The Eocene Thomas Ranch flora, Allenby Formation, Princeton, British Columbia, Canada

Botany ◽  
2013 ◽  
Vol 91 (8) ◽  
pp. 514-529 ◽  
Author(s):  
Richard M. Dillhoff ◽  
Thomas A. Dillhoff ◽  
David R. Greenwood ◽  
Melanie L. DeVore ◽  
Kathleen B. Pigg

A flora from Thomas Ranch near Princeton, British Columbia, Canada, is assessed for biodiversity and paleoclimate. This latest Early to early Middle Eocene flora occurs in the Allenby Formation. Seventy-six megafossil morphotypes have been recognized, representing at least 62 species, with 29 identified to genus or species. Common taxa include Ginkgo L., Metasequoia Miki, Sequoia Endl., Abies Mill., Pinus L., Pseudolarix Gordon, Acer L., Alnus Mill., Betula L., Fagus L., Sassafras J Presl, Macginitiea Wolfe & Wehr, Prunus L., and Ulmus L. More than 70 pollen and spore types are recognized, 32 of which are assignable to family or genus. The microflora is dominated by conifers (85%–97% abundance), with Betulaceae accounting for most of the angiosperms. The Climate Leaf Analysis Multivariate Program (CLAMP) calculates a mean annual temperature (MAT) of 9.0 ± 1.7 °C and bioclimatic analysis (BA) calculates a MAT of 12.8 ± 2.5 °C. Coldest month mean temperature (CMMT) was >0 °C. Mean annual precipitation (MAP) was >70 cm/year but is estimated with high uncertainty. Both the CLAMP and BA estimates are at the low end of the MAT range previously published for other Okanagan Highland localities, indicating a temperate climate consistent with a mixed conifer–deciduous forest.

2017 ◽  
Vol 54 (9) ◽  
pp. 981-992 ◽  
Author(s):  
Jaelyn J. Eberle ◽  
David R. Greenwood

We describe Eocene fossils of the tillodont Trogosus from the Allenby Formation in Princeton, British Columbia (B.C.), as well as teeth of Brontotheriina from the lower Australian Creek Formation near Quesnel, B.C. These fossils represent the only occurrence of Tillodontia and Brontotheriidae in B.C. Further, the presence of the largest species of Trogosus — T. latidens — as well as a smaller species identified only as Trogosus sp. supports a late early – early middle Eocene (Bridgerian) age for the Vermilion Bluffs Shale of the Allenby Formation. Based on their morphology and large size, the teeth referred here to Brontotheriina represent one of the larger, more derived brontothere genera, and suggest a Uintan–Chadronian (middle–late Eocene) age range for the lower Australian Creek Formation that is consistent with radiometric ages of underlying volcanic rocks. Paleobotanical data from sediments correlative to those that produced these Eocene mammal fossils suggest they inhabited forested landscapes interspersed with swamps and open water environments, under mild and wet temperate climates (mean annual temperature (MAT) ∼10–16 °C; cold month mean temperature (CMMT) −4–4 °C; mean annual precipitation (MAP) >100 cm/year). These mixed conifer–broadleaf forests included tree genera typical of modern eastern North American forests (e.g., Tsuga, Acer, Fagus, and Sassafras), together with genera today restricted to east Asia (e.g., Metasequoia, Cercidiphyllum, Dipteronia, and Pterocarya). The paleobotanical evidence is consistent with the hypothesized habitats of both tillodonts and brontotheres.


2018 ◽  
Vol 15 (17) ◽  
pp. 5329-5341 ◽  
Author(s):  
Jiguang Feng ◽  
Jingsheng Wang ◽  
Yanjun Song ◽  
Biao Zhu

Abstract. Soil respiration (Rs), a key process in the terrestrial carbon cycle, is very sensitive to climate change. In this study, we synthesized 54 measurements of annual Rs and 171 estimates of Q10 value (the temperature sensitivity of soil respiration) in grasslands across China. We quantitatively analyzed their spatial patterns and controlling factors in five grassland types, including temperate typical steppe, temperate meadow steppe, temperate desert steppe, alpine grassland, and warm, tropical grassland. Results showed that the mean (±SE) annual Rs was 582.0±57.9 g C m−2 yr−1 across Chinese grasslands. Annual Rs significantly differed among grassland types, and was positively correlated with mean annual temperature, mean annual precipitation, soil temperature, soil moisture, soil organic carbon content, and aboveground biomass, but negatively correlated with soil pH (p<0.05). Among these factors, mean annual precipitation was the primary factor controlling the variation of annual Rs among grassland types. Based on the overall data across Chinese grasslands, the Q10 values ranged from 1.03 to 8.13, with a mean (±SE) of 2.60±0.08. Moreover, the Q10 values varied largely within and among grassland types and soil temperature measurement depths. Among grassland types, the highest Q10 derived by soil temperature at a depth of 5 cm occurred in alpine grasslands. In addition, the seasonal variation of soil respiration in Chinese grasslands generally cannot be explained well by soil temperature using the van't Hoff equation. Overall, our findings suggest that the combined factors of soil temperature and moisture would better predict soil respiration in arid and semi-arid regions, highlight the importance of precipitation in controlling soil respiration in grasslands, and imply that alpine grasslands in China might release more carbon dioxide to the atmosphere under climate warming.


2020 ◽  
Vol 125 (7) ◽  
pp. 1077-1089
Author(s):  
Deborah W Woodcock ◽  
Herbert W Meyer

Abstract Background and Aims The Piedra Chamana fossil forest in northern Peru is an assemblage of angiosperm woods and leaves preserved in volcaniclastic rocks dated to 39 Mya (late Middle Eocene). We analysed the anatomical and morphological features of the fossils to reconstruct the palaeoenvironment during this time of global warmth, taking advantage of the co-occurrence of woods and leaves to compare different proxies and analytical approaches. Methods Wood characters analysed include vessel-related functional traits, traits linked to Baileyan trends, and quantitative features such as vessel diameter and density. Diameter-distribution and diameter and position plots are used to represent vessel diameter and arrangement. Leaf margin and area analysis provides additional climate estimates. Key Results The fossil woods show many similarities with modern tropical-forest woods and tropical fossil-wood assemblages; closest correspondence within the Neotropics is to semi-deciduous lowland tropical forest with moderate precipitation (~1000–1200 mm). Features unusual for the modern South American tropics are mainly vessel-related characters (semi-ring porosity, grouped vessels, helical vessel thickenings, short vessel elements) linked to water stress or seasonal water availability. Leaf analysis indicates mean annual temperature of 31 °C (n = 19, 100 % entire-margined) and mean annual precipitation of 1290 mm (n = 22, predominantly microphylls and notophylls). Conclusions The palaeovegetation was clearly lowland tropical forest with a dry aspect, but anomalous aspects of the wood anatomy are consistent with the high temperatures indicated by the leaves and are probably explained by differences in seasonality and water stress compared to the present-day Neotropics. A close modern analogue may be in very seasonal regions of Asia. Pronounced monsoonal (summer-rain) conditions may relate to a location (palaeolatitude of 13°S) outside the near-equatorial tropics.


2020 ◽  
Author(s):  
Lisa Zwanzig ◽  
Martin Zwanzig ◽  
Daniela Sauer

&lt;p&gt;Soil formation is controlled by climate, vegetation, organisms, topography, parent material and time. There are various hypotheses on the relative importance of these individual soil-forming factors. The quantitative influence of each soil-forming factor on the expression and rates of soil-forming processes, and in particular the influence of the different factors in combination, have not yet been sufficiently analyzed. The aim of this study was to quantify the influence of the soil-forming factors on the rates of podzolization. For this purpose, we compiled published data from 46 soil chronosequence studies in a database. These studies contained altogether 231 soil profiles of known age, on which we tested existing hypotheses on the influence of different soil-forming factors. The formation of an E horizon and its increase in thickness over time is one of the characteristic features of Podzol formation. As it is one of the few features that was described in all 46 studies, we used it as an indicator of progressive podzolization. Through statistical analysis, we investigated how E horizon thickness is affected by latitude, longitude, mean annual precipitation, mean annual temperature, range between minimum and maximum monthly temperature, annual number of days with frost, vegetation class (pioneer, deciduous and coniferous), sand content, clay content, and soil age.&lt;/p&gt;&lt;p&gt;Since E horizon thickness exhibited a zero-inflated (semi-)continuous distribution, we opted for a zero-altered gamma (ZAG) model, consisting of a Bernoulli and a Gamma part. The Bernoulli part shows, how the probability of the presence of an E horizon changes with soil age and environmental conditions. The Gamma part of the ZAG model allows for capturing the effects of the covariates on E horizon thickness. Our results indicate that vegetation is the most important factor for both (1) the soil age at which podzolization starts (used indicator: first occurrence of an E horizon), and (2) the rates of podzolization thereafter (used measure: increase of E horizon thickness with soil age). Climatic factors such as mean annual precipitation and range of temperature play subordinate roles. They are important for the soil age at which podzolization starts but less important for the rates of podzolization. We did not identify a significant influence of sand content, neither on the start nor the rates of podzolization. Thus, this statistical assessment of global data provides new insights into the relative importance of the individual soil-forming factors on the onset and temporal course of podzolization.&lt;/p&gt;


2013 ◽  
Vol 43 (1) ◽  
pp. 103-107 ◽  
Author(s):  
Björn Berg ◽  
Chunjiang Liu ◽  
Ryszard Laskowski ◽  
Matthew Davey

Using literature data, we investigated coniferous and broadleaf litter from 58 tree species using a database encompassing concentrations of N and acid-unhydrolyzable residue (AUR) (gravimetric lignin) in newly shed litter, mean annual temperature, and mean annual precipitation. Our aims were to (i) demonstrate any large-scale relationships between concentrations of N and AUR in foliar litter and (ii) determine differences in this respect among litter from Pinus and Quercus. To this end, we had collected foliar litter data for Asia and Europe, forming a climate gradient. Litter from broadleaf and coniferous trees differed significantly in concentrations of N (p < 0.0001, 9.64 versus 5.50 mg/g, respectively) and AUR (p < 0.0001, 219 versus 292 mg/g, respectively). There were highly significant positive linear relationships between concentrations of N and AUR for broadleaf (p < 0.0001) and coniferous litter (p < 0.0001). There were also significant positive relationships for AUR as a function of N concentration for the genera Pinus and Quercus but not within species. That for Scots pine (Pinus sylvestris L.) was negative and that for common oak (Quercus robur L.) not significant.


2021 ◽  
Vol 13 (12) ◽  
Author(s):  
Sara E. Rhodes ◽  
Nicholas J. Conard

AbstractEnsuring comparability between results is a key goal of all paleoecological reconstructions. Quantitative estimates of meteorological variables, as opposed to relative qualitative descriptions, provide the opportunity to compare local paleoenvironmental records against global estimates and incrementally build regional paleoclimatic records. The Bioclimatic Method provides quantitative and qualitative estimates of past landscape composition and climate along with measures of statistical accuracy by applying linear discriminant functions analysis and transfer functions to faunal taxonomic abundance data. By applying this method to the rodent data from Geißenklösterle and Hohle Fels, two Paleolithic cave sites located in the Ach Valley of southwestern Germany, we classify the regional vegetation according to Walters’ zonobiome model. We also present new estimates of meteorological variables including mean annual temperature, mean annual precipitation, and vegetative activity period of the Ach Valley for the period spanning ~ 60,000 to 35,000 cal BP. The results suggest the Ach Valley contained a non-analogous landscape of arctic tundra and temperate deciduous woodland with occasional arid steppe expansion. Meteorological estimates suggest the climate was significantly colder during the Middle and Upper Paleolithic than today, with higher annual precipitation and dramatic temperature shifts between seasons. These results fit well with climatic reconstructions from Switzerland and the Netherlands based on a variety of proxies. They also provide further evidence of a localized climatic response within southwestern Germany to the stadial-interstadial shifts preceding the Heinrich 4 event. Finally, these results reinforce our previous claims that climatic volatility was not a driving force in the loss of Neanderthal groups throughout the Swabian Jura during OIS 3.


2020 ◽  
Vol 47 (2) ◽  
pp. 418
Author(s):  
Juan M. Robledo ◽  
Maricel Y. Horn ◽  
Claudia I. Galli ◽  
Luisa M. Anzótegui

The continental sedimentary rocks that constitute the Palo Pintado Formation of the late Miocene from Salta province, presents a great paleoclimatic interest due to the environmental conditions prevailing during this geochronologic interval. The geological and paleobotanical data suggest that during the sedimentary rocks accumulation of the Palo Pintado Formation (Angastaco Basin), wetter conditions would have existed comparing with other nearby and contemporary Formations, for example the Playa del Zorro Aloformation (late Miocene of Catamarca) and the Chiquimil (late Miocene of Tucumán), Salicas and the Toro Negro Formations (both from the late Miocene of La Rioja). In this study, the margin and the foliar area of the leaves contained on rocks from the Palo Pintado Formation are analyzed, in order to obtain the mean annual temperature (MAT) and the mean annual precipitation (MAP). The resulting values were: 23.98 °C and 330.8 mm. These results are coincident by the interpretation of different authors, who consider that the Palo Pintado Formation would have been deposited under a relatively humid environment, possibly as a consequence of the rains that affected locally the Angastaco basin región.


IAWA Journal ◽  
1990 ◽  
Vol 11 (3) ◽  
pp. 261-280 ◽  
Author(s):  
Sergio R. S. Cevallos-Ferriz ◽  
Ruth A. Stockey

Several anatomieally preserved twigs, a branehing speeimen and the wood of a large axis with affinities to Rosaeeae are deseribed from the Prineeton ehert (Middle Eoeene) of British Columbia, Canada. Speeimens are eharaeterised by a heteroeellular pith with a peri-medullary rone of thiek-walled oval eells and semi-ring-porous seeondary xylem with vertieal traumatie duets, fibres with eireular bordered pits, and mostly seanty paratracheal and oeeasionally apotracheal parenehyma. Ray to vessel pitting is similar to the alternate intervaseular pitting. Seeondary phloem is eomposed of tangentially oriented diseontinuous bands of alternating fibres and thinwalled eells. Seeondary eortical tissues are represented by a phelloderm eharaeterised by rectangular eells and phellern with rectangular eoneave eells. Anatomical variation between speeimens can be related to age of the woody axes. Juvenile and mature wood of this speeies differ in vessel arrangement and presenee of scanty paratracheal parenchyma in mature wood. Vessel elements are arranged in radial multiples, oeeasional clusters as well as solitary vessels. Tyloses and dark cellular contents are present mainly in mature wood. Some twigs have a heterocellular pith with a few scattered cells with dark contents or, occasionally, short irregular rows of these cells. In the branching specimen eells of this type also are organised in longer rows. Together, these anatomical features suggest that all specimens belong to the same taxon, Prunus allenbyensis Cevallos-Ferriz ' Stockey n. sp. Anatomy of this plant reinforces the interpretation of a subtropical to temperate climate during the time of deposition.


Palaios ◽  
2019 ◽  
Vol 34 (9) ◽  
pp. 424-436 ◽  
Author(s):  
ANTHONY P. JIJINA ◽  
ELLEN D. CURRANO ◽  
KURT CONSTENIUS

ABSTRACT New collections of plant macrofossils and radiometric dates from the Herren beds of north-central Oregon provide the opportunity to document floral communities and calculate foliar-derived climate estimates from the warm early Eocene and the cooler middle Eocene. Plant macrofossils were collected from one fluvial site at East Birch Creek approximately 2 m below a 51.9 ± 0.9 Ma tuff. Collections were also made at two co-occurring fluvial sites at Arbuckle Mountain, whose ages are constrained to ca. 44.5–43.8 Ma based on a dated tuff from Willow Creek (44.5 ± 0.8 Ma) and reported ages for the overlying Clarno Formation. Floral findings show an almost complete vegetation overturn, with only two genera (Glyptostrobus and Allantodiopsis) appearing in both floras. Both floras are species poor, but the older East Birch Creek flora has higher richness and evenness than the younger Arbuckle Mountain flora. The four named genera at East Birch Creek are taxa found throughout Eocene North America; named genera at Arbuckle Mountain also include taxa restricted to the Pacific Northwest. Leaf margin analysis and leaf area analysis of the East Birch Creek community suggest a warmer and possibly wetter (mean annual temperature 23.4 ± 4.3 °C; mean annual precipitation 206 +89, -63 cm) climate than the Arbuckle Mountain flora (16.4 ± 4.2 °C; 165 +50, -71.4 cm). This research provides a framework for future research on Eocene floristic, environmental, and climatic trends of the Pacific Northwest.


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