THE SYSTEMATICS AND PHYLOGENY OF THE STENOCHIRONOMUS COMPLEX (XESTOCHIRONOMUS, HARRISIUS, and STENOCHIRONOMUS) (DIPTERA: CHIRONOMIDAE)

1984 ◽  
Vol 116 (S128) ◽  
pp. 5-270 ◽  
Author(s):  
Art Borkent

AbstractThe Stenochironomus complex contains all those species which exhibit the highly modified, mining larval form previously recognized as typical of only the genus Stenochironomus Kieffer: larvae with a dorsoventrally compressed head capsule, expanded thoracic segments and a long, flaccid abdomen. The complex includes members of Xestochironomus Sublette & Wirth (New World), Harrisius Freeman (Australian Region), Stenochironomus (cosmopolitan) and two larval forms, one from New Zealand and the other from Africa.This revision provides analysis of 12 species of Xestochironomus (five new), four species of Harrisius, and 54 species of Neotropical and Holarctic Stenochironomus (32 new named species, two new species not named, and four new combinations: S. innocuus (Williston), S. semifumosus (Edwards), S. longilobatus (Tokunaga) and S. gracilivalva (Kieffer)). Of these species, the pupae and fourth instar larvae are described for two species of Xestochironomus, Harrisius pallidus Freeman and 24 of the named species of Stenochironomus. Descriptions, diagnoses, keys, and distribution maps are provided for all species in each available stage. Stenochironomus taeniapennis (Coquillett, 1901), S. zonopterus (Mitchell, 1908), and S. exquisitus (Mitchell, 1908) are newly recognized as junior synonyms of S. hilaris (Walker, 1848).A cladistic analysis of all these species suggests that the Stenochironomus complex is monophyletic. The African larval form is the sister group of all other members; Xestochironomus is the sister group of Harrisius, the New Zealand larval form and Stenochironomus; and Stenochironomus is the sister group of Harrisius plus the New Zealand larval form. These relationships suggest possible southern continental affinities. Stenochironomus is composed of two monophyletic species groups, the subgenus Stenochironomus s. str. with species whose larvae mine wood and the new subgenus Petalopholeus (type-species, S. cinctus Townes) with species whose larvae mine leaves.Cladistic patterns show that Caribbean species are related to other New World species, and that North American species are related to Neotropical or Palaearctic species.Larval food type and ecological differences are analyzed using cladistic relationships. The plesiotypic habitat for species of the Stenochironomus complex is running waters.The phylogenetic relationships of Stenochironomus species and their mite parasites are compared and show little congruence.

2007 ◽  
Vol 21 (1) ◽  
pp. 73 ◽  
Author(s):  
Jie Wu ◽  
Hong-Zhang Zhou

The genus Priochirus Sharp (Coleoptera : Staphylinidae) is a rove beetle group with great variation in cephalic structures, especially the teeth on the frontal margin of head. In this paper, phylogenetic relationships among ten subgenera of the genus Priochirus sensu lato, and of Priochirus and other genera of the tribe Leptochirini are analysed. A cladistic analysis was conducted based on 62 characters of adult morphology. Results indicate that the genus Priochirus is not a monophyletic group and is divided into two lineages (called here Lineages I and II respectively). The subgenus Euleptarthrus Jakobson, which included three species-groups (japonicus, longicornis and malayanus), is shown to be polyphyletic. Lineage I corresponds with the Leiochirus + (Exochirus + Syncampsochirus) + (Eutriacanthus + (Plastus + (japonicus-group of Euleptarthrus + (Barychirus + Stigmatochirus)))) clade and forms a sister group to the genus Thoracochirus Bernhauer. Lineage II corresponds with the Priochirus, s. str. + (Cephalomerus + (longicornis- and malayanus-group of the Euleptarthrus)) clade. In addition, the analysis reveals sister relationships between representatives of Lineage I and the genus Thoracochirus. On the basis of phylogenetic inference, the genus Plastus Bernhauer, stat. nov. is proposed to include the subgenera of Lineage I and a new subgenus Sinumandibulus, subgen. nov., which is erected for the japonicus-group of the former subgenus Euleptarthrus. The genus Priochirus Sharp sensu novo is redefined to include the subgenera of Lineage II. Seven new species are described from China: Plastus (Sinumandibulus) curvaticornis, sp. nov. from Sichuan; Plastus (Sinumandibulus) recticornis, sp. nov. from Hubei; Priochirus (Euleptarthrus) amblyodontus, sp. nov. from Hunan; Priochirus (Euleptarthrus) baoxingensis, sp. nov. from Sichuan, Priochirus (Euleptarthrus) elongates, sp. nov. from Guizhou, Priochirus (Euleptarthrus) parvicornis, sp. nov. from Fujian and Priochirus (Euleptarthrus) oxygonus, sp. nov. from Hainan.


2021 ◽  
Vol 742 ◽  
pp. 1-77
Author(s):  
Luiz Fernando M. De Oliveira ◽  
Antonio Domingos Brescovit

The diversity of the genus Tafana Simon, 1903 is poorly known in the Neotropical regions. In this work we provide a taxonomic review of the genus as well as a phylogenetic analysis. The ingroup of the analysis is composed of sixteen species of Tafana and the outgroup is composed of five representatives of Anyphaenidae. The sister-group recovered for Tafana is the clade Aysha + Xiruana, being supported by the embolic process on the male bulb. Two species groups within Tafana are herein proposed, the silhavyi group and the riveti group, based on two exclusive synapomorphies in the male bulb. We redescribe Tafana quelchi and present a description of the previously unknown female of Tafana silhavyi, both from Venezuela. In addition, we describe the first adult specimens of Tafana straminea. Twelve new species, along with several previously described species, are described, illustrated and mapped: T. riveti, T. straminea, T. quelchi, T. kunturmarqa sp. nov., T. humahuaca sp. nov., T. pastaza sp. nov., T. nevada sp. nov., T. huatanay sp. nov. and T. ruizi sp. nov. from the riveti species group; T. maracay sp. nov., T. arawak sp. nov., T. chimire sp. nov. and T. pitieri sp. nov. from the silhavyi species group; T. oliviae sp. nov. from Argentina and T. orinoco sp. nov. from Venezuela, neither of which belongs to any species group. We also discuss the genital morphology of the species groups based on the results of the phylogenetic analysis. Furthermore, distribution maps for all species, including new records for T. riveti, T. straminea and T. quelchi, are presented.


2017 ◽  
Vol 48 (2) ◽  
pp. 97-155 ◽  
Author(s):  
Maurizio Biondi ◽  
Roberta Frasca ◽  
Elizabeth Grobbelaar ◽  
Paola D’Alessandro

The supraspecific taxonomy of the species traditionally attributed to the flea beetle genusBlepharidaChevrolat, 1836 is discussed. A cladistic analysis, based on 30 morphological characters of traditionalBlepharidaspecies, has revealed that two genera occur in Sub-Saharan Africa:CalothecaHeyden, 1887 andBlepharidinaBechyné, 1968. The latter genus is known from Africa, and probably also Madagascar, and has two subgenera:Blepharidinas.str. andAfroblepharidasubgen. nov. Twenty-seven traditionalBlepharidaspecies are here attributed to the genusCalothecaHeyden, while eighteen species are assigned to the genusBlepharidinaBechyné. FourBlepharidinaspecies,antinorii(Chapuis, 1879),gedyei(Bryant, 1948),scripta(Weise, 1904) andsomaliensis(Bryant, 1948), belong to the new subgenusAfroblepharida. The following new synonymies are established:Eutheca conradsiWeise, 1906= Eutheca erlangeriWeise, 1907 syn. nov. =Blepharidella irregularisBryant, 1945 syn. nov.;Blepharida marginalisWeise, 1902 =Blepharida monticolaWeise, 1926 syn. nov. =Blepharida ugandaeBryant, 1944 syn. nov.;Blepharida inornataJacoby, 1895 =Blepharida semisulcataAchard, 1922 syn. nov.;Blepharidella lewiniWeise in Lewin, 1912 =Blepharidella picticollisBryant, 1945 syn. nov.;Podontia nigrotessellataBaly, 1865= Blepharidella rubrosignataBryant, 1945 syn. nov.= Blepharidella variabilisBryant, 1945 syn. nov.;Blepharida ornataBaly, 1881= Blepharida freyiBechyné, 1954 syn. nov.;Podontia reticulataBaly, 1865= Blepharida guttulaBryant, 1944 syn. nov.;Blepharida antinoriiChapuis, 1879 =Blepharida sudanicaBryant, 1944 syn. nov.;Blepharida scriptaWeise, 1904= Blepharida geminataBryant, 1944 syn. nov. In addition:Blepharida plagipennisAchard, 1922, its locality certainly mislabeled, is transferred to the New World genusNotozonaChevrolat, 1837;Calotheca thunbergiis proposed as the new name forBlepharida stolida(Thunberg, 1808). Finally, an updated catalogue of the known species ofCalothecaandBlepharidinais also supplied, including new synonymies, material examined, new faunistic records, distributions and chorotypes.


1970 ◽  
Vol 102 (S69) ◽  
pp. 9-216 ◽  
Author(s):  
B. V. Peterson

AbstractThe 33 species of Prosimulium Roubaud known from Canada and Alaska are reviewed. These are segregated into four subgenera: Distosimulium new subgenus, containing one species; Parahelodon new subgenus, containing three species; Helodon Enderlein, containing seven species of which clavatum, martini, and susanae are new, and one (known only from the larva) remains undescribed; Prosimulium s. str., containing 25 species of which approximatum, constrictistylum, impostor, mysticium, neomacropyga, and woodorum are new. Species of the last subgenus are segregated into three species groups. A lectotype is designated for P. ursinum (Edwards). Descriptions, distribution maps, keys, and 158 illustrations for the identification of the known adults, pupae, and larvae are provided.


Zootaxa ◽  
2009 ◽  
Vol 2259 (1) ◽  
pp. 1-159 ◽  
Author(s):  
GARY A. P. GIBSON

Spalangia Latreille and Playaspalangia Yoshimoto, the only two genera classified in Spalangiinae (Chalcidoidea: Pteromalidae) are revised for the New World. Thirty-one species of Spalangia and a single species of Playaspalangia, P. rothi Yoshimoto, are recognized from the New World. One further name, Spalangia tarsalis Brèthes, is treated as a nomen dubium. European species of Spalangia not yet known from the New World are also discussed and illustrated relative to regional species, but are not treated formally. The following new synonymies are proposed: Spalangia erythromera brachyceps Bouček under Spalangia erythromera Förster n. syn., Spalangia bakeri Kieffer under Spalangia chontalensis Cameron n. syn., and Spalangia lanaiensis Ashmead under Spalangia impunctata Howard n. syn. Spalangia simplex Perkins is newly recorded from the New World and the following 13 species are described as new: Spalangia alyxia n. sp., Spalangia flavicrus n. sp., Spalangia imitator n. sp., Spalangia innuba n. sp., Spalangia leiopleura n. sp., Spalangia masneri n. sp., Spalangia nigroides n. sp., Spalangia noyesi n. sp., Spalangia plaumanni n. sp., Spalangia rugosifrons n. sp., Spalangia stictocephala n. sp., Spalangia stictocyla n. sp., and Spalangia xanthoscapa n. sp. The 2 genera and the 32 regional species of Spalangiinae are keyed, described and illustrated, and regional hosts and distributions are summarized for each species. Distribution maps document the known range of each of the species in the New World. For purposes of comparison of morphologically similar species, 7 informal species groups are recognized in Spalangia to include 29 of the 31 New World species: the attae-group (S. attae Burks, S. rugosifrons, S. stictocyla and S. stictocephala), the cameroni-group (S. cameroni Perkins, S. longepetiolata Bouček and S. gemina Bouček), the drosophilae- group (S. bethyloides Bouček, S. drosophilae Ashmead, S. flavicrus, S. impunctata, S. innuba, S. leiopleura and S. plaumanni), the endius group (S. endius Walker and S. nigripes), the nigra-group (S. alyxia, S. chontalensis, S. masneri, S. nigra Latreille, S. nigroides and S. nigroaenea Curtis), the noyesi-group (S. noyesi and S. xanthoscapa), and the subpunctata-group (S. erythromera, S. fuscipes, S. haematobiae Ashmead, S. imitator and S. subpunctata Förster).


Zootaxa ◽  
2008 ◽  
Vol 1737 (1) ◽  
pp. 1 ◽  
Author(s):  
S. BLAIR HEDGES ◽  
WILLIAM E. DUELLMAN ◽  
MATTHEW P. HEINICKE

New World frogs recently placed in a single, enormous family (Brachycephalidae) have direct development and reproduce on land, often far away from water. DNA sequences from mitochondrial and nuclear genes of 344 species were analyzed to estimate their relationships. The molecular phylogeny in turn was used as the basis for a revised classification of the group. The 882 described species are placed in a new taxon, Terrarana, and allocated to four families, four subfamilies, 24 genera, 11 subgenera, 33 species series, 56 species groups, and 11 species subgroups. Systematic accounts are provided for all taxa above the species level. Two families (Craugastoridae and Strabomantidae), three subfamilies (Holoadeninae, Phyzelaphryninae, and Strabomantinae), six genera (Bryophryne, Diasporus, Haddadus, Isodactylus, Lynchius, and Psychrophrynella), and two subgenera (Campbellius and Schwartzius) are proposed and named as new taxa, 13 subspecies are considered to be distinct species, and 613 new combinations are formed. Most of the 100 informal groups (species series, species groups, and species subgroups) are new or newly defined. Brachycephalus and Ischnocnema are placed in Brachycephalidae, a relatively small clade restricted primarily to southeastern Brazil. Eleutherodactylidae includes two subfamilies, four genera, and five subgenera and is centered in the Caribbean region. Craugastoridae contains two genera and three subgenera and is distributed mainly in Middle America. Strabomantidae is distributed primarily in the Andes of northwestern South America and includes two subfamilies, 16 genera, and three subgenera. Images and distribution maps are presented for taxa above the species level and a complete list of species is provided. Aspects of the evolution, biogeography, and conservation of Terrarana are discussed.


Zootaxa ◽  
2010 ◽  
Vol 2425 (1) ◽  
pp. 45 ◽  
Author(s):  
ÂNGELO PARISE PINTO ◽  
ALCIMAR DO LAGO CARVALHO

Lauromacromia melanica sp. nov. from Conceição da Barra municipality, Espírito Santo State, Brazil, is described and illustrated based on two males (both in MNRJ nº 135). The new species is similar to L. picinguaba differing from it mainly by the absence of pale spots on S3–6 and by the ellipsoid shape of metepisternal pale stripe. A key for males of all species of the genus is provided. A cladistic analysis encompassing 43 external morphological male characters carried out in two distinct procedures, the first with all characters unordered and the second with two or three state characters ordered. The unordered analysis generated only one most-parsimonious tree (66 steps of length, CI = 0.69, RI = 0.62). The hypothesis of monophyly of Lauromacromia is supported and includes three groups, one formed by the Atlantic Forest species (L. melanica sp. nov. + L. picinguaba), and another by the Cerrado species (L. flaviae + (L. bedei + L. luismoojeni)), and L. dubitalis, positioned in polytomy with these two groups. The ordered analysis also generated only one most-parsimonious tree (68 steps of length, CI = 0.70, RI = 0.67), which maintained the monophyly of Lauromacromia but L. dubitalis positioned basally as sister-group to the Atlantic Forest + Cerrado species groups. The geographic distribution of Lauromacromia is updated with a new record of L. luismoojeni based on one adult male (Brazil: Mato Grosso do Sul State) and probable first Brazilian records for L. dubitalis (Amazonas and Pará States) based on two larvae. A vicariance hypothesis is proposed to explain spatial evolution of Lauromacromia, and based on current biogeographical classifications we consider Gomphomacromia and Rialla apart from Neotropical biota. Some aspects of biology and ecology of Lauromacromia are also discussed.


1992 ◽  
Vol 23 (3) ◽  
pp. 233-248 ◽  
Author(s):  
Knut Rognes

AbstractWithin Pollenia Robineau-Desvoidy a venturii species-group is defined and revised. It consists of a single species P. venturii Zumpt. P. solitaria Grunin is proposed as a junior synonym. It is characterized by unique features in the male aedeagus and the lateral sacs of the internal female reproductive organs. Male and female terminalia are illustrated, the latter for the first time. A preliminary cladistic analysis of all known Palaearctic species of Pollenia (except P. japonica Kano & Shinonaga) suggests that the sister-group of P. venturii is a clade consisting of the viatica + vagabunda + amentaria + haeretica species-groups. A sclerotized internal wall of the lateral sacs in the internal reproductive system of female Pollenia appears to be a parallelism developed independently in the venturii, rudis, most members of the tenuiforceps and some members of the semicinerea groups, rather than an underlying synapomorphy. P. venturii is known from France, Germany, Greece, Italy and Russia. A key is provided to species-groups in Pollenia.


2009 ◽  
Vol 23 (4) ◽  
pp. 301 ◽  
Author(s):  
Mírian Nunes Morales ◽  
Luciane Marinoni

Palpada Macquart, 1834 is a large genus of tropical Eristaliini flies with distribution restricted to the New World (Diptera : Syrphidae : Eristalinae). Most species were initially placed within Eristalis Latreille, 1804. In 1972, most Neotropical species were transferred to Palpada when F.C. Thompson divided Eristalis sensu lato into three genera: Eristalis, Eoseristalis Kanervo, 1938 and Palpada. The latter genus was divided by Thompson (1981) into three groups of species: scutellaris, agrorum and vinetorum. In the present study, the species of the scutellaris group are revised and the phylogenetic relationships among its species are reconstructed based on adult morphology. The cladistic analysis of the 24 terminal species and 48 characters of external morphology and male and female genitalia, produced one cladogram under equal weights. The monophyly of the scutellaris group was corroborated. Twelve species are recognised within the group, of which one is described as new: Palpada cordiformis Morales, sp. nov., P. claudia (Curran), P. conica (Fabricius), P. doris (Curran), P. erratica (Curran), P. fuliginosa (Hull), P. megafemur Thompson, P. melanaspis (Wiedemann), P. precipua (Williston), P. rufoscutellata (Sack), P. scutellaris (Fabricius) and P. vera (Hull). Two new synonymies are proposed: P. dorothea (Hull, 1942) = P. melanaspis (Wiedemann, 1830); and P. glabella (Hull, 1942) = P. fuliginosa (Hull, 1935). The lectotype is designated for Eristalis melanaspis. An identification key, a catalogue, distribution maps, illustrations of general aspects, illustrations of male and female genitalia and scanning electron micrographs are provided.


Zootaxa ◽  
2009 ◽  
Vol 2270 (1) ◽  
pp. 1-38 ◽  
Author(s):  
IAN CHRISTOPHER STOCKS

The monophyletic agelenid genus Barronopsis Chamberlin & Ivie is revised to include 6 species. The Cuban species B. campephila Alayón and B. cesari Alayón are synonomized under B. barrowsi (Gertsch) and B. jeffersi (Muma), respectively, and B. stephaniae new species is described. Natural history observations, distribution maps, diagnoses and descriptions, and a species identification key including B. texana (Gertsch), B. arturoi Alayón, and B. floridensis (Muma) are provided. Detailed descriptions of the male palpus and female genitalia, a review and evaluation of historical terminology used to describe agelenid palpal bulbs, and a discussion of the utility of certain male palpal characters in resolving phylogeny within Agelenidae are provided. Based on the morphology of the male and female genitalia and morphometric data, two species groups are recognized: a large-bodied B. texana species group (B. texana, floridensis, arturoi, jeffersi) and a small-bodied B. barrowsi species group (B. barrowsi, B. stephaniae). A cladistic analysis of Barronopsis, using Tortolena glaucopis (F. O. P.-Cambridge), Melpomene singula (Gertsch & Ivie), and species of Agelenopsis Giebel as outgroups identified three most parsimonious trees of 37 steps. The strict consensus tree yielded the following species relationships: (Agelenopsis (((B. texana, B. jeffersi), B. floridensis, B. arturoi), (B. barrowsi, B. stephaniae))))).Key words: Agelenopsis, revision, taxonomy, phylogenetic analysis


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