Woodhouse, 1853) in mixed-conifer forests of Arizona during non-mating and mating seasons. Because Abert's squirrels are reported to depend on ponderosa pine (Pinus ponderosa P. & C. Lawson) forests, the mixed-conifer forest in our study represented a novel habitat. Home-range size, home-range overlap with females, and movement distances increased for males from non-mating to mating seasons. Home-range size and overlap characteristics of females remained consistent between seasons, but movement distances were reduced during the mating season. Males probably increased home-range size, home-range overlap with females, and movement distances during the mating season to maximize contact with scarce females. Home-range size and overlap characteristics of female Abert's squirrels likely remained stable between seasons because females do not search for mates. Restricted movements by females during the mating season may be due to changes in resource use in preparation for reproduction. Non-mating season home ranges in our study were smaller than home ranges observed in ponderosa pine forest. Abert's squirrels in mixed-conifer forest may have small home ranges because resource quality is higher than in ponderosa pine forest or competition for space with co-occurring Mount Graham red squirrels (Tamiasciurus hudsonicus grahamensis (J.A. Allen, 1894)).


2003 ◽  
Vol 12 (1) ◽  
pp. 39 ◽  
Author(s):  
Miguel G. Cruz ◽  
Martin E. Alexander ◽  
Ronald H. Wakimoto

Application of crown fire behavior models in fire management decision-making have been limited by the difficulty of quantitatively describing fuel complexes, specifically characteristics of the canopy fuel stratum. To estimate canopy fuel stratum characteristics of four broad fuel types found in the western United States and adjacent areas of Canada, namely Douglas-fir, ponderosa pine, mixed conifer, and lodgepole pine forest stands, data from the USDA Forest Service's Forest Inventory and Analysis (FIA) database were analysed and linked with tree-level foliage dry weight equations. Models to predict canopy base height (CBH), canopy fuel load (CFL) and canopy bulk density (CBD) were developed through linear regression analysis and using common stand descriptors (e.g. stand density, basal area, stand height) as explanatory variables. The models developed were fuel type specific and coefficients of determination ranged from 0.90 to 0.95 for CFL, between 0.84 and 0.92 for CBD and from 0.64 to 0.88 for CBH. Although not formally evaluated, the models seem to give a reasonable characterization of the canopy fuel stratum for use in fire management applications.


2013 ◽  
Vol 43 (4) ◽  
pp. 311-320 ◽  
Author(s):  
Jianwei Zhang ◽  
Martin W. Ritchie ◽  
Douglas A. Maguire ◽  
William W. Oliver

We analyzed 45 years of data collected from three ponderosa pine (Pinus ponderosa Douglas ex P. Lawson & C. Lawson) levels-of-growing-stock installations in Oregon (OR) and northern California (CA), USA, to determine the effect of stand density regimes on stand productivity and mortality. We found that periodic annual increment (PAI) of diameter, basal area (BA), volume, and aboveground dry mass were significantly related to stand density index (SDI) and stand age at start of the period; the quadratic trends varied among sites. Precipitation departure from the normal for each period explained a significant amount of residual variation in all PAI variables except diameter. BA production did not change significantly as SDI exceeded 270 trees·ha−1 at the OR sites and 320 trees·ha−1 at the CA site. Stand productivity was the highest at Elliot Ranch (CA) and the least at Blue Mountains (OR). A similar trend held in growth efficiency under lower stand densities (SDI < 600). Most of the mortality was caused by Dendroctonus bark beetles in stands that exceeded SDI of 500 trees·ha−1. Limiting SDI was about 900 trees·ha−1, although plots at Elliot Ranch reached much higher than that. The results demonstrate that silvicultural control of stand density can be a powerful tool for reducing bark beetle caused mortality without sacrificing stand productivity.


2019 ◽  
Vol 65 (4) ◽  
pp. 496-507 ◽  
Author(s):  
Jianwei Zhang ◽  
Kaelyn A Finley ◽  
Nels G Johnson ◽  
Martin W Ritchie

AbstractStand density affects not only structure and growth, but also the health of forests and, subsequently, the functions of forest ecosystems. Here, we integrated dendrochronology and repeated inventories for ponderosa pine research plots to determine whether long-term growth and mortality responded to climate trends and how varying stand density influenced the responses. The plots were established prior to 1975 on existing stands throughout northern California. Although annual temperature increased consistently for the last 65 years, ring-width indices produced by eliminating age and thinning effects failed to detect radial trend regardless of site quality. However, interannual variation for the indices was substantial, reflecting a strong influence of climate on tree growth. Plot-level basal area increments were significantly affected by tree mortality. Stand density index explained most variation of mortality. Lowering stand density enhanced remaining tree growth, reduced mortality, and increased stand resiliency to disturbances and climate change. Besides higher climate moisture indices or lower vapor pressure deficits, any treatments that improve tree vigor and reduce stress will have a similar effect to reducing stand density. Although neither biotic disturbances nor abiotic conditions can be controlled, forest managers can manage stand density appropriately to enhance resilience to climate change and disturbances.


1995 ◽  
Vol 10 (3) ◽  
pp. 91-94
Author(s):  
Darrell W. Ross

Abstract Second-growth ponderosa pine (Pinus ponderosa) stands with outbreak populations of the pandora moth (Coloradia pandora) were thinned from below removing about half of the basal area. Thinning had no effect on pandora moth pupal density or weight, or emerging adult density in the following generation. However, adult emergence and egg hatch occurred 7-10 days earlier in thinned plots compared with unthinned plots. Egg and larval densities on a foliage weight basis were not significantly different between thinned and unthinned plots. Thinning stands infested with pandora moth will not significantly affect the course of an outbreak for at least one generation. Timing of direct controls for the pandora moth should consider the effect of stand density on insect phenology. West. J. Appl. For. 10(3):91-94.


1994 ◽  
Vol 4 (1) ◽  
pp. 3 ◽  
Author(s):  
PZ Fule ◽  
WW Covington

Stand and natural fuel conditions were sampled in ponderosa pine forests in northern and central Arizona to develop predictive fuel depth and loading equations. Litter and duff depths can be estimated from measurements of stand density (basal area, stand density index). Although woody fuel loading did not correlate well with stand variables, correlations were found among loadings of different woody fuel size classes, so that results from a planar intersect tally of certain single woody fuel size classes may be used to predict the loadings in certain other size classes. The relatively low precision of estimates from these predictive equations can be substantially increased by applying them in a double sampling scheme. Making use of these predictive relationships, managers can devise simple, rapid, arid cost-effective fuel inventories that focus on the fuel category of interest. Fuel loads can be estimated at a desired precision with reduced investment of time and funds compared to a more comprehensive direct fuel inventory.


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