scholarly journals Fourteen new species ofOecetisMcLachlan, 1877 (Trichoptera: Leptoceridae) from the Neotropical region

PeerJ ◽  
2017 ◽  
Vol 5 ◽  
pp. e3753 ◽  
Author(s):  
Fabio B. Quinteiro ◽  
Ralph W. Holzenthal
Keyword(s):  
New Species ◽  
Neotropical Region ◽  
Valid Species ◽  
Light Traps ◽  
Larval Stages ◽  
The Past ◽  
Biogeographical Region ◽  
Species Groups ◽  
Malaise Traps ◽  
Maxillary Palps

BackgroundThe caddisfly genusOecetiscurrently contains 534 valid species. Its larval stages are found in freshwaters around the world. The adults can be distinguished from other Leptoceridae by the unbranched forewing M vein and the exceptionally long maxillary palps. In the Neotropical region, 55 species ofOecetishave been recorded and most of them can be placed in one of the six species groups known from this biogeographical region: theavara-,falicia-,inconspicua-,punctata-,punctipennis-, andtestacea-groups. More than 50% of the known diversity of NeotropicalOecetishas been described in the past 40 years. Here, we describe an additional 14 new species ofOecetisto further document the diversity of this genus in the Neotropical region.MethodsThe descriptions and illustrations presented here are based on male specimens. Specimens were collected with Malaise traps or ultraviolet light traps. They were preserved in alcohol or pinned as stated in material examined section. Specimens had their genitalia prepared in 85% lactic acid to better observe internal characters and illustrations were aided by the use of a microscope with drawing tube attached.Results and DiscussionThis study raises the number of species ofOecetisin the Neotropics from 55 to 69. Eight of the new species presented here could not be reliably placed in one of the known species groups (Oecetis acuticlaspern. sp.,Oecetis flintin. sp.,Oecetis carinatan. sp.,Oecetis cassicoleatan. sp.,Oecetis blahnikin. sp.,Oecetis gibbosan. sp.,Oecetis licinan. sp., andOecetis pertican. sp.). The others are placed in thepunctata-group (Oecetis bidigitatan. sp.,Oecetis quasipunctatan. sp.),testacea-group (Oecetis plenuspinosan. sp.), andfalicia-group (Oecetis calorin. sp.,Oecetis hastapullan. sp.,Oecetis machaeran. sp.). Most of the diagnostic characters rely on structures of the inferior appendages and phallic apparatus, and the shape of tergum X.

Review of Gonatocerus (Hymenoptera: Mymaridae) in the Neotropical region, with description of eleven new species

Zootaxa ◽  
2010 ◽  
Vol 2456 (1) ◽  
pp. 1 ◽  
Author(s):  
SERGUEI V. TRIAPITSYN ◽  
JOHN T. HUBER ◽  
GUILLERMO A. LOGARZO ◽  
VLADIMIR V. BEREZOVSKIY ◽  
DANIEL A. AQUINO
Keyword(s):  
New Species ◽  
Costa Rica ◽  
Dominican Republic ◽  
Neotropical Region ◽  
Species Group ◽  
The Other ◽  
Valid Species ◽  
Neotropical Species ◽  
Distribution Ranges ◽  
Species Groups

The described Neotropical species of the common and speciose fairyfly genus Gonatocerus Nees ab Esenbeck (Hymenoptera: Mymaridae) are reviewed and re-diagnosed. Eighty-four valid species are recognized including 11 newly described ones. Illustrated identification keys are provided to identify the 5 subgenera recognized in Gonatocerus and to females of each subgenus. The keys include 79 of the treated species, the other five species are known from males only. The known distribution ranges of the species, including new records of extralimital specimens examined, are given. Host associations of the treated species are also given, with emphasis on those species that parasitize eggs of proconiine sharpshooters (Hemiptera: Cicadellidae: Cicadellinae: Proconiini), all of which belong to the ater and morrilli subgroups of the ater species group of G. (Cosmocomoidea Howard), stat. rev. The other four recognized subgenera within Gonatocerus are the nominate subgenus, G. (Gonatocerus) (= sulphuripes species group of authors), G. (Lymaenon Walker), stat. rev. (= litoralis and straeleni species groups of authors), G. (Gastrogonatocerus Ogloblin), stat. rev. (= membraciphagus species group of authors), and G. (Gahanopsis Ogloblin), syn. n. & stat. n. (= deficiens species group of authors). Newly treated as synonyms of G. (Lymaenon) are Rachistus Foerster, Oophilus Enock, Agonatocerus Girault, Gonatoceroides Girault, and Decarthrius Debauche, all syn. n. Two species groups are recognized within G. (Cosmocomoidea) in the New World: the ater group, to which most Neotropical species belong (with the ater, bucculentus, morrilli, and the newly defined chusqueicolus subgroups) and the masneri group, with two described species from the Dominican Republic. The 11 new species described are G. (Gahanopsis) arkadak Triapitsyn sp. n. (Colombia), G. (Cosmocomoidea) barbos Triapitsyn sp. n. (Costa Rica and Mexico), G. (Cosmocomoidea) blefuscu Triapitsyn sp. n. (Costa Rica), G. (Cosmocomoidea) cuscus Triapitsyn sp. n. (Peru), G. (Cosmocomoidea) garchamp Triapitsyn sp. n. (Argentina), G. (Cos- mocomoidea) gerasim Triapitsyn sp. n. (Mexico), G. (Cosmocomoidea) hispaniolus Triapitsyn & Huber sp. n. (Dominican Republic), G. (Cosmocomoidea) kiskis Triapitsyn sp. n. (Argentina), G. (Cosmocomoidea) logarzoi Triapitsyn sp. n. (Argentina), G. (Cosmocomoidea) mumu Triapitsyn sp. n. (Argentina), and G. (Cosmocomoidea) rakitovi Triapitsyn sp. n. (Costa Rica). Seven new synonymies are proposed: Gonatocerus h-luteum (Ogloblin) syn. n. under G. (Cosmoco- moidea) nigriflagellum (Girault); G. enicmophilus Huber syn. n. and G. necator (Ogloblin) syn. n. under G. (Cosmoco- moidea) bonariensis (Brèthes); G. dimorphus (Ogloblin) syn. n. and G. monrosi (Ogloblin) syn. n. under G. (Gastrogonatocerus) margiscutum Girault; and G. setulosus (Ogloblin) syn. n. and G. dorsiniger (Ogloblin) syn. n. under G. (Gastrogonatocerus) membraciphagus Ogloblin. Litus maculipennis Ashmead, Gahanopsis deficiens (Ogloblin), and Gahanopsis straeleni (Debauche) are transferred to Gonatocerus as, respectively, G. (Cosmocomoidea) maculipennis (Ashmead) comb. n., G. (Gahanopsis) deficiens (Ogloblin) comb. n., and G. (Lymaenon) straeleni (Debauche) comb. n. Lectotypes are designated for the following 23 species: G. (Gahanopsis) acanophorae (Ogloblin), G. (Gahanopsis) aethalionis (Ogloblin), G. (Cosmocomoidea) annulicornis (Ogloblin), G. (Gastrogonatocerus) anomocerus Crawford, G. (Gonatocerus) appendiculatus (Ogloblin), G. (Gonatocerus) bonaerensis (Ogloblin), G. (Cosmocomoidea) caudatus (Ogloblin), G. (Cosmocomoidea) concinnus (Ogloblin), Lymaenon (Gastrogonatocerus) dimorphus Ogloblin, G. (Gona- tocerus) excisus (Ogloblin), G. (Cosmocomoidea) gracilicornis (Ogloblin), G. (Cosmocomoidea) grandis (Ogloblin), Lymaenon h-luteum Ogloblin, G. (Cosmocomoidea) inauditus (Ogloblin), G. (Gastrogonatocerus) juvator Perkins, G. (Gastrogonatocerus) margiscutum Girault, G. (Cosmocomoidea) metanotalis (Ogloblin), G. (Cosmocomoidea) nasutus (Ogloblin), Lymaenon necator Ogloblin, G. (Cosmocomoidea) nigrithorax (Ogloblin), G. (Lymaenon) pratensis (Ogloblin), G. (Gonatocerus) stenopterus (Ogloblin), and G. (Gonatocerus) urocerus (Ogloblin).

scholarly journals Morphology and life cycle of a new species of Psilocladus Blanchard, 1846 (Coleoptera, Lampyridae, Psilocladinae), the first known bromeliad-inhabiting firefly

2020 ◽  
Author(s):  
Stephanie Vaz ◽  
Luiz Felipe Lima Da Silveira ◽  
Simone Policena Rosa
Keyword(s):  
New Species ◽  
Ground Level ◽  
Neotropical Region ◽  
Immature Stages ◽  
Larval Biology ◽  
Larval Stages ◽  
First Time ◽  
A New Species ◽  
Brazilian Atlantic Rainforest ◽  
Malaise Traps

Fireflies (Coleoptera: Lampyridae) are soft-bodied beetles usually associated with mesic and hydric habitats. As such, terrestrial firefly larvae are commonly found in marshy environments and stream banks, while aquatic larvae might dwell in ponds, streams, mangroves, and even brackish water. Larval biology is especially important in fireflies, as the majority of species are extremely semelparous – that is, adults rely on resources gathered during larval stages. Despite their crucial relevance in firefly biology, larvae of only near 1% firefly species have been studied, and the majority of species remain known only from adult stages. That is especially true in the Neotropical region, where they are most diverse. Here, we describe Psilocladus costae sp. nov. after the study of adults and immature stages, the latter reported for the first time for the monotypic subfamily Psilocladinae McDermott, 1964. Interestingly, adults were first obtained by rearing the larvae, the former usually fly fairly high (ca. 10 m) and are therefore seldom collected at ground level by conventional methods (e.g., active search, Malaise traps). The new species is found in the Brazilian Atlantic Rainforest, inhabiting canopy bromeliads, an unprecedented habit for fireflies.

New species of Rineloricaria (Siluriformes: Loricariidae) from the Paranaíba River basin, Brazil

Zootaxa ◽  
2021 ◽  
Vol 5072 (6) ◽  
pp. 531-540
Author(s):  
GUILHERME JOSÉ DA COSTA-SILVA ◽  
CLAUDIO OLIVEIRA ◽  
GABRIEL DE SOUZA DA COSTA E SILVA
Keyword(s):  
New Species ◽  
River Basin ◽  
Color Pattern ◽  
Neotropical Region ◽  
Valid Species ◽  
Southeastern Brazil ◽  
Dark Spot ◽  
Caudal Fin ◽  
Fin Ray ◽  
First Ray

Rineloricaria is a genus of armored catfish encompassing 67 valid species that are widely distributed throughout the Neotropical region. A new species of Rineloricaria is described from the Paranaíba River, Upper Paraná River basin, southeastern Brazil. Rineloricaria rodriquezae sp. n. is distinguished from its congeners by the combination of the following characters: caudal-fin color pattern with basal dark spot and subterminal dark bar on branched rays interspersed with a hyaline area; five series of lateral plates with two keeled in the mid-dorsal series around the insertion of the first ray of dorsal fin; and unbranched caudal-fin ray extended as long filaments.  

A new species of Imparfinis Eigenmann & Norris, 1900 (Siluriformes: Heptapteridae) from the Tapajós basin, Brazil

Zootaxa ◽  
2019 ◽  
Vol 4701 (5) ◽  
pp. 461-472
Author(s):  
ÍTHALO DA SILVA CASTRO ◽  
WOLMAR BENJAMIN WOSIACKI
Keyword(s):  
New Species ◽  
Neotropical Region ◽  
Valid Species ◽  
Distal Margin ◽  
Caudal Peduncle ◽  
The Andes ◽  
Brown Band ◽  
Northern Brazil ◽  
Uruguay River ◽  
A New Species

Imparfinis comprises 20 valid species in the Heptapteridae, being the most diverse taxonomic group of catfishes of the Nemuroglanis subclade. The genus has one of the widest geographical distributions in the neotropical region, found on both sides of the Andes, from Costa Rica to the Paraná and Uruguay river basins in Argentina. A new species of Imparfinis is described from streams of the upper Rio Tapajós and its tributary Teles Pires in northern Brazil. The new species is diagnosed from all congeners by the presence of a vertical dark brown band W-shaped at the base of the caudal-fin rays, a thick dark brown lateral stripe from the snout to the end of the caudal peduncle, dark brown head, long maxillary barbel surpassing the distal margin of the pectoral fin, and presence of 39 or 40 total vertebrae. 

An update of Micromyinae (Diptera: Cecidomyiidae) taxonomy, with descriptions of a new genus and 13 new species from Northern Europe

Zootaxa ◽  
2020 ◽  
Vol 4750 (3) ◽  
pp. 349-369
Author(s):  
MATHIAS JASCHHOF ◽  
CATRIN JASCHHOF
Keyword(s):  
New Species ◽  
Russian Federation ◽  
New Genus ◽  
Valid Species ◽  
Northern Europe ◽  
Ongoing Research ◽  
The Past ◽  
Previously Treated ◽  
The Republic ◽  
First Time

While about 100 new species of Micromyinae have been described from Sweden in the past ten years, ongoing research into the country’s fauna continues to unveil previously unknown taxa. This paper provides scientific descriptions of another 13 new species from various parts of Sweden, belonging to six different genera, including a new genus. The new taxa, all to be attributed to both authors, are named as follows: Antennardia suorkensis sp. nov., Aprionus mossbergi sp. nov., Apr. oljonsbynensis sp. nov., Ladopyris baltica gen. et sp. nov. (found also in Estonia), Monardia (M.) lapponica sp. nov., Monardia (Xylopriona) abbreviata sp. nov., Mon. (Xyl.) obscura sp. nov., Neurolyga simillima sp. nov., N. taigensis sp. nov. (found also in the Republic of Karelia, Russian Federation), Peromyia elongatula sp. nov., P. lindstroemi sp. nov. and P. sofielundensis sp. nov. Two new junior synonyms of Aprionus Kieffer, 1894 were identified: Azygotricha Plakidas, 2017 syn. nov. and Ampullomyia Plakidas, 2018 syn. nov., both introduced for Nearctic species. The generic rank of Antennardia Mamaev, previously treated as a subgenus of Monardia Kieffer, is restored. Aprionus internuntius Jaschhof, 2003, previously regarded as synonymous with A. stylifer Mamaev, 1998, is restituted as a valid species. Peromyia assimilis nom. nov. is established as the new substitute name for P. despecta Jaschhof, 2017, found to be a junior homonym of P. despecta Jaschhof, 2010. Micromyinae discovered in Sweden for the first time are Antennardia saxonica Jaschhof, Aprionus heothinos Jaschhof, Apr. internuntius, Monardia (M.) lignivora (Felt), Monardia (Trichopteromyia) relicta Jaschhof, Neurolyga venusta (Mamaev & Rozhnova), Peromyia abdita Jaschhof, and P. brandenburgensis Jaschhof. 

Sobarocephala (Diptera: Clusiidae: Sobarocesphalinae)—Subgeneric classification and Revision of the New World species

Zootaxa ◽  
2012 ◽  
Vol 3370 (1) ◽  
pp. 1
Author(s):  
OWEN LONSDALE ◽  
STEPHEN A. MARSHALL
Keyword(s):  
New Species ◽  
New World ◽  
Neotropical Region ◽  
Species Level ◽  
World Species ◽  
Old World ◽  
Neotropical Fauna ◽  
Species Groups

Sobarocephala Czerny is divided into nine newly erected species groups: the S. archisobarocephala (4 spp.), S. erwini (3spp.), S. flava (8 New World spp.; 2 Old World spp.), S. flaviseta (86 New World spp.; 1 Old World sp.), S. interrupta (86spp.), S. isla (25 spp.), S. latipennis (16 spp.), S. plumicornis (9 Old World spp.), and S. ruebsaameni (11 spp.) groups; 1species (S. setulocetra spec. nov.) remains unplaced. There are 240 species of Sobarocephala in the New World, including17 Nearctic (2 extend into the Neotropical Region) and 170 new species, some of which were previously considered va-rieties of S. humeralis Melander & Argo. The Neotropical fauna is here described, illustrated, keyed, and placed in a phy-logenetic context. Sobarocephala peruana Soós stat. nov., formerly a subspecies of S. nigronota Melander & Argo, israised to the species level, S. imitans Curran syn. nov. and S. diversipes Curran syn. nov. are synonymized with S. liturataMelander & Argo, S. annulata albiventris Soós syn. nov. is synonymized with S. annulata Melander & Argo, S. albitarsisCzerny syn. nov. is synonymized with S. humeralis, and S. pruinosa pallidor Steyskal syn. n. is synonymized with S. pru-inosa Soós. Lectotypes are designated for S. bistrigata (Kertesz) , S. lumbalis Williston, S. plumicornis Lamb and S. variegata Melander & Argo. The egg of S. uberis spec. nov. is described.

Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae)

Zootaxa ◽  
2010 ◽  
Vol 2364 (1) ◽  
pp. 1 ◽  
Author(s):  
PAULO PASSOS ◽  
RONALDO FERNANDES ◽  
RENATO S. BÉRNILS ◽  
JULIO C. DE MOURA-LEITE
Keyword(s):  
New Species ◽  
Atlantic Forest ◽  
Taxonomic Status ◽  
Taxonomic Revision ◽  
Neotropical Region ◽  
Brazilian Atlantic Forest ◽  
Mountainous Regions ◽  
Species Groups ◽  
Three New Species ◽  
Maxillary Dentition

Dipsadine snakes of the genus Atractus are endemic to the Neotropical region, occurring from Panama to Argentina. Currently, the taxonomic status of most species of the genus is unclear and previous attempts of taxonomic revisions have been local in scale. In this paper we evaluate the taxonomic status of the Brazilian Atlantic Forest species of Atractus based on meristic, morphometric, maxillary dentition, and hemipenis characters. Quantitative and qualitative analyses suggest the recognition of one new species (A. caete sp. nov.) from the state of Alagoas, another (A. francoi sp. nov.) from the mountainous regions of the states of Rio de Janeiro and São Paulo, and the synonymy of A. kangueryensis with A. thalesdelemai. Specimens previously assigned to A. taeniatus in Argentina and Brazil are here considered A. paraguayensis. A key to the Atlantic Forest Atractus is provided and three new species groups are proposed for some cisAndean Atractus, mainly on the basis of hemipenial morphology: the A. emmeli, A. maculatus, and A. pantostictus species groups.

scholarly journals Taxonomic revision of the Pheidole sikorae species group (Hymenoptera, Formicidae) from Madagascar

ZooKeys ◽  
2020 ◽  
Vol 949 ◽  
pp. 1-185
Author(s):  
Sebastian Salata ◽  
Brian L. Fisher
Keyword(s):  
New Species ◽  
Taxonomic Revision ◽  
Species Group ◽  
Species Level ◽  
Junior Synonym ◽  
Identification Key ◽  
Valid Species ◽  
Species Groups ◽  
Worker Castes

The present study represents a taxonomic revision of the Pheidole sikorae species group from Madagascar. Forty-four members of this group are recognised and described, and an illustrated identification key to this group is also presented. One species is raised to species level: P. litigiosa Forel, 1892 stat. nov.Pheidole veteratrix angustinoda Forel, 1892 syn. nov. is proposed as a junior synonym of Pheidole veteratrix Forel, 1891. Worker castes are also described and lectotypes designated for P. litigiosa Forel, 1892, P. sikorae Forel, 1891, and P. veteratrix Forel, 1891. The following 41 new species are described: P. alinasp. nov., P. ambohimangasp. nov., P. analavelonasp. nov., P. andohahelasp. nov., P. anomalasp. nov., P. anosyennesp. nov., P. antranohofasp. nov., P. beankasp. nov., P. befotakasp. nov., P. dasossp. nov., P. flavominutasp. nov., P. gracilissp. nov., P. habokasp. nov., P. havoanasp. nov., P. hazosp. nov., P. itremosp. nov., P. joffrevillesp. nov., P. kelysp. nov., P. lavasoasp. nov., P. mahamavosp. nov., P. maintysp. nov., P. mamiratrasp. nov., P. mananteninasp. nov., P. masoandrosp. nov., P. mavohavoanasp. nov., P. midongysp. nov., P. mikrossp. nov., P. mivorysp. nov., P. nitidobrunasp. nov., P. parvulasp. nov., P. parvulogibbasp. nov., P. reniranosp. nov., P. savasp. nov., P. sofiasp. nov., P. sparsasp. nov., P. tamponysp. nov., P. trichotossp. nov., P. tsaravonianasp. nov., P. vadumsp. nov., P. volontanysp. nov., and P. vonysp. nov. At present, there are 109 valid species and subspecies of Pheidole known from Madagascar, but this number is expected to increase with upcoming taxonomic revisions of the species groups not revised in this study.

A new species and the first record of the genus Micrapemon Edwards (Diptera: Keroplatidae) from Mexico

Zootaxa ◽  
2019 ◽  
Vol 4604 (3) ◽  
pp. 539
Author(s):  
HERON HUERTA
Keyword(s):  
New Species ◽  
First Record ◽  
Neotropical Region ◽  
Adult Males ◽  
A New Species ◽  
Malaise Traps ◽  
Natural Reserves

The first record of the genus Micrapemon Edwards from Mexico is presented, with the description and illustration of a new species, Micrapemon maya spec. nov., based on adult males collected with Malaise traps from Yucatan (natural reserves of Ria Celestun and Ria Lagartos). The new species from Mexico is compared with the two previously known species from the Neotropical region, Micrapemon majusculum (Edwards) and Micrapemon parvum (Williston). A key is included to the species of the genus. 

scholarly journals Taxonomy and phylogeny of New World Polyplectropus Ulmer, 1905 (Trichoptera: Psychomyioidea: Polycentropodidae) with the description of 39 new species

Zootaxa ◽  
2010 ◽  
Vol 2582 (1) ◽  
pp. 1 ◽  
Author(s):  
MARIA LOURDES CHAMORRO ◽  
RALPH W. HOLZENTHAL
Keyword(s):  
New Species ◽  
Bayesian Methods ◽  
New World ◽  
Morphological Characters ◽  
Neotropical Region ◽  
Species Group ◽  
Biogeographic Region ◽  
Parsimony Tree ◽  
Group A ◽  
Species Groups

The taxonomy of the New World species of Polyplectropus (Ulmer, 1905a) is revised to include detailed male and female diagnoses, descriptions, illustrations, distribution records, and keys to males of all species and species groups. A phylogenetic analysis based on 59 morphological characters, 89 of 92 (97%) New World Polyplectropus species, and 2 outgroup taxa was inferred using parsimony and Bayesian methods, which resulted in minor topological differences. Conflicting estimates of relationship among and within most species groups led to a less resolved Bayesian tree (vs. parsimony tree) due to high variation in rates of change among characters and an overall low number of characters. A new classification for New World Polyplectropus is proposed with revised characterization of 10 recognized species groups, 6 newly established. Four species remain unassigned to species group. A key to genera of New World Polycentropodidae, including a redescription of Polyplectropus is provided. The homology of the male genitalia of Polyplectropus is discussed. Ninety-two species are treated. The following 39 new species are described: Polyplectropus adamsae (Peru), P. alatespinus (Brazil), P. amazonicus (Brazil), P. andinensis (Argentina, Bolivia), P. blahniki (Venezuela), P. bolivianus (Bolivia), P. brasilensis (Brazil), P. brborichorum (Ecuador), P. cressae (Venezuela), P. colombianus (Colombia), P. corniculatus (Peru), P. cuzcoensis (Peru), P. ecuadoriensis (Ecuador), P. flintorum (Venezuela), P. gaesum (Brazil), P. guyanae (Guyana, Venezuela), P. hollyae (Brazil), P. hystricosus (Brazil), P. insularis (Panama), P. juliae (Brazil), P. kanukarum (Guyana), P. maculatus (Venezuela), P. manuensis (Peru), P. matatlanticus (Brazil), P. minensium (Brazil), P. novafriburgensis (Brazil), P. peruvianus (Peru), P. petrae (Brazil), P. pratherae (Brazil), P. puyoensis (Ecuador), P. robertsonae (Bolivia), P. rodmani (Brazil), P. rondoniensis (Brazil), P. tragularius (Brazil), P. tripunctatum (Peru), P. venezolanus (Venezuela), P. woldai (Panama), P. zamoranoensis (Honduras), and P. zuliae (Venezuela). Polyplectropus buchwaldi (Ulmer, 1911) is designated as a nomen dubium.Key words: phylogeny, taxonomy, Neotropics, Nearctric, Bayesian methods, parsimony, classification, cladistics, morphology, Trichoptera, Polycentropodidae, Polyplectropus, new speciesThe polycentropodid caddisfly genus Polyplectropus Ulmer contains 219 species found primarily in the tropical and subtropical regions of the world, with 125 species recorded from the Oriental biogeographic region, 53 from the Neotropical region (6 species extending their ranges into the Nearctic region), 37 from the

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