scholarly journals Prootic anatomy of a juvenile tyrannosauroid from New Jersey and its implications for the morphology and evolution of the tyrannosauroid braincase

2018 ◽  
Author(s):  
Chase D Brownstein
Keyword(s):  
Phylogenetic Analysis ◽  
North America ◽  
Northern Hemisphere ◽  
Late Cretaceous ◽  
Coastal Plain ◽  
Fossil Record ◽  
Atlantic Coastal Plain ◽  
Western Interior Seaway ◽  
Large Predator ◽  
Atlantic Coastal

Among the most recognizable theropods are the tyrannosauroids, a group of small to large carnivorous coelurosaurian dinosaurs that inhabited the majority of the northern hemisphere during the Cretaceous and came to dominate large predator niches in North American and Asian ecosystems by the end of the Mesozoic era. The clade is among the best-represented of dinosaur groups in the notoriously sparse fossil record of Appalachia, the Late Cretaceous landmass that occupied the eastern portion of North America after its formation from the transgression of the Western Interior Seaway. Here, the prootic of a juvenile tyrannosauroid collected from the middle-late Campanian Marshalltown Formation of the Atlantic Coastal Plain is described, remarkable for being the first concrete evidence of juvenile theropods in that plain during the time of the existence of Appalachia and the only portion of theropod braincase known from the landmass. Phylogenetic analysis recovers the specimen as an “intermediate” tyrannosauroid of similar grade to Dryptosaurus and Appalachiosaurus. Comparisons with the corresponding portions of other tyrannosauroid braincases suggest that the Ellisdale prootic is more similar to Turonian forms in morphology than to the derived tyrannosaurids of the Late Cretaceous, thus supporting the hypothesis that Appalachian tyrannosauroids and other vertebrates were relict forms surviving in isolation from their derived counterparts in Eurasia.

scholarly journals Prootic anatomy of a juvenile tyrannosauroid from New Jersey and its implications for the morphology and evolution of the tyrannosauroid braincase

2018 ◽  
Author(s):  
Chase D Brownstein
Keyword(s):  
Phylogenetic Analysis ◽  
North America ◽  
Northern Hemisphere ◽  
Late Cretaceous ◽  
Coastal Plain ◽  
Fossil Record ◽  
Atlantic Coastal Plain ◽  
Western Interior Seaway ◽  
Large Predator ◽  
Atlantic Coastal

Among the most recognizable theropods are the tyrannosauroids, a group of small to large carnivorous coelurosaurian dinosaurs that inhabited the majority of the northern hemisphere during the Cretaceous and came to dominate large predator niches in North American and Asian ecosystems by the end of the Mesozoic era. The clade is among the best-represented of dinosaur groups in the notoriously sparse fossil record of Appalachia, the Late Cretaceous landmass that occupied the eastern portion of North America after its formation from the transgression of the Western Interior Seaway. Here, the prootic of a juvenile tyrannosauroid collected from the middle-late Campanian Marshalltown Formation of the Atlantic Coastal Plain is described, remarkable for being the first concrete evidence of juvenile theropods in that plain during the time of the existence of Appalachia and the only portion of theropod braincase known from the landmass. Phylogenetic analysis recovers the specimen as an “intermediate” tyrannosauroid of similar grade to Dryptosaurus and Appalachiosaurus. Comparisons with the corresponding portions of other tyrannosauroid braincases suggest that the Ellisdale prootic is more similar to Turonian forms in morphology than to the derived tyrannosaurids of the Late Cretaceous, thus supporting the hypothesis that Appalachian tyrannosauroids and other vertebrates were relict forms surviving in isolation from their derived counterparts in Eurasia.

scholarly journals Dinosaurs from the Santonian–Campanian Atlantic coastline substantiate phylogenetic signatures of vicariance in Cretaceous North America

2021 ◽  
Vol 8 (8) ◽  
pp. 210127
Author(s):  
Chase Doran Brownstein
Keyword(s):  
North America ◽  
Late Cretaceous ◽  
Coastal Plain ◽  
Fossil Record ◽  
Atlantic Coastal Plain ◽  
Terrestrial Vertebrates ◽  
Small Juvenile ◽  
The Poor ◽  
Atlantic Coastal ◽  
Early Late

During the Cretaceous, diversifications and turnovers affected terrestrial vertebrates experiencing the effects of global geographical change. However, the poor fossil record from the early Late Cretaceous has concealed how dinosaurs and other terrestrial vertebrates responded to these events. I describe two dinosaurs from the Santonian to Early Campanian of the obscure North American paleolandmass Appalachia. A revised look at a large, potentially novel theropod shows that it likely belongs to a new clade of tyrannosauroids solely from Appalachia. Another partial skeleton belongs to an early member of the Hadrosauridae, a highly successful clade of herbivorous dinosaurs. This skeleton is associated with the first small juvenile dinosaur specimens from the Atlantic Coastal Plain. The tyrannosauroid and hadrosaurid substantiate one of the only Late Santonian dinosaur faunas and help pinpoint the timing of important anatomical innovations in two widespread dinosaur lineages. The phylogenetic positions of the tyrannosauroid and hadrosaurid show Santonian Appalachian dinosaur faunas are comparable to coeval Eurasian ones, and the presence of clades formed only by Appalachian dinosaur taxa establishes a degree of endemism in Appalachian dinosaur assemblages attributable to episodes of vicariance.

scholarly journals An elongate hadrosaurid forelimb with biological traces informs the biogeography of the Lambeosaurinae

2020 ◽  
pp. 1-9
Author(s):  
Chase Doran Brownstein ◽  
Immanuel Bissell
Keyword(s):  
New Jersey ◽  
North America ◽  
Late Cretaceous ◽  
North American ◽  
American West ◽  
Atlantic Coastal Plain ◽  
Western Interior Seaway ◽  
Terrestrial Vertebrates ◽  
Both Bones ◽  
Atlantic Coastal

Abstract Although the fossil record of the Late Cretaceous eastern North American landmass Appalachia is poor compared to that from the American West, it includes material from surprisingly aberrant terrestrial vertebrates that may represent relictual forms persisting in relative isolation until the end of the Mesozoic. One intriguing question is to what extent eastern and western North American faunas interspersed following the closure of the Western Interior Seaway during the Maastrichtian Stage of the Late Cretaceous ca. 70 Ma. Isolated remains from the Atlantic Coastal Plain in New Jersey have been preliminarily identified as the bones of crested lambeosaurine hadrosaurids, a derived clade known from the Cretaceous of Asia, western North America, and Europe, but have not been formally described. We describe the partial forelimb of a large hadrosaurid from the late Maastrichtian New Egypt Formation of New Jersey. The ulna preserves multiple deep scores identifiable as shark feeding marks, and both bones show ovoid and circular marks attributable to invertebrates. This forelimb is very similar to another partial antebrachium from the same area that shows evidence of septic arthritis. Both these specimens and a complete humerus from the same unit are closely comparable to the lower forelimbs of lambeosaurines among hadrosaurid dinosaurs. Although the absence of lambeosaurine synapomorphies observable on the New Egypt Formation forelimbs precludes their definite referral to Lambeosaurinae, they show that a morphotype of large hadrosauromorph with distinctly elongate forelimbs existed in the latest Maastrichtian of eastern North America and allow for a revision of the latest Cretaceous biogeography of crested herbivorous dinosaurs.

scholarly journals A remarkable short-snouted horned dinosaur from the Late Cretaceous (late Campanian) of southern Laramidia

2013 ◽  
Vol 280 (1766) ◽  
pp. 20131186 ◽  
Author(s):  
Scott D. Sampson ◽  
Eric K. Lund ◽  
Mark A. Loewen ◽  
Andrew A. Farke ◽  
Katherine E. Clayton
Keyword(s):  
Phylogenetic Analysis ◽  
North America ◽  
Late Cretaceous ◽  
Fossil Record ◽  
Strong Support ◽  
Northern Region ◽  
New Taxon ◽  
Western North America ◽  
Postcranial Skeleton ◽  
Western Interior Seaway

The fossil record of centrosaurine ceratopsids is largely restricted to the northern region of western North America (Alberta, Montana and Alaska). Exceptions consist of single taxa from Utah ( Diabloceratops ) and China ( Sinoceratops ), plus otherwise fragmentary remains from the southern Western Interior of North America. Here, we describe a remarkable new taxon, Nasutoceratops titusi n. gen. et sp., from the late Campanian Kaiparowits Formation of Utah, represented by multiple specimens, including a nearly complete skull and partial postcranial skeleton. Autapomorphies include an enlarged narial region, pneumatic nasal ornamentation, abbreviated snout and elongate, rostrolaterally directed supraorbital horncores. The subrectangular parietosquamosal frill is relatively unadorned and broadest in the mid-region. A phylogenetic analysis indicates that Nasutoceratops is the sister taxon to Avaceratops , and that a previously unknown subclade of centrosaurines branched off early in the group's history and persisted for several million years during the late Campanian. As the first well-represented southern centrosaurine comparable in age to the bulk of northern forms, Nasutoceratops provides strong support for the provincialism hypothesis, which posits that Laramidia—the western landmass formed by inundation of the central region of North America by the Western Interior Seaway—hosted at least two coeval dinosaur communities for over a million years of late Campanian time.

Escalation in Late Cretaceous-early Paleocene oysters (Gryphaeidae) from the Atlantic Coastal Plain

Paleobiology ◽  
2000 ◽  
Vol 26 (2) ◽  
pp. 215-237 ◽  
Author(s):  
Gregory P. Dietl ◽  
Richard R. Alexander ◽  
Walter F. Bien
Keyword(s):  
Late Cretaceous ◽  
Coastal Plain ◽  
Selective Advantage ◽  
Atlantic Coastal Plain ◽  
Independent Evidence ◽  
Evolutionary Response ◽  
Frequency Independent ◽  
Early Paleocene ◽  
Surface Ornamentation ◽  
Atlantic Coastal

More than 1600 valves of Late Cretaceous and early Paleocene Northern Atlantic Coastal Plain gryphaeid oysters (Exogyrinae and Pycnodonteinae) were examined for breakage-induced shell repair and morphologic variability to evaluate the hypothesis of escalation. The Exogyrinae show disproportionately higher average repair frequency (0.41) relative to the ecologically and functionally similar unornamented pycnodonts (0.19). An increase in repair frequency (independent evidence of the action of a selective agent, e.g., predation) through the stratigraphic interval supports escalation. Variation in repair frequencies may reflect differences in oyster morphology and in the strength and diversity of shell crushers across an onshore-offshore gradient. Escalation of antipredatory adaptation characterized the evolutionary response of gryphaeid oysters to their durophagous predators. Adaptation generally occurred by the enhancement of existing traits in both oyster lineages. Characters that confer a selective advantage against predators are not all expressed or improved concurrently in both oyster lineages. Morphologic adaptations to minimize shell breakage include the development of expansive, broad commissural shelves, thickened valves, and surface ornamentation (Exogyrinae). Surface ornament in the Exogyrinae gradually increased with time. For some characters, such as thickness, conflicting functional demands (e.g., valve stabilization) may have limited adaptation to predators.

New Eclipidrilus species (Annelida, Clitellata, Lumbriculidae) from southeastern North America

Zootaxa ◽  
2012 ◽  
Vol 3194 (1) ◽  
pp. 51 ◽  
Author(s):  
STEVEN FEND ◽  
DAVID R. LENAT
Keyword(s):  
North Carolina ◽  
New Species ◽  
North America ◽  
Coastal Plain ◽  
Atlantic Coastal Plain ◽  
Middle Atlantic ◽  
Three New Species ◽  
Atlantic Coastal ◽  
Plain Streams ◽  
Coastal Plain Streams

Three new species of Lumbriculidae from southeastern North America are attributed to Eclipidrilus Eisen. All are small worms (diameter 0.2–0.5 mm), having semi-prosoporous male ducts with the atria in X, and spermathecae in IX. Eclipidrilus breviatriatus n. sp. and E. microthecus n. sp. have crosshatched atrial musculature, similar to some E. (Eclipidrilus) species, but they differ from congeners in having small, compact spermathecal ampullae. Eclipidrilus macphersonae n. sp. has a single, median atrium and spermatheca. The new species have been collected only in Sandhills and Middle Atlantic Coastal Plain streams of North Carolina.

scholarly journals A tyrannosauroid metatarsus from the Merchantville formation of Delaware increases the diversity of non-tyrannosaurid tyrannosauroids on Appalachia

PeerJ ◽  
2017 ◽  
Vol 5 ◽  
pp. e4123 ◽  
Author(s):  
Chase D. Brownstein
Keyword(s):  
Phylogenetic Analysis ◽  
North America ◽  
Late Cretaceous ◽  
Fossil Record ◽  
Morphological Features ◽  
The West ◽  
Intermediate Grade

During the Late Cretaceous, the continent of North America was divided into two sections: Laramidia in the west and Appalachia in the east. Although the sediments of Appalachia recorded only a sparse fossil record of dinosaurs, the dinosaur faunas of this landmass were different in composition from those of Laramidia. Represented by at least two taxa (Appalachiosaurus montgomeriensis and Dryptosaurus aquilunguis), partial and fragmentary skeletons, and isolated bones, the non-tyrannosaurid tyrannosauroids of the landmass have attracted some attention. Unfortunately, these eastern tyrants are poorly known compared to their western contemporaries. Here, one specimen, the partial metatarsus of a tyrannosauroid from the Campanian Merchantville Formation of Delaware, is described in detail. The specimen can be distinguished from A. montgomeriensis and D. aquilunguis by several morphological features. As such, the specimen represents a potentially previously unrecognized taxon of tyrannosauroid from Appalachia, increasing the diversity of the clade on the landmass. Phylogenetic analysis and the morphology of the bones suggest the Merchantville specimen is a tyrannosauroid of “intermediate” grade, thus supporting the notion that Appalachia was a refugium for relict dinosaur clades.

scholarly journals Biogeographic, stratigraphic, and environmental distribution of Basilosaurus (Mammalia, Cetacea) in North America with a review of the late Eocene shoreline in the southeastern coastal plain

2021 ◽  
pp. 1-13
Author(s):  
Kathlyn M. Smith ◽  
Alexander K. Hastings ◽  
Ryan M. Bebej ◽  
Mark D. Uhen
Keyword(s):  
North America ◽  
Coastal Plain ◽  
Late Eocene ◽  
Atlantic Coastal Plain ◽  
Thoracic Vertebrae ◽  
Environmental Distribution ◽  
Mississippi Embayment ◽  
Southeastern Coastal Plain ◽  
Flint River ◽  
Atlantic Coastal

Abstract A new specimen of Basilosaurus cetoides was discovered on the banks of the Flint River in Albany, Georgia, USA, in 2010. This fossil, which was the most complete specimen of the species from Georgia to date, consisted of five nearly complete and two partial post-thoracic vertebrae, tentatively identified as S4 through Ca6. During excavation, however, the site was looted and most of the specimen was lost to science. Nonetheless, we use this discovery as an opportunity to update the current state of knowledge on the stratigraphic, biogeographic, and environmental distribution of Basilosaurus in North America, as well as the position of the late Eocene shoreline in the southeastern United States. The results show that Basilosaurus was most abundant across the southeastern coastal plain during the early to middle Priabonian, coincident with the late Eocene maximum marine transgression. The decline in Basilosaurus localities is associated with the retreating shoreline of the terminal Eocene. The majority of Basilosaurus localities fall well south of the position of the late Eocene shoreline hypothesized in this study, suggesting the genus favored middle to outer neritic zones of the epicontinental sea. The comparatively low number of Priabonian specimens in the Atlantic Coastal Plain versus the Gulf Coastal Plain, then, suggests the presence of shallow zones in the Atlantic Coastal Plain that may have limited the distribution of Basilosaurus across the region. The hypothesized shoreline of this study ultimately differs from earlier reconstructions by extending the Mississippi embayment at the Bartonian/Priabonian boundary farther north than previously noted.

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