Response to selection in Australian Merino sheep. VIII.* Further results on selection for high clean wool weight with attention to quality

1978 ◽  
Vol 29 (3) ◽  
pp. 615 ◽  
Author(s):  
HN Turner ◽  
N Jackson

Results of selection for high clean wool weight per head with control of quality are reported for two selection groups over the period 1966–74. Results for the same experiment for the periods 1950–1959 and 1961–64 were reported earlier. Both groups were selected for high clean wool weight, one (S) with a ceiling on fibre diameter and degree of skin wrinkle, and the other (MS) with a lower limit on staple crimp frequency and a ceiling on skin wrinkle. Genetic progress in clean wool weight was greater in S than in MS over the 1966–74 period (0.12–0.15 lb/annum, compared with 0.06–0.09). This was to be expected from genetic correlations of clean wool weight with fibre diameter (low positive) and staple crimp frequency (high negative). The result supports the previous recommendation that staple crimp frequency is an inefficient way of controlling wool quality while attempting to improve quantity by selection, because its use severely restricts the likely progress in quantity. The actual rate of progress in the S group was similar to that in the period 1950–59, which was followed by a fall in superiority of the selected over the control group animals born during 1961–64. The recovery of response in the 1966–74 period negates the suggestion that the loss of response during the 1961-64 period was due to a 'plateau'. The most likely explanation is that a genotype x environment interaction occurred, such that the genetic gain made in the 1950-59 period could not be expressed in the poorer environments of 1960–65, but reappeared gradually under the improving environment of the 1966–74 period. Attempts to remove this interaction by regression of response on the mean clean wool weight of the unselected control group (as an index of the level of the environment) for each year, were not successful. The interaction is, therefore, not simply a case of all selection groups being equal when the environment is poor. ________________ *Part VII, Aust. J. Agric. Res., 26: 937 (1975).

1963 ◽  
Vol 14 (4) ◽  
pp. 460 ◽  
Author(s):  
SSY Young ◽  
HN Turner ◽  
CHS Dolling

Phenotypic and genetic parameters for fertility in sheep, in terms of number of lambs born and number weaned, have been estimated in a flock of medium Peppin Merinos. Repeatability of fertility traits over all ages is low, but there is an age effect, the record at 3 years of age having a higher value than those at 2 or 4 years. The regression of subsequent performance on a difference of 1 lamb at the initial lambing was higher for the difference between 1 and 2 lambs than between 0 and 1 lamb, which indicated that selection for twins is likely to raise fertility in the current flock more rapidly than selection against barrenness. The estimate of heritability for 2 years of age was negligible for each fertility trait, but the estimates for the 3-year-old record were higher, the value for lambs born being over 0.3. Mass selection for number of lambs born at 3 years of age would thus be expected to lead to appreciable genetic progress, while there would be at least some progress in number of lambs weaned. Phenotypic and genetic correlations were also estimated between fertility at different ages and 10 sheep and wool traits measured at 15–16 months of age. Phenotypically, fertility is positively correlated with body weight and negatively with skin wrinkle score. Genetically, it is suggested from the estimates that fertility is positively correlated with body weight and staple length and negatively correlated with fibre diameter, clean scoured yield, and wrinkle score. No phenotypic or genetic correlation was found between greasy or clean wool weight and either measure of fertility. The application of the findings to breeding for higher fertility is discussed. In an appendix, a maximum likelihood method of estimating heritability for all-or-none traits in half-sib data is presented.


2002 ◽  
Vol 53 (3) ◽  
pp. 271 ◽  
Author(s):  
S. W. P. Cloete ◽  
J. C. Greeff ◽  
R. P. Lewer

(Co)variance estimates for hogget liveweight, greasy fleece weight, clean fleece weight, clean yield, fibre diameter, and the coefficient of variation of fibre diameter were obtained for a Western Australian Merino resource flock. The flock encompassed 16 medium wool bloodlines and data were available for the period 1982–93. Direct additive genetic variances (h2) — expressed as a ratio of the total phenotypic variance within bloodlines — were estimated at 0.52 for hogget liveweight, 0.44 for greasy fleece weight, 0.42 for clean fleece weight, 0.63 for clean yield, 0.71 for fibre diameter, and 0.62 for coefficient of variation of fibre diameter. Maternal genetic variance estimates were significant (P < 0.05) only in hogget liveweight and fibre diameter, but components within bloodlines were low (0.05 for liveweight and 0.02 for fibre diameter). Direct within-bloodline genetic correlations of hogget liveweight as well as greasy and clean fleece weight with fibre diameter were positive (0.17, 0.31, and 0.31, respectively), suggesting that selection for bigger and heavier cutting sheep would generally lead to a broader fibre diameter. Liveweight was unrelated to clean yield and negatively related to coefficient of variation of fibre diameter (–0.17). Greasy fleece weight was negatively related to clean yield (–0.20). The genetic correlation of clean fleece weight with clean yield was positive (0.37). Wool quantity was, in general, positively related to coefficient of variation of fibre diameter, although the estimated genetic correlations were low (0.12 for greasy fleece weight and 0.07 for clean fleece weight). The genetic correlation between fibre diameter and coefficient of variation of fibre diameter was negative, and fairly low (–0.10). These results are discussed with reference to sheep breeding.


1970 ◽  
Vol 21 (5) ◽  
pp. 837 ◽  
Author(s):  
N Jackson ◽  
JW James

Data from two-tooth rams and ewes representing seven Australian Merino studs were analysed to provide estimates of between-stud genetic variances and between-stud genetic correlations for 20 wool and body traits. The estimates were used to compare two methods of choosing foundation animals for a new stud: selection within one stud or selection within each of several studs. Where only one trait was considered in selection, and provided that accurate estimates of stud mean breeding values were available, selection from a single stud was superior, although there were some differences between traits in the degree of superiority. Where more than one trait was considered the conclusion depended on the relative magnitudes and signs of the between and within-stud genetic and phenotypic correlations. In the particular case of selection for high clean fleece weight and fine fibre diameter, a strong unfavourable between-stud genetic correlation shifted the emphasis more toward selection from several studs, but selection from a single stud was still superior when accurate estimates of stud mean breeding values for clean fleece weight were available. When response to subsequent selection, as well as immediate gain in choice of founders, was considered, the conclusions were reversed. For a single trait, selection from several studs was always superior in the long term (three or more generations), and also in the short term when accurate estimates of stud mean breeding values were not available.


1966 ◽  
Vol 17 (4) ◽  
pp. 557 ◽  
Author(s):  
GH Brown ◽  
HN Turner ◽  
SSY Young ◽  
CHS Dolling

Estimates were made of the effects of the following factors on 10 fleece and body characteristics measured on breeding ewes aged 1½ to 10½ years in three mating groups over a period of 15 years: age of ewe, single or twin birth, age of dam, the ewe's own lambing performance, the year in which measurements were made, and the year in which each set of ewes was born. Two groups (S and MS) were under selection for high clean wool weight at 15–16 months, with a ceiling on wrinkle score and fibre diameter, while the third (C) was a random control. Changes with age were present in all characteristics and were similar in the three groups. The finding that selection on wool weight at an early age had no effect on subsequent age changes in any characteristic is of considerable importance. Greasy and clean wool weight reached a maximum at 34 years, then declined by 0.3–0.2 1b per year. Percentage clean yield, fibre diameter, body weight, and wrinkle score had maxima at 5½ to 6½ years. Staple length fell consistently by approximately 0.2 cm per year, while face cover rose consistently but slightly. Crimp number rose, fell, and rose again, while fibre number rose, fell, and remained constant from 4½ years. The chief source of increase in wool weight from l½ to 3½ years was an increase in the total number of fibres. The chief source of the subsequent fall was a decrease in fibre volume, with a minor contribution from a fall in total fibre number after 6½ years. Twin-born ewes cut 0.21 lb (4.2% of the mean) less clean wool per year over their lifetime than single-born ewes, while the progeny of 2-year-old ewes cut 0.32 lb (6.4%) less than the progeny of adults. The main source of lower weight in each case was a lower total fibre number. Pregnancy lowered clean wool weight more than lactation, the separate effects being 0.87 and 0.38 lb respectively (17.4 and 7.7% of the mean) and the combined effect 1.25 1b or 25.1%. Pregnancy lowered total fibre number but lactation had no further effect. Mean clean wool weights over all ages in the C group varied from year to year, the range being from 1.08 lb (21.6%)below the mean to 0.97 lb (19.4%) above. Differences in total fibre number contributed between one-third and two-thirds of the variation. Ewes born in consecutive years in the S and MS groups showed marked upward trends in clean wool weight, fibre number, and staple length, with a marked downward trend in crimp number and a slight upward trend in body weight. These trends demonstrate direct and correlated responses to the strong selection for high clean wool weight at 15–16 months of age, and the associated slight selection against fibre diameter and wrinkle score. The mean annual increases in clean wool weight were 0.15 and 0.11 Ib (3.0 and 2.2%) in the S and MS groups, approximately 40% of the increase arising from increased total fibre number and 40% from increased staple length. The effects of age and lambing performance can be used to predict productivity in flocks of differing age structures. As the casting age rises to 54 years changes in productivity are negligible. With a rise in casting age to 7½ years the average clean wool weight of the flock would fall by 0.14 lb, with a slight decrease in staple length and crimp number. These changes need to be balanced against any increased lambing percentage or decreased annual genetic gain due to increased generation interval. Comparison with other available figures indicates that age changes may vary from one area to another.


2016 ◽  
Vol 56 (1) ◽  
pp. 87 ◽  
Author(s):  
Andrew A. Swan ◽  
Daniel J. Brown ◽  
Julius H. J. van der Werf

Genetic variation within and between Australian Merino subpopulations was estimated from a large breeding nucleus in which up to 8500 progeny from over 300 sires were recorded at eight sites across Australia. Subpopulations were defined as genetic groups using the Westell–Quaas model in which base animals with unknown pedigree were allocated to groups based on their flock of origin if there were sufficient ‘expressions’ for the flock, or to one of four broad sheep-type groups otherwise (Ultra/Superfine, Fine/Fine-medium, Medium/Strong, or unknown). Linear models including genetic groups and additive genetic breeding values as random effects were used to estimate variance components for 12 traits: yearling greasy and clean fleece weight (ygfw and ycfw), yearling mean and coefficient of variation of fibre diameter (yfd and ydcv), yearling staple length and staple strength (ysl and yss), yearling fibre curvature (ycuv), yearling body wrinkle (ybdwr), post-weaning weight (pwt), muscle (pemd) and fat depth (pfat), and post-weaning worm egg count (pwec). For the majority of traits, the genetic group variance ranged from approximately equal to two times larger than the additive genetic (within group) variance. The exceptions were pfat and ydcv where the genetic group to additive variance ratios were 0.58 and 0.22, respectively, and pwec and yss where there was no variation between genetic groups. Genetic group correlations between traits were generally the same sign as corresponding additive genetic correlations, but were stronger in magnitude (either more positive or more negative). These large differences between genetic groups have long been exploited by Merino ram breeders, to the extent that the animals in the present study represent a significantly admixed population of the founding groups. The relativities observed between genetic group and additive genetic variance components in this study can be used to refine the models used to estimate breeding values for the Australian Merino industry.


1970 ◽  
Vol 21 (6) ◽  
pp. 955 ◽  
Author(s):  
HN Turner ◽  
MG Brooker ◽  
CHS Dolling

Direct and correlated responses are reported for 10 traits in eight pairs of lines under selection for high and low values of single characteristics : clean wool weight per head, clean wool weight per unit skin area (measured only in the groups under selection for it), body weight, wrinkle score, fibre number per unit skin area (adjusted for body weight), fibre diameter, staple length, and percentage clean yield. All groups showed overall divergence in the character under selection, though response was not always symmetrical, and in two cases (long staple and low fibre number) there was no continuing response after the initial response to extreme selection of the base parents. In all except one case, whenever previous estimates of genetic correlation were at the level of 0.2 or higher, correlated responses were in the predicted direction, though not always symmetrical in magnitude. The exception was a previously reported negative correlation between fibre number and staple length, which was not exhibited under selection for either character. Where two characters of a pair were under selection, reciprocal responses agreed in sign; magnitude will be examined in a later paper. Wool per unit area was shown to have a greater influence on clean wool weight than that of surface area, with staple length on this occasion being the most important component of wool per unit area. Reasons for an apparent decrease in the relative importance of fibre number are discussed. Three other points of importance in sheep breeding are again emphasized by these results : 1. Crimp in several pairs of groups was a poor indicator of fibre diameter. 2. Increases in clean wool weight were associated with falls in crimp number, even though fibre diameter actually became finer; diameter, not crimp, should therefore be used as a guide to wool quality. 3. The absence of genetic correlations DL and NL indicates that selection can be for high N (fibre number) with high L (staple length) and low D (fibre diameter) -in other words, a desirable fleece -with no impeding genetic correlations.


1970 ◽  
Vol 10 (46) ◽  
pp. 544 ◽  
Author(s):  
PD Mullaney ◽  
ID Sanderson

The economic weights of various quality traits of both Merino and crossbred wool types hive been estimated. Australian averages of clean wool prices for Australian Wool Board wool types, into which mid-side samples of fleece were classified, have been used.For each of fifteen seasons, 1953-54 to 1967-68, the mean clean wool price for each wool type was expressed as a percentage of the mean wool price for all types in that season. Rank correlations for price-type means were high between seasons, varying between 0.91 and 0.98. Further analyses were therefore performed using prices averaged for the 15 seasons. Multiple regression analyses were carried out, the dependent variable being the mean price for each fleece type, expressed as a percentage of the average price of all types for al) seasons and the independent variables being the measurement of fibre diameter, crimps per inch and length, and the subjective gradings of colour, handle, character, and quality number. For both Merino and crossbred wool types, the major determinants of price were quality number, colour, and handle, with quality number being of overwhelming importance. The remaining variables were not important as joint determinants of price, and were deleted from the analyses. Quality number, colour, and handle controlled about 81 and 86 per cent of the variation in price for Merino and crossbred wool types. It is likely that most progress in selection for wool production would be made by incorporating these findings into a selection index.


2017 ◽  
Vol 47 (4) ◽  
Author(s):  
Flavia de Almeida Lucas ◽  
Adriana Lima Kandrotas ◽  
Eugenio Nardin Neto ◽  
Carlos Eduardo de Siqueira ◽  
Gabriela Santo André ◽  
...  

ABSTRACT: The aim of this study was to evaluate the effects of 10% copaiba oil in experimentally induced wounds in horses. Four wounds were made in the lumbar and metacarpal regions of eight adult horses. In the treatment group, the wounds received 10% copaiba oil and in the control group 0.9% sodium chloride, in the daily dressing for 21 days. The wounds were evaluated three, 7, 14, and 21 days postoperatively. No significant differences were observed between the groups. The mean lumbar wound contraction rates were 80.54% and 69.64%, for the control and treated groups, respectively. For the wounds in the metacarpal region, these averages were 44.15% and 52.48%, respectively. Under the experimental conditions of the present study, it is concluded that 10% copaiba oil has beneficial in wound healing in the equine species and suggest that copaiba oil can be used as a therapeutic possibility in equine wound therapy.


1955 ◽  
Vol 6 (1) ◽  
pp. 77 ◽  
Author(s):  
FHW Morley

Estimates of heritability, genetic correlations, phenotypic correlations, and other parameters among eight characteristics were obtained from Merino sheep at Trangie Agricultural Experiment Station. One set of estimates was obtained by parent-offspring regressions from data obtained between 1943 and 1952. Another set was obtained by half-sib analyses from data from sheep with first adult shearings in 1953. Estimates of heritability were: greasy fleece weight, 0.40 ± 0.06; yield (per cent.), 0.39 ± 0.07; clean fleece weight, 0.47 ± 0.07; staple length, 0.56 � 0.07; crimps per inch, 0.47 ± 0.11; folds, 0.50 ± 0.07; body weight, 0.36 ± 0.08; birthcoat, 0.80 ± 0.10. The most important genetic correlation limiting genetic improvement in clean fleece weight is that between this trait and crimps per inch. Because of this the rate of improvement in fleece weight could be reduced by about 30 per cent. whenever crimps per inch is also selected. Other genetic correlations indicate antagonisms between characteristics, but these antagonisms appear to be of minor importance except that between clean fleece weight and crimps per inch. This sheep population does not appear to have reached a state of genetic homeostasis, a t least with respect to the characteristics studied. However, the findings emphasize the necessity for careful evaluation of the economic importance of different traits, and for accurate measurement, if maximum genetic progress is to be achieved.


1997 ◽  
Vol 65 (2) ◽  
pp. 275-283 ◽  
Author(s):  
R. E. Crump ◽  
C. S. Haley ◽  
R. Thompson ◽  
J. Mercer

AbstractEstimates of heritabilities, common litter of birth effects and additive maternal genetic effects were produced for ultrasonic backfat depth, average daily food intake, average daily gain and food conversion ratio of Landrace boars and gilts. Boars and gilts were performance tested under different regimes. A bivariate derivative-free restricted maximum likelihood procedure was used to estimate genetic correlations between the performance test traits as recorded in the two sexes.Heritability estimates from the analysis including the common litter of birth effect tended to be towards the low end of the range of recently published estimates. This may reflect either population specific effects, such as effects of long-term selection, or the use of an individual animal model.Estimates of the common litter of birth effect were around 0·05, and generally had a significant effect upon the fit of the model, while additive maternal genetic effect estimates were negligible. Therefore, it is expected that omission of maternal effects from models for evaluation by best linear unbiased prediction will not hinder genetic progress. Inclusion of common litter of birth effects would be recommended, although this result may not hold for populations given food ad libitum.The estimates of genetic correlations between performance test traits of boars and gilts indicate that the levels of genotype-environment interaction (G × E) and genotype-sex interaction were low across most traits and data sets, with all genetic correlation estimates lying between 0·8 and 1·0. The lowest estimates of the genetic correlations, which were observed in data sets where the environments appeared to differ most, indicate that G × E interactions may be a problem in populations where males and females are subject to test regimes with greater differences than those seen here.


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