Systematics and biology of the Ichnestomina, including new genera and species (Coleoptera: Scarabaeidae, Cetoniinae)

2020 ◽  
Vol 52 (2) ◽  
pp. 217-320
Author(s):  
Renzo Perissinotto

The subtribe Ichnestomina, endemic to southern Africa, exhibits a suite of plesiomorphic features that reflects a particularly old evolutionary age among the Goliathini. Sexual dimorphism is extreme and females are generally brachypterous and unable to fly. Also, adult stages do not feed, and therefore remain active for only a few days after emerging from the soil, devoting all their energy reserves to reproduction. Consequently, populations are range-bound to small areas, often completely isolated on high mountain peaks or ridges, leading to rapid speciation. Results of recent investigations and historical data are hereby combined to provide an updated and revised structure of the systematics of this subtribe. As a result, four genera are now recognised in place of the perceived uniformity earlier expressed with much reservation by Holm (1992). This involves the rehabilitation of the original name of Gariep Gory & Percheron, 1833, to accommodate species with fully winged and flying females as well as elongate and incision-free dorsal lobes of parameres (Gariep patera and G. perstriata. It is further proposed that the genus Ichnestoma Gory & Percheron, 1833 be restricted to species with moderate female brachyptery and males with cretaceous markings on their body surface. Consequently, two new genera are erected (Karooida gen. nov., Mzansica gen. nov.) and four species-complexes (Ichnestoma cuspidata, I. albomaculata, Mzansica nasula and M. luridipennis), are added in order to account for key differences among species, including degree of female brachyptery (wing:elytron length ratio), presence/absence of cretaceous markings, aedeagal structure and biogeographic distribution. Eleven new species (Ichnestoma carbonaria sp. nov., I. dealbata sp. nov., I. furcata sp. nov., I. karoominoris sp. nov., I. spatulata sp. nov., Karooida balli sp. nov., K. sagittata sp. nov., Mzansica botswanica sp. nov., M. clarki sp. nov., M. denticulata sp. nov., M. falcipata sp. nov.) are added to the 13 species and three subspecies currently recognised in this subtribe. The three subspecies are elevated to species status (Ichnestoma cochleata stat. nov., I. fuscipennis stat. nov., Karooida kikvorsti comb. et stat. nov.). The diagnostic characters of each taxon are described along with all available knowledge on the habitat characteristics and general biology of each species.

Zootaxa ◽  
2020 ◽  
Vol 4722 (2) ◽  
pp. 157-174
Author(s):  
ALLEN F. SANBORN

A recent genetic analysis has shown that the genus Pycna Amyot & Audinet-Serville, 1843 is polyphyletic. New genera are proposed for the species that have been classified together in the genus previously but form distinct clades from the clade containing the true species of Pycna. Pycnoides n. gen. is described for the species inhabiting continental Africa. Pycna antinorii (Lethierry, 1881), Pycna baxteri Distant, 1914, Pycna becarrii (Lethierry, 1881), Pycna dolosa Boulard, 1975a, Pycna hecuba Distant, 1904b, Pycna moniquae Boulard, 2012, Pycna natalensis Distant, 1905b, Pycna neavei Distant, 1912b, Pycna passosdecarvalhoi Boulard, 1975b, Pycna quanza (Distant, 1899a), Pycna semiclara (Germar, 1834), Pycna sylvia (Distant, 1899b), Pycna umbelinae Boulard, 1975b, Pycna vitrea Schumacher, 1913, and Pycna vitticollis (Jacobi, 1904) are reassigned to Pycnoides n. gen. to become Pycnoides antinorii (Lethierry, 1881) n. comb., Pycnoides baxteri (Distant, 1914) n. comb., Pycnoides becarrii (Lethierry, 1881) n. comb., Pycnoides dolosa (Boulard, 1975a) n. comb., Pycnoides hecuba (Distant, 1904b) n. comb., Pycnoides moniquae (Boulard, 2012) n. comb., Pycnoides natalensis (Distant, 1905b) n. comb., Pycnoides neavei (Distant, 1912b) n. comb., Pycnoides passosdecarvalhoi (Boulard, 1975b) n. comb., Pycnoides quanza (Distant, 1899a) n. comb., Pycnoides semiclara (Germar, 1834) n. comb., Pycnoides sylvia (Distant, 1899b) n. comb., Pycnoides umbelinae (Boulard, 1975b) n. comb., Pycnoides vitrea (Schumacher, 1913) n. comb., and Pycnoides vitticollis (Jacobi, 1904) n. comb. Pycnoides zambiaensis n. sp. is described as new to increase the diversity of the new genus further and represents the first record of a species in the group for Zambia. Eopycna n. gen. is described for the Asian species previously assigned to Pycna. Pycna coelestia Distant, 1904a, Pycna concinna Boulard, 2005, Pycna himalayana (Naruse, 1977), Pycna indochinensis Distant, 1913, Pycna minor Liu, 1940, Pycna montana Hayashi, 1978, Pycna repanda (Linnaeus, 1758), and Pycna verna Hayashi, 1982 are reassigned to Eopycna n. gen. to become Eopycna coelestia (Distant, 1904a) n. comb., Eopycna concinna (Boulard, 2005) n. comb., Eopycna himalayana (Naruse, 1977) n. comb., Eopycna indochinensis (Distant, 1913) n. comb., Eopycna minor (Liu, 1940) n. comb., Eopycna montana (Hayashi, 1978) n. comb., Eopycna repanda (Linnaeus, 1758) n. comb., and Eopycna verna (Hayashi, 1982) n. comb. Pycna angusta (Butler, 1882) rev. stat. is removed from junior synonymy of Pycna madagascarensis (Distant, 1881). Pycna schmitzi Boulard, 1979 is reassigned to Canualna Boulard, 1985a to become Canualna schmitzi (Boulard, 1979) n. comb. and Pynca tangana (Strand, 1910) is reassigned to Orapa Distant, 1905c to become Orapa tangana (Strand, 1910) n. comb. 


Zootaxa ◽  
2009 ◽  
Vol 2102 (1) ◽  
pp. 1-74 ◽  
Author(s):  
STEWART B. PECK ◽  
JOYCE COOK

This paper is a review and revision of the tribe Sogdini (Coleoptera: Leoididae: Leiodinae) of North and Central America. It covers the following genera: Triarthron Märkel, 1840, with one species; Hydnobius Schmidt, 1841, with seven species; Stereus Wollaston, 1857, new for North America, with one species; Sogda Lopatin, 1961, new for North America, with two species; Kalohydnobius new genus, with three species; Macrohydnobius new genus, with six species; and Platyhydnobius new genus, with eight species. The species are inhabitants of northern and mountain forests or sandy habitats. Larvae and adults probably feed on subterranean fungi. The following new synonyms are recognized: Triarthron pennsylvanicum Horn, 1883 = T. lecontei Horn, 1868; Hydnobius luggeri Hatch, 1927 = H. substriatus LeConte, 1863; Hydnobius lobatus Hatch, 1936 = H. longidens LeConte, 1879; Hydnobius longulus LeConte, 1879 = Sogda obtusa (LeConte,1879); Hydnobius femoratus Hatch, 1936 = Kalohydnobius strigilatus (Horn, 1880); and Hydnobius stacesmithi Hatch, 1957 = Macrohydnobius matthewsii (Crotch, 1874). Hydnobius kiseri Hatch, 1936 and H. longidens LeConte, 1879 are resurrected from synonymy to valid species status. The following new combinations are established: Sogda obtusa (LeConte, 1879), ex Hydnobius; Kalohydnobius strigilatus (Horn, 1880), ex Hydnobius; Macrohydnobius contortus (Hatch, 1957), ex Hydnobius; M. crestonensis (Hatch, 1957), ex Hydnobius; M. matthewsii (Crotch, 1874), ex Hydnobius; M. simulator (Brown, 1932), ex Hydnobius; Platyhydnobius arizonensis (Horn, 1885), ex Hydnobius; and P. validus (Brown1932), ex Hydnobius. Lectotypes are designated for Hydnobius substriatus LeConte, 1863; H. pumilus LeConte, 1879; H. obtusus LeConte, 1879; H. longulus LeConte, 1879; H. strigilatus Horn, 1880 and H. matthewsii Crotch, 1874. New species are Hydnobius acarinus, H. autumnalis, Stereus arenarius, Sogda enigma, Kalohydnobius californicus, K. dentatus, Macrohydnobius montanus, M. tibiocalcaris, Platyhydnobius andersoni, P. bicolor, P. calvario, P. howdenorum, P. latigra, and P. newtonorum.


2004 ◽  
Vol 18 (4) ◽  
pp. 377 ◽  
Author(s):  
Jean Just ◽  
George D. F. Wilson

This paper presents a global review of the current unwieldy concept of the genus Paramunna Sars, 1866. The study is based mainly on large new collections of material from Australia and subantarctic islands south of Tasmania and New Zealand. Of the four genera previously synonymised with Paramunna, Austrimunna Richardson, 1906 is revived (type species A. antarctica), Leptaspidia Bate & Westwood, 1867 and Metamunna Tattersall, 1905 are considered indeterminable, while Austronanus Hodgson, 1910 is not considered part of the Paramunna complex. Forty-three species, 27 new, are treated, mostly from Australia and adjacent subantarctic islands. Fifteen species currently in Paramunna do not belong in this complex. The type genus Paramunna Sars, 1866 comprises four known species, P. bilobata Sars, 1866, P. capensis Vanh�ffen, 1914, P. integra Nordenstam, 1933 and P. koreana Malyutina & Ushakova, 2001, and four new species. Nine new genera are created based on a cladistic analysis (type species, original combination): Ascionana (A. darwinia, sp. nov.), Epipedonana (E. profunda, sp. nov.), Harrietonana (Austrimunna subtriangulata Richardson, 1908), Kiklonana (Paramunna arnaudi Amar & Roman, 1974), Omonana (O. brachycephala, sp. nov.), Pagonana (Paramunna rostrata Hodgson, 1910), Palanana (Austrimunna serrata Richardson, 1908), Spiculonana (S. platysoma, sp. nov) and Sporonana (S. robusta, sp. nov.). Six species of Paramunna are transferred to other genera in the complex: P. simplex Menzies, 1962 and P.�parasimplex Winkler, 1994 to Omonana, gen. nov.; P. dilatata Vanh�ffen, 1914 to Pagonana, gen. nov.; P. gaini (Richardson, 1913) to Palanana gen. nov.; P. laevifrons Stebbing, 1910 and P. rhipis Shimomura & Mawatari, 1999 to Ascionana, gen. nov. Paramunna shornikovi Malyutina & Ushakova, 2001, is synonymised with P. rhipis. Keys to genera and species (if more than two in a genus) are given. Terminal males (males with elongated cephalon and massively enlarged pereonite 1) are documented in several genera. The distribution of the complex confirms that this part of the Paramunnidae is a Southern Hemisphere, shallow water group. Species previously thought to be circumpolar prove to be species complexes, with each species having a small distribution.


Zootaxa ◽  
2021 ◽  
Vol 4937 (1) ◽  
pp. 1-79
Author(s):  
ALLEN F. SANBORN

The known cicada fauna of Madagascar is identified. Yanga mahajangaensis n. sp., Nelbroma betrokaensis n. gen., n. sp., Trismarcha parva n. sp., Abroma bezamahafalyensis n. sp., Abroma falcata n. sp., Timbaltransversa pronotosigillata n. gen., n. sp., Malagasia argentea n. sp., Malagasia italivianaensis n. sp., Malagasia rubrastragala n. sp., Deremeces andringitraensis n. gen., n. sp., Malgachialna rubracosta n. sp., and Cephalalna tulearensis n. sp. are described as new. The genera Mouldspsaltria n. gen., and Orientafroinsularis n. gen. are described as new with the reassignment of Mouldspsaltria madegassa (Boulard, 1996) n. comb., Orientafroinsularis elenae n. comb., rev. stat., Orientafroinsularis hova (Distant, 1905b) n. comb., Orientafroinsularis loici (Boulard, 2000) n. comb., Orientafroinsularis martini (Distant, 1905b) n. comb., Orientafroinsularis matilei (Boulard, 2000) n. comb., Orientafroinsularis nigrans (Distant, 1904) n. comb., Orientafroinsularis pulverulenta (Distant, 1905b) n. comb., Orientafroinsularis seychellica (Dmitriev, 2020) n. comb., rev. stat., and Orientafroinsularis seychellensis (Boulard, 1999) n. comb., rev. stat. to these new genera. Yanga argyrea (Melichar, 1896) rev. stat. is removed from subspecies status of Yanga pulverea (Distant, 1882) and returned to species rank. Antankaria pulverulenta madegassa Boulard, 1999 n. syn. and Chremistica pulverulenta madagascariensis Boulard, 2001 n. syn. are considered synonyms of Orientafroinsularis pulverulenta (Distant, 1905b) n. comb. eliminating the potential homonym based on “madagascariensis”. Antankaria pulverulenta seychellensis Boulard, 1999 and Chremistica pulverulenta seychellensis Boulard, 2001 n. syn. are considered synonyms, elevated to species rank and reassigned to the new genus to become Orientafroinsularis seychellensis (Boulard, 1999) n. comb., rev. stat. Chremistica elenae elenae Boulard, 2001 rev. stat. is shown to be a synonym of Chremistica elenae madagascariensis Boulard, 2001 n. syn. and is a distinct species from Chremistica elenae seychellica Dmitriev, 2020 rev. stat. with both elevated to species rank and transferred to the new genus to become Orientafroinsularis elenae (Boulard, 2001) n. comb., rev. stat. and Orientafroinsularis seychellica (Dmitriev, 2020) n. comb., rev. stat. Antankariini n. tribe is formed for former Malagasy members of the Cryptotympanini Handlirsch, 1925. Four genera are reassigned to Malagasiini Moulds & Marshall, 2018, Anopercalna Boulard, 2008 from Taphurini Distant, 1905d, Cephalalna Boulard, 2006a from Chlorocystini Distant, 1905c, Malgotilia Boulard, 1980b from Parnisini Distant, 1905c and Viettealna Boulard, 1980a from Lamotialnini Boulard, 1976. There are no remaining members of Chlorocystini or Taphurini in Madagascar after the generic reassignments and these tribes are removed from the Madagascar fauna. Anopercalnina Boulard, 2008 n. syn. is made a junior synonym to Malagasiini Moulds & Marshall, 2018. An updated faunal list is provided to summarize recent changes to the assignment of taxa. The currently known cicada fauna of Madagascar is comprised of 65 species from 25 genera, eight tribes and three subfamilies. 


Zootaxa ◽  
2007 ◽  
Vol 1603 (1) ◽  
pp. 1-78 ◽  
Author(s):  
JUAN J. SCHMITTER-SOTO

The cichlid genus Archocentrus has been considered one of the most promising (i.e., possibly natural) genera resurrected to receive some of the species formerly included in Cichlasoma. Evidence is presented to justify generic recognition of Archocentrus, as well as eight other closely related genera (Caquetaia, Hypsophrys, Parachromis, Amphilophus, Archocentrus, Cryptoheros, Amatitlania, and Rocio). Of these, Amatitlania (type species, A. nigrofasciata) and Rocio (type species, R. octofasciata) are described as new. The present revision treats all nominal species ever assigned to Archocentrus, as well as species that have been included in or near the same clade as Archocentrus centrarchus (type species of the genus) in available phylogenetic analyses. Geographical variation in morphology of the more widespread species was examined, which has resulted in the description of six new species (Cryptoheros chetumalensis, Amatitlania coatepeque, A. kanna, A. siquia, Rocio gemmata, and R. ocotal) with a seventh resurrected from synonymy (Cryptoheros cutteri). Archocentrus includes the type species (Ar. centrarchus), plus Ar. spinosissimus and Ar. multispinosus. Cryptoheros is restricted to the species complexes of Cr. spilurus (= subgenus Cryptoheros, including also Cr. chetumalensis and Cr. cutteri) and Cr. septemfasciatus (= Bussingius n. subgen., including also Cryptoheros altoflavus, Cr. nanoluteus, Cr. myrnae, and Cr. sajica); Cryptoheros panamensis is placed in Panamius n. subgen. Herotilapia is synonymized with Archocentrus, and Neetroplus is synonymized with Hypsophrys, which now includes the type species H. nicaraguensis and H. nematopus. Lectotypes are designated for Amatitlania nigrofasciata, Archocentrus spinosissimus, Cryptoheros septemfasciatus, Cr. spilurus, and Rocio octofasciata. Cichlasoma immaculatum is considered to be a synonym of Archocentrus spilurus, not of Ar. spinosissimus.


Zootaxa ◽  
2009 ◽  
Vol 2255 (1) ◽  
pp. 1-100 ◽  
Author(s):  
DONALD G. BROADLEY ◽  
VAN WALLACH

All eastern and southern African species of Typhlopidae (excluding Letheobia) are reviewed with synonymies, diagnoses, and distribution maps. The 27 species from this region are arranged as follows: three species remain in Typhlops (with T. cuneirostris calabresii elevated to full species status), 16 species are transferred from Typhlops to a new genus endemic to Africa (including a new species described from Kenya), three species remain in Rhinotyphlops, four species are transferred from Rhinotyphlops to a new genus endemic to Africa (with R. schlegelii brevis and R. s. mucruso elevated to full species status), and one invasive species of Ramphotyphlops is included, which has been established in a few areas. Keys to the included species and all six African typhlopid genera are provided. A current taxonomic synopsis is provided for extralimital African and Southwest Asian typhlopid species.


2011 ◽  
Vol 20 (1) ◽  
pp. 161-173
Author(s):  
A.P. Kassatkina

Resuming published and own data, a revision of classification of Chaetognatha is presented. The family Sagittidae Claus & Grobben, 1905 is given a rank of subclass, Sagittiones, characterised, in particular, by the presence of two pairs of sac-like gelatinous structures or two pairs of fins. Besides the order Aphragmophora Tokioka, 1965, it contains the new order Biphragmosagittiformes ord. nov., which is a unique group of Chaetognatha with an unusual combination of morphological characters: the transverse muscles present in both the trunk and the tail sections of the body; the seminal vesicles simple, without internal complex compartments; the presence of two pairs of lateral fins. The only family assigned to the new order, Biphragmosagittidae fam. nov., contains two genera. Diagnoses of the two new genera, Biphragmosagitta gen. nov. (type species B. tarasovi sp. nov. and B. angusticephala sp. nov.) and Biphragmofastigata gen. nov. (type species B. fastigata sp. nov.), detailed descriptions and pictures of the three new species are presented.


Zootaxa ◽  
2009 ◽  
Vol 2205 (1) ◽  
pp. 1-36 ◽  
Author(s):  
MICHAEL HOFFMANN ◽  
PETER GRUBB ◽  
COLIN P. GROVES ◽  
RAINER HUTTERER ◽  
ERIK VAN DER STRAETEN ◽  
...  

We provide a synthesis of all mammal taxa described from the African mainland, Madagascar and all surrounding islands in the 20 years since 1988, thereby supplementing the earlier works of G.M. Allen (1939) and W.F.H. Ansell (1989), and bringing the list of African mammals described over the last 250 years current to December 2008. We list 175 new extant taxa, including five new genera, one new subgenus, 138 new species and 31 new subspecies, including remarks, where relevant, on the current systematic position of each taxon. Names of seven species of primates are emended, according to the requirements of the ICZN. The taxonomic group in which the largest number of new taxa has been described is the Primates, with two new genera, 47 new species and 11 new subspecies, while geographically the biggest increase in new species descriptions has been on the island of Madagascar, accounting for roughly half (67) of all new species described in the past 20 years. Nearly half of all new species listed currently are assessed as Data Deficient on the IUCN Red List of Threatened Species (49 of 101 listed species) suggesting further research is urgently needed to help clarify the status of those recently described species.


1992 ◽  
Vol 11 (1) ◽  
pp. 85-93 ◽  
Author(s):  
Martin R. Langer

Abstract. Two new genera and eight new species of benthic foraminifera are described from the shallow water, tropical lagoon of Madang, Papua New Guinea. The new hauerinid genus Pseudolachlanella is characterized by juvenile cryptoquinqueloculine, adult almost massiline arranged chambers, and a slitlike, curved aperture with parallel sides and a long, slender, curved miliolid tooth. Pitella haigi n. gen., n. sp. is a new foraminifera with cryptoquinqueloculine arranged chambers, an almost entirely pitted shell surface (pseudopores) and a rounded aperture with a short simple tooth. Among the other species described as new are four hauerinids and two agglutinated foraminifera All new species described here occur sporadically in the shallow water back- and forereef environments of the lagoon (0–55m), and live infaunally and epifaunally in well-oxygenated, fine and coarse grained biogenic sediments. They are absent in muddy, organic-rich, low-oxygen sedimentary environments within bay inlets where variations of salinity are considerable.


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