Underplanted conifer seedling survival and growth in thinned Douglas-fir stands

2001 ◽  
Vol 31 (2) ◽  
pp. 302-312 ◽  
Author(s):  
Thomas J Brandeis ◽  
Michael Newton ◽  
Elizabeth C Cole
Keyword(s):  
Seedling Survival ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Understory Vegetation ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Herbicide Application ◽  
Western Redcedar ◽  
Second Growth ◽  
Conifer Seedling

In a multilevel study to determine limits to underplanted conifer seedling growth, Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco), grand fir (Abies grandis (Dougl. ex D. Don) Lindl.), western redcedar (Thuja plicata Donn ex D. Don), and western hemlock (Tsuga heterophylla (Raf.) Sarg.) seedlings were planted beneath second-growth Douglas-fir stands that had been thinned to basal areas ranging from 16 to 31 m2/ha. Understory vegetation was treated with a broadcast herbicide application prior to thinning, a directed release herbicide application 2 years later, or no treatment beyond harvest disturbance. Residual overstory density was negatively correlated with percent survival for all four species. Broadcast herbicide application improved survival of grand fir and western hemlock. Western redcedar, grand fir, and western hemlock stem volumes were inversely related to overstory tree density, and this effect increased over time. There was a strong indication that this was also the case for Douglas-fir. Reduction of competing understory vegetation resulted in larger fourth-year stem volumes in grand fir and western hemlock.

Ionic balance and organic acids in western redcedar, western hemlock, and Douglas-fir seedlings grown in low- and high-N soils

1999 ◽  
Vol 29 (6) ◽  
pp. 669-678 ◽  
Author(s):  
J E Graff, Jr. ◽  
R K Hermann ◽  
J B Zaerr
Keyword(s):  
Organic Acids ◽  
Quinic Acid ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
N Availability ◽  
Ionic Balance ◽  
Total N ◽  
Western Redcedar

Seedlings of western redcedar (Thuja plicata Donn ex. D. Don), western hemlock (Tsuga heterophylla (Raf.) Sarg.), and Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) were transplanted into soils with low and high levels of available NO3-(and total N). Current-year foliage was sampled after 10 weeks to determine the effect of N availability on foliar cation-anion balance (C-A) and the concentrations of low molecular weight organic acids of the three species. Carboxylate concentrations were estimated by using the difference between sums of cations and anions (C-A): 750 mequiv.·kg-1for western redcedar, 351 mequiv.·kg-1for western hemlock, and 266 mequiv.·kg-1for Douglas-fir. Quinic acid was a primary constituent, accounting for 40% of the total for western redcedar and 75% for western hemlock and Douglas-fir. Oxalic acid was present in greatest concentration in the foliage of western redcedar (65 mequiv.·kg-1) but was a minor constituent in western hemlock and Douglas-fir. The quantified acids accounted for only 15% of the C-A of western redcedar but >80% of the C-A of western hemlock and Douglas-fir. A considerable portion of the C-A balance not accounted for in redcedar may be associated with the accumulation of CaCO3. Litterfall deposition of CaCO3may lead to the consumption of H+ions and enrichment of exchangeable soil Ca in the rooting zone of long-lived western redcedar trees. No statistically significant differences among the soils were detected with regard to C-A or the concentration of organic acids.

Response of Gaultheria shallon and Epilobium angustifolium to large additions of nitrogen and phosphorus fertilizer

2004 ◽  
Vol 34 (2) ◽  
pp. 502-506 ◽  
Author(s):  
Jennifer N Bennett ◽  
Brent M Lapthorne ◽  
Leandra L Blevins ◽  
Cindy E Prescott
Keyword(s):  
Seedling Survival ◽  
Tsuga Heterophylla ◽  
Thuja Plicata ◽  
Ground Vegetation ◽  
N Availability ◽  
Gaultheria Shallon ◽  
Nitrogen And Phosphorus ◽  
N Application ◽  
Epilobium Angustifolium ◽  
Conifer Seedling

A study was established in coastal British Columbia to determine if repeated nitrogen (N) and phosphorus (P) fertilization negatively influences the reestablishment of salal (Gaultheria shallon Pursh) on cleared and burned cedar–hemlock (Thuja plicata Donn ex D. Don – Tsuga heterophylla (Raf.) Sarg.) forests. Fertilizers were applied for 3 years, and the biomass of ground vegetation and conifer seedling survival and growth were measured. Salal biomass decreased with high levels of N application (1000 kg N/ha), but not when 400 kg P/ha was added with 1000 kg N/ha. The addition of 500 kg N/ha, with or without P, stimulated salal growth. The biomass of fireweed (Epilobium angusti folium L.) increased with the addition of N + P but not with N alone. In the high N and N + P treatments, conifer seedling survival and heights were reduced. These results confirm earlier reports that salal responds negatively to high N applications and that this negative response can be alleviated with simultaneous additions of P. The response of fireweed to N + P, but not to N alone, suggests that the abundance of this species is more indicative of P than N availability.

Ectomycorrhizal mushroom response to partial cutting in a western hemlock - western redcedar forest

2001 ◽  
Vol 31 (6) ◽  
pp. 978-987 ◽  
Author(s):  
J M Kranabetter ◽  
P Kroeger
Keyword(s):  
Stand Structure ◽  
Basal Area ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Soil Conditions ◽  
Taxon Richness ◽  
Partial Cutting ◽  
Western Redcedar ◽  
Significant Block

We examined epigeous ectomycorrhizal mushroom richness and productivity after partial cutting in a western hemlock (Tsuga heterophylla (Raf.) Sarg.) - western redcedar (Thuja plicata Donn ex D. Don in Lamb.) forest of northwestern British Columbia. Mushrooms were collected throughout the fruiting season (July to October) for 3 years, starting 5 years after partial cutting, from plots with mesic soil conditions and residual basal areas ranging from 23 to 69 m2/ha for western hemlock and 0 to 26 m2/ha for western redcedar. Partial cutting had no apparent effect on mushroom phenology over the 3 years. Significant block interactions demonstrated that reductions in basal area of western hemlock could lead to positive, neutral, and negative responses in mushroom richness, biomass, and number of fruiting bodies. These responses were related to stand structure and the potential differences in tree vigour after partial cutting. In addition, there was weak evidence that western redcedar, a host for vesicular-arbuscular mycorrhiza, had a negative effect on average taxon richness. The study demonstrated that partial-cutting systems could allow some timber removal without necessarily reducing ectomycorrhizal mushroom communities.

Palynology and paleoecology of postglacial sediments in an anoxic basin, Saanich Inlet, British Columbia

1983 ◽  
Vol 20 (5) ◽  
pp. 873-885 ◽  
Author(s):  
Linda E. Heusser
Keyword(s):  
British Columbia ◽  
Pinus Contorta ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Alnus Rubra ◽  
Red Cedar ◽  
Saanich Inlet ◽  
Anoxic Basin

Varved, black clayey silts deposited in the marine waters of Saanich Inlet yield unusually abundant and diverse pollen assemblages derived from the coastal Douglas-fir (Pseudotsuga) and western hemlock (Tsuga heterophylla) forests of southwestern British Columbia. The 12 000 year palynological record chronicles the development of vegetation since ice left Saanich Inlet: the succession of pine (Pinus contorta) and alder (Alnus rubra) woodlands by forests characterized by Douglas-fir and oak (Quercus) and later by western hemlock and red cedar (Thuja plicata). Rapid deposition of annual layers of pollen, charcoal, and other terrigenous particles provides detailed evidence of changes in land use during the past few hundred years: settlement, logging, farming, and urbanization. Vegetational and climatic changes inferred from pollen spectra in the marine sediments of Saanich Inlet compare favorably with changes inferred from correlative pollen assemblages previously described from adjacent parts of Vancouver Island and the Fraser River valley.

Leaf area index of an old-growth Douglas-fir forest estimated from direct structural measurements in the canopy

2000 ◽  
Vol 30 (12) ◽  
pp. 1922-1930 ◽  
Author(s):  
Sean C Thomas ◽  
William E Winner
Keyword(s):  
Leaf Area Index ◽  
Leaf Area ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Understory Vegetation ◽  
Western Hemlock ◽  
Old Growth ◽  
Area Index ◽  
Contact Frequency ◽  
Canopy Access

Leaf area index (LAI) in old-growth Douglas-fir (Pseudotsuga menziesii var menziesii (Mirb.) Franco) forests exceeds that of any other forest ecosystem by some estimates; however, LAI determinations in coniferous forests have generally been indirect, involving extrapolations of patterns observed in younger stands. Aided by a 75-m construction crane for canopy access, we used a vertical line-intercept method to estimate LAI for a [Formula: see text]450-year-old Douglas-fir - western hemlock (Tsuga heterophylla (Raf.) Sarg.) forest in southwestern Washington state. LAI was calculated as the product of foliage contact frequency and an "extinction coefficient" accounting for foliage angular distribution, geometry, and the ratio of "interceptable" to total leaf area. LAI estimates were 9.3 ± 2.1 (estimate ± 95% confidence interval), 8.5 ± 2.2, and 8.2 ± 1.8 in 1997, 1998, and 1999, respectively, or 8.6 ± 1.1 pooled across years. Understory vegetation, including foliage of woody stems <5 cm diameter, represented 20% of this total. Sample points in which Douglas-fir was dominant had a higher total LAI than points dominated by western hemlock, including a higher LAI of understory vegetation. Our results do not support the contention that old-growth Douglas-fir - western hemlock forests maintain an appreciably higher LAI than do other forest ecosystems. Moreover, LAI in very old stands may decline as western hemlock replaces Douglas-fir through the course of succession.

Overturning resistance of western redcedar and western hemlock in mixed-species stands in coastal British Columbia

2007 ◽  
Vol 37 (5) ◽  
pp. 931-939 ◽  
Author(s):  
Kenneth E. Byrne ◽  
Stephen J. Mitchell
Keyword(s):  
British Columbia ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Specific Information ◽  
Mixed Species ◽  
Stem Form ◽  
Western Redcedar ◽  
Applied Forces ◽  
Coastal British Columbia

Specific information about the applied forces that cause trees to fail is required to validate mechanistic models of windthrow in different forest types. Static tree-pulling tests were conducted to examine the overturning resistance of western redcedar ( Thuja plicata Donn ex D. Don) and western hemlock ( Tsuga heterophylla (Raf.) Sarg.) in a mixed species second-growth stand in coastal British Columbia. Although widely used, tree-pulling techniques are not standardized. Data from three inclinometers were used to estimate stem deflection, which was found to increase with tree slenderness. Differing methods of fitting stem curvature had a small effect on estimates of self-loading at failure. The distance of the pivot point from the centre of the stem base increased with tree diameter. Accounting for the correct self-loading at failure produced a small difference in the overall turning moment regressions but did not improve the fit of these regressions. However, this difference increased with tree size and warrants consideration in future tree-pulling tests with large or plate-rooted trees. The stem mass – overturning resistance relationship had the best fit and was not significantly different for these species in spite of their differences in wood density and stem form.

Influence of millipedes on litter decomposition, N mineralization, and microbial communities in a coastal forest in British Columbia, Canada

2000 ◽  
Vol 30 (5) ◽  
pp. 817-826 ◽  
Author(s):  
H A Cárcamo ◽  
T A Abe ◽  
C E Prescott ◽  
F B Holl ◽  
C P Chanway
Keyword(s):  
Litter Decomposition ◽  
N Mineralization ◽  
Sitka Spruce ◽  
Douglas Fir ◽  
Picea Sitchensis ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Coastal Forests ◽  
Western Redcedar ◽  
Litter Consumption

Laboratory experiments were conducted with the millipede Harpaphe haydeniana haydeniana Wood (Polydesmida: Xystodesmidae) to determine (i) its litter feeding preferences, (ii) rates of leaf litter consumption, (iii) feeding effects on available nitrogen, and (iv) functional microbial diversity. The millipede exhibited a preference for Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and, to a lesser extent, Sitka spruce (Picea sitchensis (Bong.) Carrière) litter compared with western hemlock (Tsuga heterophylla (Raf.) Sarg.) and western redcedar (Thuja plicata Donn ex D. Don) litter when given a choice. When only one litter type was provided, millipedes consumed considerably more western redcedar than Douglas-fir, Sitka spruce, or western hemlock. Among the six broadleaf species tested, paper birch (Betula papyrifera Marsh.), bigleaf maple (Acer macrophyllum Pursh), vine maple (Acer circinatum Pursh), and red alder (Alnus rubra Bong.) were consumed at much higher rates than swordfern (Polystichum munitum (Kaulf.) Presl.) or salal (Gaultheria shallon Pursh). Daily rates of conifer litter consumption ranged between 10 and 20% of the millipede's fresh biomass and may translate to 36% of the annual litter fall. Our results suggest that transformation of conifer litter into millipede frass can increase rates of litter decomposition and N mineralization, as well as influence microbial activity and diversity in coastal forests.

Life History and Habits of Pseudohylesinus nebulosus (Leconte) (Coleoptera: Scolytidae) in the Interior of British Columbia

1956 ◽  
Vol 88 (5) ◽  
pp. 197-202 ◽  
Author(s):  
J. Walters ◽  
L. H. McMullen
Keyword(s):  
British Columbia ◽  
Pinus Ponderosa ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Western North America ◽  
The West ◽  
Red Cedar ◽  
Yellow Pine

The Douglas-fir hylesinus, Pseudohylesinus nebulosus (Leconte), is a common bark beetle in western North America from British Columbia to Mexico. Although Douglas fir, Pseudotsuga menziesii (Mirb.) Franco, is the preferred host, the beetle has been collected also from western red cedar, Thuja plicata Donn, grand fir, Abies grandis (Dougl.) Lindl., amabilis fir, Abies amabilis (Dougl.) Forb., and western hemlock, Tsuga heterophylla (Raf.) Sarg., on the west coast of British Columbia, and from western hemlock and western yellow pine, Pinus ponderosa Laws., in the interior of the Province. Swaine (1918) describes Pseudohylesinus nebulosus (Lec.) as “a slender species, with strong colour-markings in dark and light reddish-brown; the male very densely clothed with stout scales; the epistomal lobe strongly developed; length, 2.8 mm.; width, 1.2 mm. The supposed female has interspace 9 on the declivity less strongly serrate, and the elytral scales decidedly elongate and becoming plumose towards the base.”

Old-Growth Douglas-Fir and Western Hemlock: A 36-Year Record of Growth and Mortality

1987 ◽  
Vol 2 (4) ◽  
pp. 111-114 ◽  
Author(s):  
Dean S. DeBell ◽  
Jerry F. Franklin
Keyword(s):  
Volume Growth ◽  
Tsuga Heterophylla ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Silver Fir ◽  
Pinus Monticola ◽  
White Pine ◽  
Old Growth ◽  
Stand Volume

Abstract Growth and mortality were measured at 6-year intervals in a 1,180-acre old-growth stand in southwestern Washington. Principal tree species were Douglas-fir (Pseudotsuga menziesii), western hemlock (Tsuga heterophylla), Pacific silver fir (Abies amabilis), western redcedar (Thuja plicata), and western white pine (Pinus monticola). They composed 59, 27, 6, 6, and 1%, respectively, of the total cubic volume (13,290 ft³) in 1947. Gross volume growth averaged 94 ft³ per acre per year, and mortality averaged 86 ft³ per acre per year. Net growth was therefore minimal, and total stand volume remained nearly constant for 36 years. Douglas-fir, which accounted for only one-third of the gross growth and nearly one-half of the mortality, is losing dominance to western hemlock, which provided nearly one-half the gross growth and only 28% of the mortality. Pacific silver fir increased in importance in the lower canopy and composed 60% of the in-growth. Thus, although net gain in timber volume was nil, substantial changes occurred in stand characteristics during the 1947-1983 period. West. J. Appl. For. 2(4):111-114, October 1987.

Tenth-year survival and size of underplanted seedlings in the Oregon Coast Range

2009 ◽  
Vol 39 (3) ◽  
pp. 580-595 ◽  
Author(s):  
Elizabeth Cole ◽  
Michael Newton
Keyword(s):  
Stand Structure ◽  
Site Preparation ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Coast Range ◽  
Oregon Coast Range ◽  
Western Redcedar ◽  
Oregon Coast ◽  
Forest Stand Structure

This study initiated a two-aged forest stand structure by underplanting 50-year-old stands, primarily of Douglas-fir ( Pseudotsuga menziesii (Mirb.) Franco var. menziesii) and Douglas-fir – western hemlock ( Tsuga heterophylla (Raf.) Sarg.), thinned to 19–33 m2/ha on interior and coastal sites in the Oregon Coast Range. Douglas-fir, grand fir ( Abies grandis (Dougl. ex D. Don) Lindl.) (interior site only), western hemlock, and western redcedar ( Thuja plicata Donn ex D. Don) were planted following thinning either uniformly or in gaps of 0.06 or 0.1 ha. Understory vegetation treatments included (i) a preharvest site preparation herbicide application and an untreated control at both sites and (ii) a postharvest herbicide release treatment at the interior site. Planting conditions and stock at the interior site were not ideal, so survival was adjusted for first-year mortality. Adjusted 10 year survival ranged from 27% to 56% for Douglas-fir, 47% to 65% for western hemlock, 61% to 80% for grand fir, and 78% to 96% for western redcedar. Tenth-year survival at the coastal site ranged from 79% to 92% for Douglas-fir, 61% to 75% for western hemlock, and 67% to 86% for western redcedar. All species grew moderately well beneath the lowest-density overstories, and size was better within gaps than matrices for most species. Understory site preparation improved size for most species. Browsing on Douglas-fir and western redcedar impacted size on both sites.

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