Evolution in sown populations of subterranean clover (Trifolium subterraneum L. ) in South Australia

1992 ◽  
Vol 43 (7) ◽  
pp. 1583 ◽  
Author(s):  
PS Cocks

The seed banks of three pastures at Kingscote, Parndana and Waterloo, S.A., were sampled to determine the frequency of divergent genotypes in subterranean clover, and the direction of natural selection. The seeds were grown in nursery rows at Adelaide, and the resulting plants classified into one of the commercial cultivars, or as divergent genotypes. The divergent genotypes from one locality, Kingscote, were described in terms of 17 variables, and compared with strains collected from southern Australia and the Mediterranean basin. At each locality there was more genetic diversity than had been sown, both in terms of additional cultivars and the presence of divergent genotypes. The percentage of divergent genotypes appeared to be proportional to the age of the pasture, and was greatest at Kingscote, where it reached 67% of the population. The Kingscote genotypes were genetically related to Mt Barker and Dwalganup, only five out of 283 genotypes having leaf markings that differed from both cultivars. The 17 variables fell between the values recorded for Mt Barker and Dwalganup, and there was evidence of directional selection. For example, formononetin content was less, and genistein more than would be expected in the absence of natural selection. Mean flowering time was about equal to that of Seaton Park, and closer to Mt Barker than to Dwalganup. Flowering time was related to elevation above the lowest point in the paddock, the latest genotypes tending to inhabit sites at the foots of slopes, and the earliest genotypes sites on the tops. Seed and burr weight were slightly larger than expected. Hybridization was the most important, but not the sole source of genetic divergence at Kingscote, whereas contamination was the main source at Waterloo. It is likely that only a minority of the divergent genotypes was generated by mutation. The Australian subterranean clovers were as diverse as those from the Mediterranean. Those from Kingscote were also diverse, but, on the whole, had longer peduncles, shorter internodes and were taller than strains from other parts of Australia and from the Mediterranean basin.

2013 ◽  
Vol 140 (682) ◽  
pp. 1563-1577 ◽  
Author(s):  
P. Martinet ◽  
L. Lavanant ◽  
N. Fourrié ◽  
F. Rabier ◽  
A. Gambacorta

1987 ◽  
Vol 27 (4) ◽  
pp. 551 ◽  
Author(s):  
SG Clark ◽  
JR Hirth

Forty-two midseason-late maturing subterranean clover genotypes from the Mediterranean basin, 12 commercial cultivars, 2 bulk treatments and a blank control were sown in small plots in 1983 at the Pastoral Research Institute, Hamilton, Vic. (10.5 m2), and at the Rutherglen Research Institute, Rutherglen, Vic. (7.0 m2), to assess seasonal yield and persistence over 3 years. Perennial ryegrass cv. Ellett was sown as a companion grass at Hamilton. At both sites, but particularly at Hamilton, several genotypes were identified which showed considerable improvement over commercial cultivars. Some genotypes, although late maturing, were as productive as the best commercial cultivars in autumn and winter (e.g 2.06-2.38 t DM/ha) and were also highly productive (e.g. 3.46-3.94 t DM/ha) in spring (e.g. CPI 89822 H, 89774 F [Sardinia], 68103 H [Portugal] and GF 073.2 [Corsica] at Hamilton and CPI 89880 F, 89855 G, 89880 J [Sardinia] and GF 144.3 [France] at Rutherglen). Seed production of some genotypes was superior to that of the commercial cultivars (P= 0.01). Trikkala, Larisa and Karridale were the cultivars with the largest seed banks at both sites at the end of the third year. Nine of the 13 genotypes measured had seed banks at least as large as Trikkala (36 1 kg/ha) at Hamilton (e.g. 89774 F, 431 kg/ha; 89777 C, 402 kg/ha). All 10 genotypes measured at Rutherglen had seed banks at least as large as Karridale (1298 kg/ha) (e.g. 89880 J, 1717 kg/ ha; and 89820 D, 1578 kg/ha). As a result of this work and similar studies in Western Australia, New South Wales, South Australia and Tasmania, 13 late-maturing genotypes have been chosen for advanced evaluation prior to cultivar release about 1990.


2013 ◽  
Vol 64 (4) ◽  
pp. 312 ◽  
Author(s):  
P. G. H. Nichols ◽  
K. J. Foster ◽  
E. Piano ◽  
L. Pecetti ◽  
P. Kaur ◽  
...  

Subterranean clover (Trifolium subterraneum L.) is the most widely sown annual pasture legume species in southern Australia, valued in the livestock and grains industries as a source of high-quality forage and for its ability to fix atmospheric nitrogen. From its initial accidental introduction into Australia in the 19th Century and subsequent commercialisation in the early 1900s, 45 cultivars have been registered in Australia. These consist of 32 cultivars of ssp. subterraneum, eight of ssp. yanninicum, and five of ssp. brachycalycinum and range in flowering time from 77 to 163 days from sowing, enabling the species to be grown in a diversity of rainfall environments, soil types, and farming systems. Eleven of these cultivars are introductions from the Mediterranean region, 15 are naturalised strains collected in Australia, 18 are the products of crossbreeding, and one is derived from mutagenesis. Cultivars developed in Italy have been commercialised for the local market, whereas other cultivars developed in Spain, Portugal, and France have not had commercial seed production. Important traits exploited include: (i) selection for low levels of the oestrogenic isoflavone formononetin, which causes reduced ewe fertility; (ii) increased levels of dormancy imposed by seed-coat impermeability (hard seeds) for cultivars aimed at crop rotations or unreliable rainfall environments; (iii) strong burr-burial ability to maximise seed production; (iv) resistance to important disease pathogens for cultivars aimed at medium- and high-rainfall environments, particularly to Kabatiella caulivora and root rot pathogens; (v) resistance to pests, particularly redlegged earth mites; and (vi) selection for unique leaf markings and other morphological traits (where possible) to aid cultivar identification. Cultivar development has been aided by a large genetic resource of ~10 000 accessions, assembled from its centre of origin in the Mediterranean Basin, West Asia, and the Atlantic coast of Western Europe, in addition to naturalised strains collected in Australia. The development of a core collection of 97 accessions, representing almost 80% of the genetic diversity of the species, and a genetic map, provides a platform for development of future cultivars with new traits to benefit the livestock and grains industries. New traits being examined include increased phosphorous-use efficiency and reduced methane emissions from grazing ruminant livestock. Economic analyses indicate that future trait development should focus on traits contributing to increased persistence and autumn–winter productivity, while other potential traits include increased nutritive value (particularly of senesced material), increased N2 fixation ability, and tolerance to cheap herbicides. Beneficial compounds for animal and human health may also be present within the species for exploitation.


Author(s):  
Joshua M. White

This book offers a comprehensive examination of the shape and impact of piracy in the eastern half of the Mediterranean and the Ottoman Empire’s administrative, legal, and diplomatic response. In the late sixteenth and seventeenth centuries, piracy had a tremendous effect on the formation of international law, the conduct of diplomacy, the articulation of Ottoman imperial and Islamic law, and their application in Ottoman courts. Piracy and Law draws on research in archives and libraries in Istanbul, Venice, Crete, London, and Paris to bring the Ottoman state and Ottoman victims into the story for the first time. It explains why piracy exploded after the 1570s and why the Ottoman state was largely unable to marshal an effective military solution even as it responded dynamically in the spheres of law and diplomacy. By focusing on the Ottoman victims, jurists, and officials who had to contend most with the consequences of piracy, Piracy and Law reveals a broader range of piratical practitioners than the Muslim and Catholic corsairs who have typically been the focus of study and considers their consequences for the Ottoman state and those who traveled through Ottoman waters. This book argues that what made the eastern half of the Mediterranean basin the Ottoman Mediterranean, more than sovereignty or naval supremacy—which was ephemeral—was that it was a legal space. The challenge of piracy helped to define its contours.


Author(s):  
Matthew D. C. Larsen

The concept of textual unfinishedness played a role in a wide variety of cultures and contexts across the Mediterranean basin in antiquity and late antiquity. Chapter 2 documents examples of Greek, Roman, and Jewish writers reflecting explicitly in their own words about unfinished texts. Many writers claimed to have written unfinished texts on purpose for specific cultural reasons, while others claimed to have written texts that slipped out of their hands somehow with their permission.


Author(s):  
Madadh Richey

The alphabet employed by the Phoenicians was the inheritor of a long tradition of alphabetic writing and was itself adapted for use throughout the Mediterranean basin by numerous populations speaking many languages. The present contribution traces the origins of the alphabet in Sinai and the Levant before discussing different alphabetic standardizations in Ugarit and Phoenician Tyre. The complex adaptation of the latter for representation of the Greek language is described in detail, then some brief attention is given to likely—Etruscan and other Italic alphabets—and possible (Iberian and Berber) descendants of the Phoenician alphabet. Finally, it is stressed that current research does not view the Phoenician and other alphabets as inherently simpler, more easily learned, or more democratic than other writing systems. The Phoenician alphabet remains, nevertheless, an impressive technological development worthy, especially by virtue of its generative power, of detailed study ranging from paleographic and orthographic specifications to social and political contextualization.


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