scholarly journals Species-groups and cladistic analysis of the cleptoparastic [sic] bee genus Nomada (Hymenoptera: Apoidea)

1994 ◽  
Vol 55 ◽  
pp. 175-236 ◽  
1990 ◽  
Vol 64 (4) ◽  
pp. 600-614 ◽  
Author(s):  
Jonathan M. Adrain ◽  
Brian D. E. Chatterton

Odontopleura (Odontopleura) arctica, a new species of odontopleurine trilobite, is described from the Canadian Arctic. A method of cladistic analysis is detailed. Parsimony analysis should be performed treating all characters as unordered. The universe of directed trees implied by the resulting rootless network(s) can then be examined and a preferred tree selected by a criterion of congruency. Namely, the most parsimonious directed tree that accommodates the most congruent arrangement of character-states should be taken as the preferred cladogram. Since this is essentially a general congruency method operating within the constraints of parsimony, it is termed “constrained congruency.” The method is applied to the genus Odontopleura, resulting in the recognition of two major species groups, the nominate subgenus and Sinespinaspis n. subgen. Odontopleura (Ivanopleura) dufrenoyi Barrande is tentatively included in the genus, but considered too poorly known for cladistic analysis. Species assigned to Odontopleura (Odontopleura) include Odontopleura ovata Emmrich, Odontopleura brevigena Chatterton and Perry, Odontopleura (Odontopleura) arctica n. sp., and Diacanthaspis serotina Apollonov. Species assigned to Sinespinaspis n. subgen. include Taemasaspis llandoveryana Šnajdr, Odontopleura greenwoodi Chatterton and Perry, Odontopleura maccallai Chatterton and Perry, and Odontopleura nehedensis Chatterton and Perry. Odontopleura bombini Chatterton and Perry is tentatively placed in synonymy with Odontopleura nehedensis. The genus had a wide distribution throughout the Early and Middle Silurian, due to preferences for deep-water, distal shelf or shelf-slope transition zone habitats.


2018 ◽  
Vol 49 (2) ◽  
pp. 103-129 ◽  
Author(s):  
Rogério Botion Lopes ◽  
Fernando Barbosa Noll

Zethus is the largest genus in Eumeninae, with over 250 species. Currently, it is divided in four subgenera: Z. (Zethus), Z. (Zethusculus), Z. (Zethoides) and Z. (Madecazethus). Z. (Zethoides), with 42 species, is subdivided in eight species groups, each considered a phylogenetic unit, that were created without any phylogenetic analysis. Eighteen species of Z. (Zethoides) corresponding to different groups were examined, altogether with terminals from distinct lineages of Zethus, Zethini and Eumenini, to perform a cladistics analysis to verify the proposed divisions. Zethus (Zethoides) and all of its species groups, except for the Z. biglumis group, were monophyletic. Zethus s.s. was paraphyletic in relation to Z. (Madecazethus), Z. (Zethoides) and Ctenochilus. Z. (Zethusculus) was also retrieved paraphyletic. Despite the subgeneric incongruences, the outgroups were too poorly represented to carry a taxonomic modification. Thus, the only alteration was the inclusion of the Z. clypearis group in the Z. biglumis group.


2001 ◽  
Vol 15 (3) ◽  
pp. 353 ◽  
Author(s):  
Brian E. Heterick

The Australian ants of the genus Monomorium are revised. Fifty-nine species are recognised. Of these, 41 are described as new: Monomorium aithoderum, M. albipes, M. anderseni, M. anthracinum, M. arenarium, M. bifidum, M. bihamatum, M. brachythrix, M. burchera, M. capito, M. carinatum, M. castaneum, M. crinitum, M. decuria, M. disetigerum, M. draculai, M. durokoppinense, M. elegantulum, M. eremophilum, M. euryodon, M. flavonigrum, M. lacunosum, M. legulus, M. longinode, M. macarthuri, M. majeri, M. megalops, M. micula, M. nanum, M. nightcapense, M. nigriceps, M. parantarcticum, M. petiolatum, M. pubescens, M. ravenshoense, M. rufonigrum, M. shattucki, M. silaceum, M. stictonotum, M. striatifrons, and M. xantheklemma. Thirteen species pass into synonymy: M. armstrongi with M. whitei, M. broomense and M. ilia with M. laeve, M. donisthorpeiand M. fraterculus with M. fieldi, M. flavipes and M. insularis with M. leae, M. foreli with M. sordidum, M. howense with M. tambourinense, M. macareaveyi with M. bicorne, M. sanguinolentum with M. rubriceps, M. subapterum with M. rothsteini, and M. turneri withM. gilberti. Sixteen infraspecific forms are also synonymised: M. kilianii obscurelluminto M. kilianii, M. laeve nigriusand M. laeve fraterculus into M. fieldi, M. ilia lamingtonensisinto M. laeve, M. rothsteini humilior, M. rothsteini leda, M. rothsteini doddi and M. subapterum bogischi into M. rothsteini, M. rothsteini squamigena, M. rothsteini tostum and M. sordidum nigriventris into M. sordidum, M. fraterculus barretti and M. sydneyense nigella into M. sydneyense, M. gilberti mediorubra into M. gilberti, and M. rubriceps cinctumand M. rubriceps rubrum into M. rubriceps. Seventeen species and one subspecies are unchanged. Monomorium kiliani reverts to M. kilianii, M. kilianii tambourinenseis raised to species status, M. occidaneus is here treated as a species inquirenda, and M. flavigaster is removed from the genus Monomorium. Since the generic status of the latter taxon is uncertain, M. flavigaster is here regarded as incertae sedis. The supposedly extralimitalMonomorium talpa is synonymised under Monomorium australicum. At a higher taxonomic level the South American genus Antichthonidris is synonymised under Monomorium. Seven species-groups are proposed for the Australian fauna, (the bicorne-, falcatum-, insolescens-, kilianii-, longinode-, monomorium-, and rubriceps-groups). A cladistic analysis was undertaken of species for which all castes were examined (identifiable males and/or queens were lacking for all members of the falcatum-, insolescens- and longinode-groups). In all, fifteen species of Australian Monomorium were examined (M. bicorne, M. whitei, M. striatifrons and M. rufonigrum from the bicorne-group, M. crinitumand M. kilianii from the kilianii-group, M. fieldi, M. laeve, M. rothsteini, M. sordidum and M. sydneyense from the monomorium-group, and M. centrale, M. leae, M. euryodon and M. rubriceps from the rubriceps-group), together with Monomorium antarcticum(from New Zealand) and the Neotropical Antichthonidris denticulatus. The taxon used for the outgroup was the Neotropical ant Megalomyrmex modestus. Using the PAUP program, 37 characters for worker, queen and male castes were analysed. The clade incorporating the tiny generalists (M. fieldi, M. laeve, M. sordidum, and M. sydneyense), together with M. rothsteini, was found to be the clade most strongly supported as a monophyletic grouping. In this analysis M. euryodon was the sister taxon to the above clade. These ants were shown on this analysis to share a common ancestor with the other members of the rubriceps-group, with M. antarcticum and A. denticulatus, and with thekilianii-group. The relationships between these latter four sets of species were left unresolved, except that M. crinitum was shown to be the sister taxon to M. kilianii. The large, arid zone species in thebicorne-group were also shown as ancestral to the other Australian Monomorium. A key is provided to enable researchers to identify the workers of all Australian Monomorium, as well as extralimital species established in Australia.


1993 ◽  
Vol 125 (S168) ◽  
pp. 1-193 ◽  
Author(s):  
Valerie M. Behan-Pelletier

AbstractThe oribatid family Eremaeidae is represented in North America by two genera, Eremaeus and Eueremaeus, both widely distributed throughout the Palaearctic and Nearctic regions. In North America species in both genera are found in moist to arid habitats from New Mexico to the High Arctic. Reproduction is sexual, and both immatures and adults feed mainly on fungi.Revised diagnoses are presented for the Eremaeidae and genera Eremaeus and Eueremaeus. Eighteen species of Eremaeus, of which 14 are newly proposed, and 24 species of Eueremaeus, of which 15 are newly proposed, are recognized. Identification keys are provided for the world genera of Eremaeidae, and for adults of Eremaeus and Eueremaeus of North America. All but one North American species of these genera are described, and their geographical distributions mapped.North American Eremaeus species include E. appalachicus sp. no v., E. boreomontanus sp. nov., E. brevitarsus (Ewing), E. californiensis sp. nov., E. gracilis sp. nov., E. grandis Hammer, E. kananaskis sp. nov., E. kevani sp. nov., E. megistos sp. nov., E. monticolus sp. nov., E. nortoni sp. nov., E. occidentalis sp. nov., E. oresbios sp. nov., E. plumosus Woolley, E. porosus sp. nov., E. salish sp. nov., E. translamellatus Hammer, and E. walteri sp. nov. The immatures of four of these, E. kananaskis, E. occidentalis, E. oresbios, and E. translamellatus, are described.North American Eueremaeus include Eu. acostulatus sp. nov., Eu. aridulus sp. nov., Eu. columbianus (Berlese), Eu. foveolatus (Hammer), Eu. marshalli sp. nov., Eu. masinasin sp. nov., Eu. michaeli sp. nov., Eu. nahani sp. nov., Eu. nemoralis sp. nov., Eu. proximus (Berlese) comb, nov., Eu. woolleyi (Higgins) comb, nov., Eu. yukonensis sp. nov., and three informal species groups with the following included species in North America: (1) Eu. trionus group—Eu. trionus (Higgins) comb, nov., (2) Eu. stiktos group—Eu. carinatus sp. nov., Eu. higginsi sp. nov., Eu. stiktos (Higgins) comb, nov., Eu. tetrosus (Higgins) comb, nov., (3) Eu. chiatous group—Eu. alvordensis sp. nov., Eu. aysineep sp. nov., Eu. chiatous (Higgins) comb, nov., Eu. danos sp. nov., Eu. lindquisti sp. nov., Eu. magniporosus (Wallwork) comb, nov., and Eu. osoyoosensis sp. nov. The immatures of nine of these, Eu. masinasin, Eu. nahani, Eu. carinatus, Eu. higginsi, Eu. columbianus, Eu. proximus, Eu. woolleyi, Eu. stiktos, and Eu. tetrosus, are described. Kartoeremaeus reevesi Higgins and Eremaeus politus Banks are considered junior subjective synonyms of Eueremaeus columbianus (Berlese).A cladistic analysis of the genera comprising Eremaeidae: Eremaeus, Tricheremaeus, Eueremaeus (and included species groups), Proteremaeus, Carinabella, and Asperemaeus, indicates that Eremaeus is the sister taxon of Carinabella, and that Eueremaeus is the sister taxon of Proteremaeus. Tricheremaeus is the sister taxon of Eremaeus + Carinabella, and Asperemaeus is the sister taxon of Eueremaeus + Proteremaeus. The relationship of the Eremaeidae to the Megeremaeidae and Zetorchestidae is presented. Finally, I discuss the ecology and distribution of North American species of Eremaeidae.


Zootaxa ◽  
2010 ◽  
Vol 2425 (1) ◽  
pp. 45 ◽  
Author(s):  
ÂNGELO PARISE PINTO ◽  
ALCIMAR DO LAGO CARVALHO

Lauromacromia melanica sp. nov. from Conceição da Barra municipality, Espírito Santo State, Brazil, is described and illustrated based on two males (both in MNRJ nº 135). The new species is similar to L. picinguaba differing from it mainly by the absence of pale spots on S3–6 and by the ellipsoid shape of metepisternal pale stripe. A key for males of all species of the genus is provided. A cladistic analysis encompassing 43 external morphological male characters carried out in two distinct procedures, the first with all characters unordered and the second with two or three state characters ordered. The unordered analysis generated only one most-parsimonious tree (66 steps of length, CI = 0.69, RI = 0.62). The hypothesis of monophyly of Lauromacromia is supported and includes three groups, one formed by the Atlantic Forest species (L. melanica sp. nov. + L. picinguaba), and another by the Cerrado species (L. flaviae + (L. bedei + L. luismoojeni)), and L. dubitalis, positioned in polytomy with these two groups. The ordered analysis also generated only one most-parsimonious tree (68 steps of length, CI = 0.70, RI = 0.67), which maintained the monophyly of Lauromacromia but L. dubitalis positioned basally as sister-group to the Atlantic Forest + Cerrado species groups. The geographic distribution of Lauromacromia is updated with a new record of L. luismoojeni based on one adult male (Brazil: Mato Grosso do Sul State) and probable first Brazilian records for L. dubitalis (Amazonas and Pará States) based on two larvae. A vicariance hypothesis is proposed to explain spatial evolution of Lauromacromia, and based on current biogeographical classifications we consider Gomphomacromia and Rialla apart from Neotropical biota. Some aspects of biology and ecology of Lauromacromia are also discussed.


Zootaxa ◽  
2012 ◽  
Vol 3577 (1) ◽  
pp. 1 ◽  
Author(s):  
HIROYUKI TAKAOKA

Simulium (Gomphostilbia) Enderlein, the third largest (206 named species included) in the genus Simulium Latreille s. l., is one of the two most abundant and diverse subgenera in the Oriental Region. To provide a classification scheme to facilitate morphological identification of the species within this subgenus, its diagnostic characters are redefined, and nine known species-groups within it are reviewed. Based on putative lineages explored by using certain adult morphological characters, seven more species-groups are proposed: asakoae, darjeelingense, epistum, gombakense, heldsbachense, hemicyclium and palauense species-groups, while the trirugosum species-group is merged in the varicorne species-group. Subgroups are also introduced to represent apparently different lineages within certain species-groups based on certain pupal morphological characters: two in the banauense species-group, seven in the batoense species-group redefined, four in the ceylonicum species-group redefined, four in the epistum species-group, two in the hemicyclium species-group, two in the sherwoodi species-group and four in the varicorne species-group redefined. A new checklist of species of the subgenus Gomphostilbia, and a key to all 15 species-groups within it are provided. The eastward expansion of the geographical distribution of the subgenus Gomphostilbia is inferred on the basis of the more frequent occurrence of apomorphic characters of certain adult and pupal morphological features in insular species-groups than in continental species-groups. A preliminary attempt using a cladistic analysis of morphological characters shows that among 10 subgenera examined, Gomphostilbia has a sister-taxon relationship with the Australasian subgenus Morops Enderlein, and this clade, together with the Central-Western Pacific subgenus Inseliellum Rubtsov, is positioned closest to the most derived clade formed by Daviesellum Takaoka & Adler and Simulium Latreille s. str.


Zootaxa ◽  
2005 ◽  
Vol 1027 (1) ◽  
pp. 55 ◽  
Author(s):  
ROXANA ACOSTA ◽  
JUAN J. MORRONE

A new flea species, Hystrichopsylla cryptotis, is described from the Sierra Madre Oriental and Sierra Madre del Sur, Mexico. The host of this new species is the shrew Cryptotis mexicana (Coues, 1877) (Mammalia: Soricidae). This flea species is easily recognized by its large size, seven genal combs, and the slender sternum IX, with 13 pairs of thick spiniform setae of different sizes. A key to the Mexican and Guatemalan species of Hystrichopsylla is given. The cladistic analysis indicates that Mexican species of Hystrichopsylla may be arranged in two different species groups: the H. orophila species group (H. orophila Barrera 1952 and H. cryptotis) and the H. dippiei species group (H. dippiei Rotshchild 1902, H. llorentei Ayala and Morales 1990, and H. kris Traub and Johnson 1952). The resolved area cladogram based on their phylogenetic relationships indicates the following relationships: (Sierra Madre Oriental, (Sierra Madre del Sur, Transmexican Volcanic Belt)).


Zootaxa ◽  
2012 ◽  
Vol 3369 (1) ◽  
pp. 1 ◽  
Author(s):  
PAKAWIN DANKITTIPAKUL ◽  
RUDY JOCQUÉ ◽  
TIPPAWAN SINGTRIPOP

The systematics status of the spider genus Mallinella Strand, 1906 (Araneae, Zodariidae), the phylogenetic relationshipof the species within the genus and its relationships to other zodariids were investigated by means of cladistic analysis ofmorphological data. Mallinella is redefined and characterized by a single synapomorphy: the presence of posterior ventralspines situated in front of the spinnerets arranged in a single row. The genus is clearly palaeotropical, occurring in Africa,Indian subcontinent, Indo-Burma, Sundaland, Wallacea and Polynesia-Micronesia.Two hundred and two (202) Mallinella species are treated. One hundred and one (101) species are described as newand placed in twenty-two (22) species-groups, making Mallinella the largest zodariid genus. Nineteen (19) species are redescribed, the conspecific sex of seven (7) species is discovered and described for the first time. Fifteen (15) new com-binations are proposed. Nine (9) Storena species are here transferred to Mallinella: M. beauforti (Kulczyński, 1911) comb.nov., M. sciophana (Simon, 1901) comb. nov., M. sobria (Thorell, 1890) comb. nov., M. fasciata (Kulczyński, 1911)comb. nov., M. vicaria (Kulczyński, 1911) comb. nov., M. redimita (Simon, 1905) comb. nov., M. melanognatha (van Has-selt, 1882) comb. nov., M. nilgherina (Simon, 1906) comb. nov., M. vittata (Thorell, 1890) comb. nov. Two Storena spe-cies are transferred to Asceua: A. dispar (Kulczyński, 1911) comb. nov., A. quinquestrigata (Simon, 1905) comb. nov. OneStorena species is transferred to Oedignatha (Liocranidae): O. aleipata (Marples, 1955) comb. nov. One Storena speciesis transferred to Cybaeodamus: C. lentiginosus (Simon, 1905) comb. nov. Storena tricolor Simon, 1908 is transferred tothe Asteron complex of Australia. Three Storena and two Mallinella species are misplaced; they belong to undescribedgenera (S. kraepelini Simon, 1905; S. lesserti Berland, 1938; S. parvula Berland, 1938; M. khanhoa Logunov, 2010; M.rectangulata Zhang et al., 2011). Mallinella vittata (Thorell, 1890) comb. nov. is revalidated and removed from the syn-onymy with M. zebra (Thorell, 1881). Storena vittata Caporiacco, 1955 is removed from homonym replacement (S. ca-poriaccoi Brignoli, 1983) with S. vittata Thorell, 1890 (= M. vittata comb. nov.). Storena annulipes Thorell, 1892 isremoved from its preoccupied name with S. annulipes (L. Koch, 1867) in Storena and transferred to Mallinella; its re-placement name S. cinctipes Simon, 1893 is suppressed.Zodarion luzonicum Simon, 1893, Storena multiguttata Simon, 1893, S. semiflava Simon, 1893 and S. obnubila Si-mon, 1901 are regarded as nomina dubia. Six Indian species were misplaced in Storena; they belong to one of the follow-ing genera: Mallinella, Heliconilla gen. nov., Workmania gen. nov., Heradion, or Euryeidon. These taxa are S. arakuensisPatel & Reddy, 1989, S. debasrae Biswas & Biswas, 1992, S. dibangensis Biswas & Biswas, 2006, S. gujaratensis Tikader& Patel, 1975, S. indica Tikader & Patel, 1975 and S. tikaderi Patel & Reddy, 1989. They are regarded as species incertaesedis.A new genus, Heliconilla gen. nov., is proposed for nine species, six of which are new to science while the otherthree are transferred from Mallinella and Storena. These taxa are: H. irrorata (Thorell, 1887) comb. nov., H. oblonga(Zhang & Zhu, 2009) comb. nov., H. thaleri (Dankittipakul & Schwendinger, 2009) comb. nov.Workmania gen. nov. is established to accommodate two species from Southeast Asia; W. juvenca (Workman, 1896)comb. nov. is transferred from Storena.It is unlikely that the origin of Mallinella dates back more than 100 MYA. Mallinella or its ancestor is believed tohave evolved during the Cretaceous, after the separation of South America from Gondwana, and the greater part of itsevolution took place during the Tertiary. The Asian-Australian lineages of Mallinella could migrate to India via GreaterSomalia before or after the K-T extinction (65 MYA), before the Indian subcontinent joined Asia (ca. 45 MYA).The bio-geographic history of the genus involves plate tectonics during the Cretaceous and the Cenozoic in combination with cli-matic changes and alternating climatic cycles which might have led to episodes of range expansion, isolation of populations and allopatric speciation.


1992 ◽  
Vol 23 (3) ◽  
pp. 233-248 ◽  
Author(s):  
Knut Rognes

AbstractWithin Pollenia Robineau-Desvoidy a venturii species-group is defined and revised. It consists of a single species P. venturii Zumpt. P. solitaria Grunin is proposed as a junior synonym. It is characterized by unique features in the male aedeagus and the lateral sacs of the internal female reproductive organs. Male and female terminalia are illustrated, the latter for the first time. A preliminary cladistic analysis of all known Palaearctic species of Pollenia (except P. japonica Kano & Shinonaga) suggests that the sister-group of P. venturii is a clade consisting of the viatica + vagabunda + amentaria + haeretica species-groups. A sclerotized internal wall of the lateral sacs in the internal reproductive system of female Pollenia appears to be a parallelism developed independently in the venturii, rudis, most members of the tenuiforceps and some members of the semicinerea groups, rather than an underlying synapomorphy. P. venturii is known from France, Germany, Greece, Italy and Russia. A key is provided to species-groups in Pollenia.


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