Vegetation responses to stand structure and prescribed fire in an interior ponderosa pine ecosystemThis article is one of a selection of papers from the Special Forum on Ecological Studies in Interior Ponderosa Pine — First Findings from Blacks Mountain Interdisciplinary Research.

2008 ◽  
Vol 38 (5) ◽  
pp. 909-918 ◽  
Author(s):  
Jianwei Zhang ◽  
Martin W. Ritchie ◽  
William W. Oliver

A large-scale interior ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws.) study was conducted at the Blacks Mountain Experimental Forest in northeastern California. The primary purpose of the study was to determine the influence of structural diversity on the dynamics of interior pine forests at the landscape scale. High structural diversity (HiD) and low structural diversity (LoD) treatments were created with mechanical thinning on 12 main plots. Each plot was then split in half with one-half treated with prescribed fire. During the 5 year period after the treatments, the LoD treatments showed slightly higher periodic annual increments for basal area (BA) and significantly higher diameter increments than did the HiD treatments, although HiD carried twice as much BA as LoD did immediately after the treatments. Prescribed fire did not affect growth, but killed and (or) weakened some trees. No interaction between treatments was found for any variable. Stand density was reduced from the stands before treatments, but species composition did not change. Old dominant trees still grew and large snags were stable during the 5 year period. Treatments had minor impacts on shrub cover and numbers. These results suggest that ponderosa pine forest can be silviculturally treated to improve stand growth and health without sacrificing understory shrub diversity.

2013 ◽  
Vol 43 (4) ◽  
pp. 311-320 ◽  
Author(s):  
Jianwei Zhang ◽  
Martin W. Ritchie ◽  
Douglas A. Maguire ◽  
William W. Oliver

We analyzed 45 years of data collected from three ponderosa pine (Pinus ponderosa Douglas ex P. Lawson & C. Lawson) levels-of-growing-stock installations in Oregon (OR) and northern California (CA), USA, to determine the effect of stand density regimes on stand productivity and mortality. We found that periodic annual increment (PAI) of diameter, basal area (BA), volume, and aboveground dry mass were significantly related to stand density index (SDI) and stand age at start of the period; the quadratic trends varied among sites. Precipitation departure from the normal for each period explained a significant amount of residual variation in all PAI variables except diameter. BA production did not change significantly as SDI exceeded 270 trees·ha−1 at the OR sites and 320 trees·ha−1 at the CA site. Stand productivity was the highest at Elliot Ranch (CA) and the least at Blue Mountains (OR). A similar trend held in growth efficiency under lower stand densities (SDI < 600). Most of the mortality was caused by Dendroctonus bark beetles in stands that exceeded SDI of 500 trees·ha−1. Limiting SDI was about 900 trees·ha−1, although plots at Elliot Ranch reached much higher than that. The results demonstrate that silvicultural control of stand density can be a powerful tool for reducing bark beetle caused mortality without sacrificing stand productivity.


1995 ◽  
Vol 10 (3) ◽  
pp. 91-94
Author(s):  
Darrell W. Ross

Abstract Second-growth ponderosa pine (Pinus ponderosa) stands with outbreak populations of the pandora moth (Coloradia pandora) were thinned from below removing about half of the basal area. Thinning had no effect on pandora moth pupal density or weight, or emerging adult density in the following generation. However, adult emergence and egg hatch occurred 7-10 days earlier in thinned plots compared with unthinned plots. Egg and larval densities on a foliage weight basis were not significantly different between thinned and unthinned plots. Thinning stands infested with pandora moth will not significantly affect the course of an outbreak for at least one generation. Timing of direct controls for the pandora moth should consider the effect of stand density on insect phenology. West. J. Appl. For. 10(3):91-94.


2002 ◽  
Vol 32 (8) ◽  
pp. 1360-1371 ◽  
Author(s):  
R C Godfree ◽  
R O Tinnin ◽  
R B Forbes

We investigated the importance of lodgepole pine dwarf mistletoe (Arceuthobium americanum Nutt.) in determining the height to crown top (HCT), height to crown base (HCB), and live crown ratio (LCR) of 2025 lodgepole pine (Pinus contorta var. murrayana (Grev. & Balf.) Engelm.) growing over a 24-km2 study site in central Oregon. We compared the effects of infection and associated witches' brooms with those of site topography, soil type, shrub cover, stand density, and the abundance of mature ponderosa pine (Pinus ponderosa Dougl. ex P. Laws. & C. Laws). using multiple regression and path analysis. The density of dominant-size P. contorta was consistently the most important factor influencing HCT, HCB, and LCR across the study site. In dense stands, trees tended to have elevated crown bases due to self-pruning and, hence, lower values of LCR. Dwarf mistletoe and related witches' brooms uniquely explained 6.9% of the variance in LCR, which was close to that of dominant P. contorta (7.1%) and more than that of soil type (3.0%), but explained only 2.6% of the variance in HCB, which was less than that of dominant P. contorta (6.5%) and soil type (4.6%). Regression models suggest that heavily infected trees should be 18% shorter and have crown bases 37% lower than uninfected trees, while moderately infected trees should have an LCR over 20% larger than that of uninfected and heavily infected trees. We also found that the largest 25 heavily infected trees sampled were approximately 19% shorter and 11–13% smaller in diameter than the largest 25 uninfected trees. The results suggest that dwarf mistletoe can be an important factor in determining the crown dimensions of P. contorta but that these effects may be interpreted only in the context of site characteristics and stand structure.


1997 ◽  
Vol 12 (4) ◽  
pp. 122-130 ◽  
Author(s):  
William W. Oliver

Abstract A 20 yr old ponderosa pine (Pinus ponderosa var. ponderosa) plantation on a productive site on the west slope of the Sierra Nevada in northern California was thinned four times over a 25 yr period. Stand densities tested were Stand Density Indexes (SDI) of 73, 128, 183, 238, and 293 (equivalent to 40,70,100,130, and 160 ft2/ac of basal area), replicated three times in a randomized design. Growth was analyzed for each of five 5 yr periods. Periodic annual increments (PAI) of diameter, net basal area, and net total volume differed significantly among periods and, in the earlier periods, among stocking levels. Mortality from winter storms and bark beetles was largely confined to the higher stand densities and in periods 3 and 4 caused PAIs of net basal area and net total volume to decline below that of lower densities. The sensitivity of mortality to stand density suggests a thinning target of SDI 183 (about 100 ft²/ac of basal area) for similar stands—no higher than that recommended for eastside stands of much lower site productivity. This sensitivity coupled with rapid growth suggests that multiple thinnings will be necessary in similar stands to maintain healthy, vigorous trees. West. J. Appl. For. 12(4):122-130.


2008 ◽  
Vol 38 (5) ◽  
pp. 969-980 ◽  
Author(s):  
Nancy E. Gillette ◽  
Richard S. Vetter ◽  
Sylvia R. Mori ◽  
Carline R. Rudolph ◽  
Dessa R. Welty

We assessed spider (Arachnida: Araneae) responses to prescribed fire following stand structure treatments in ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws.) stands in the Cascade Range of California. Stands were logged or left untreated to create three levels of structural diversity. We logged one treatment to minimize old-growth characteristics (low diversity) and one to enhance old-growth characteristics (high diversity) and we used unlogged Research Natural Areas (RNAs) as old-growth, highest-diversity reference stands. We conducted low-intensity prescribed fire on half of each plot following harvest. Spider assemblages in unburned, logged stands were similar to one another but diverged from those in RNAs, with increased abundance, species richness, and diversity in more structurally diverse stands. Prescribed fire, which altered habitat in the organic soil layer where many spiders forage, resulted in altered spider assemblages and population declines in most plots. Fire generally reduced spider species richness, evenness, and diversity. Several taxa were potential indicators of fire and old-growth structure, and we discovered one species and one genus that were previously unknown. There was evidence that old-growth characteristics intensified the effects of fire on spider abundance. This outcome probably results from the deep litter layers in high-diversity stands and RNAs, which constituted greater fuel loads than low-diversity stands.


2005 ◽  
Vol 20 (4) ◽  
pp. 240-246
Author(s):  
Nicholas Vaughn ◽  
Martin W. Ritchie

Abstract We evaluated the relationship between crown cover measured with a vertical sight tube and stand basal area per acre in treated (thinned, burned, and thinned and burned) and untreated interior ponderosa pine (Pinus ponderosa P. & C. Lawson) stands in northeastern California. Crown cover was significantly related to basal area at the plot level and stand level. In addition, the relationship was not affected by two extremely different thinning regimes. However, the predicted crown cover was generally lower, for a given level of basal area, in a recently thinned stand than in a stand that had not been recently thinned. Prescribed fire had no detectable effect on the relationship. The maximum measured value of stand level crown cover in untreated stands in this study was about 60 percent. West. J. Appl. For. 20(4): 240–246.


2008 ◽  
Vol 38 (5) ◽  
pp. 943-955 ◽  
Author(s):  
Chris C. Maguire ◽  
Douglas A. Maguire ◽  
Tom E. Manning ◽  
Sean M. Garber ◽  
Martin W. Ritchie

A common, but largely untested, strategy for maintaining forest biodiversity is to enhance stand structural complexity. A silvicultural experiment was implemented from 1996 to 1998 at Blacks Mountain Experimental Forest, California, to test the efficacy of two levels of structural diversity (high versus low) and two levels of prescribed underburning (burn versus no burn) for maintaining or restoring biodiversity. Small mammals were trapped and tagged in experimental units for 2 noncontiguous weeks in fall 2003 and 2004. Total number of captures and number of captured individuals varied by year (P < 0.002). No treatment effects were detected for all species lumped together or for the three most frequent species analyzed separately ( Tamias amoenus J.A. Allen, 1890, Peromyscus maniculatus (Wagner, 1845), and Spermophilus lateralis (Say, 1823)), with the exception that T. amoenus was captured more often in burned units in 2004 (P = 0.004 for year × burn interaction). Mixed-effects regression models indicated that the number of captures and captured individuals of T. amoenus and P. maniculatus decreased with increasing residual basal area of overstory trees, but opposite results were obtained for S. lateralis. After accounting for residual stand density differences, T. amoenus was captured more frequently in units of low structural diversity and S. lateralis in units of high structural diversity.


1993 ◽  
Vol 8 (4) ◽  
pp. 126-132 ◽  
Author(s):  
P. H. Cochran ◽  
James W. Barrett

Abstract A spacing study in ponderosa pine (Pinus ponderosa) was established in 1959 by thinning plots in a 33-yr-old plantation near John Day, Oregon. The influence of 4 spacings (17.2, 12.5, 10.1, and 8.7 ft) on stand and tree growth for a 31-yr period was examined. Study plots were remeasured five times after establishment. Periodic annual increments (PAI) of gross basal area, gross volume, and average height differed with period but not with spacing (P ≤ 0.10). The PAIs of mean diameter differed with period and decreased with increasing density. Annual height growth and annual gross and net growth of basal area and volume did not differ with spacing. Annual diameter growth was much greater for trees at the widest spacing. Annual volume growth of the largest 90 trees/ac was greatest at the widest spacing. Thirty-one years after thinning, the largest 90 trees/ac on the widest spacing had 73% of the volume of all the trees on the narrowest spacing. Mortality due to mountain pine beetle (Dendroctonus ponderosae) increased markedly when values for stand density index exceeded 200. Wide spacings increased average tree volumes, increased mean diameters, and reduced the probability of mortality without sacrificing gross cubic volume growth potential. West. J. Appl. For. 8(4):126-132.


2016 ◽  
Vol 46 (1) ◽  
pp. 101-113 ◽  
Author(s):  
Kathryn I. McGown ◽  
Kevin L. O’Hara ◽  
Andrew Youngblood

We used six metrics of size and growth variation (standard deviation (SD), coefficient of variation (CV), skewness coefficient (S), Gini coefficient (G), Lorenz asymmetry coefficient (LAC), and growth dominance coefficient (GD)) to describe changes in two long-term ponderosa pine (Pinus ponderosa Douglas ex P. Lawson & C. Lawson) initial spacing trials in Oregon and Washington, USA. Trends were examined over a 35-year time period and across a range of initial stand densities (from 154 to 2470 trees·ha−1) for four measures of tree size: diameter at breast height (dbh, 1.37 m), basal area (BA), height, and volume. Unlike many previous studies of size variation in monospecific stands, our results suggest that variation declined or remained relatively stable for all treatments at both study areas. This suggests that these stands are experiencing size symmetric competition for belowground resources. We found that a combination of metrics is necessary to provide a complete picture of size variability and differentiation in developing stands. We recommend using the CV or G, as there were clear trends with increasing density for all size variables. If the objective of the assessment was to track changes in absolute size within an individual stand, we would recommend using the SD, as there were consistent trends with time for all size variables. S, LAC, and GD may be less suited for comparing differentiation during the early stages of stand development because of a lack of clear trends with stand density and time.


2020 ◽  
Vol 66 (No. 5) ◽  
pp. 197-207 ◽  
Author(s):  
Vilém Podrázský ◽  
Zdeněk Vacek ◽  
Stanislav Vacek ◽  
Jan Vítámvás ◽  
Josef Gallo ◽  
...  

Scots pine (Pinus sylvestris L.) is one of the most important tree species in Eurasia. During the past centuries, it has been extensively introduced into artificial monocultures, but is currently experiencing a number of problems related to climate change and extreme droughts. There is a large-scale disintegration of its stands and, in addition to its replacement by other native trees, it is possible to use a wide range of introduced species of the same genus. The aim of the investigation was to compare production parameters, structure and diversity of pine stands at the age of 35 years in school Arboretum of Faculty of Forestry and Wood Science in Central Bohemia (320 m a.s.l., medium rich habitats, water deficit site). Seven species of pine were compared: ponderosa pine (Pinus ponderosa Douglas ex C. Hawson), Jeffrey pine (Pinus jeffreyi Balf.), black pine (Pinus nigra J.F.Arnold), eastern white pine (Pinus strobus L.), Lodgepole pine (Pinus contorta Douglas), Macedonian pine (Pinus peuce Griseb.) and the only native Scots pine. The results showed that significantly (P &lt; 0.001) highest height, diameter at breast height and mean stem volume were achieved in Pinus ponderosa and P. strobus stands, while these parameters were lowest in P. peuce and P. nigra. In contrast, the lowest stand volume was calculated for P. strobus (112 m3·ha–1) due to the lower stand density, while the highest production was again in P. ponderosa (430 m3·ha–1). In terms of structural variability, the highest diversity was found in P. jeffreyi and P. peuce. The introduced pine species, especially P. ponderosa, could therefore play an important role in terms of production and economic potential and even replace native P. sylvestris on suitable sites.


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