GENERIC REVISION OF THE CLINOCERINAE (EMPIDIDAE), AND DESCRIPTION AND PHYLOGENETIC RELATIONSHIPS OF THE TRICHOPEZINAE, NEW STATUS (DIPTERA: EMPIDOIDEA)

1995 ◽  
Vol 127 (5) ◽  
pp. 665-752 ◽  
Author(s):  
Bradley J. Sinclair

AbstractSixteen genera are recognized in the Clinocerinae (Diptera: Empididae): Dipsomyia Bezzi, Proagomyia Collin, Asymphyloptera Collin, Proclinopyga Melander, Rhyacodromia Saigusa, Trichoclinocera Collin, Clinocera Meigen, Oreothalia Melander, Roederiodes Coquillett, Clinocerella Engel, new status, Hypenella Collin, Kowarzia Mik, Dolichocephala Macquart, Phaeobalia Mik, Bergenstammia Mik, and Wiedemannia Zetterstedt. The subgenera of Clinocera are no longer recognized and the genus is divided into six tentative informal species groups. The following new generic synonyms are proposed: Hydrodromia Macquart = Clinocera; Archiclinocera Frey and Hypoclinocera Frey = Kowarzia; Lamposoma Becker, Fur Garrett Jones, and Obstinocephala Garrett Jones = Dolichocephala. A key to genera is provided for the identification of adult specimens of Clinocerinae, all genera are described, several lectotypes are designated, and male and female terminalia are illustrated. A world list of species is included for each genus, identifying new combinations. Cladistic relationships of the genera of the Clinocerinae are presented.Zanclotus Wilder is transferred from the Clinocerinae to the Oreogetoninae. Dryodromia Rondani, also formerly in Clinocerinae, is considered to be most closely related to Hesperempis Melander, both of which are hypothesized to share apomorphies with the Empidinae + Hemerodromiinae. The genera Boreodromia Coquillett, Ceratempis Melander, Ephydrempis Saigusa, Niphogenia Melander, Sabroskyella Wilder, and Trichopeza Rondani also are removed from the Clinocerinae and transferred to the subfamily Trichopezinae, new status. In addition, Apalocnemis Philippi, Gloma Meigen, Hyperperacera Collin (all formerly in Oreogetoninae) and Heleodromia Haliday (formerly in Hemerodromiinae) are transferred to the Trichopezinae. Heterophlebus Philippi (formerly in Oreogetoninae) and Rubistella Garrett Jones (formerly in Brachystomatinae) are removed from synonymy and also transferred to the Trichopezinae. This subfamily is described, and the phylogenetic relationships and.geographic distribution of the included genera are discussed. The Trichopezinae is considered most closely related to the Brachystomatinae (exclusive of Homalocnemis Philippi) and Ceratomerinae. These three subfamilies are hypothesized to represent the sister group to the Microphorinae + Dolichopodidae.

1992 ◽  
Vol 124 (6) ◽  
pp. 951-998 ◽  
Author(s):  
J.M. Cumming ◽  
B.E. Cooper

AbstractThis revision recognizes 13 species of Stilpon Loew in the Nearctic Region, including nine new species. A key is provided for the identification of adult specimens, all species are described, and their geographical distributions are mapped. Two informal species groups are recognized in the Nearctic Region, with the following included species: (1) S. graminum group — S. campestris Cumming sp.nov. (type-locality Assiniboia, Saskatchewan, Canada); (2) S. varipes group — S. chillcotti Cumming sp.nov. (type-locality 10 km E Evergreen, Alabama, USA), S. ctenistes Cumming sp.nov. (type-locality Salmon Creek, New York, USA), S. curvipes Melander, S. limitaris Cumming sp.nov. (type-locality Sapelo Island,Georgia,USA), S. pauciseta Melander, S. pilomus Cumming sp.nov. (type-locality Franconia, New Hampshire, USA), S. spinipes Melander, S. tribulosus Cumming sp.nov. (type-locality Gatlinburg, Tennessee, USA), S. tyconyx Cumming sp.nov. (type-locality Bedford, Massachusetts, USA), S. varipes Loew, S. vockerothi Cumming sp.nov. (type-locality Ottawa, Ontario, Canada), and S. wirthi Cumming sp.nov. (type-locality Dennisport, Massachusetts, USA). A world list of described species of Stilpon, including synonyms, is provided. Stilpon pectiniger Melander is a new junior synonym of S. varipes Loew and S. demnatensis Vaillant is indicated as a nomen nudum. Stilpon pleuriticus Melander, previously considered to belong within Stilpon, is excluded from the genus, and S. obscuripes Adams is transferred as Crossopalpus obscuripes (Adams) comb.nov. Lectotypes are designated for S. pauciseta Melander and S. pectiniger Melander. Homologies of previously confused structures of the male and female terminalia are summarized with reference to other Empidoidea. The monophyly of Stilpon is justified, including discussion of the phylogenetic relationships of the genus to the remainder of the Tachydromiinae. All described species of Stilpon, including extralimital species, are assigned to one of three proposed informal species groups (S. divergens, S. graminum, or S. varipes group), and zoogeographic patterns are discussed.


Zootaxa ◽  
2007 ◽  
Vol 1488 (1) ◽  
pp. 1-164 ◽  
Author(s):  
IGOR V. SHAMSHEV ◽  
PATRICK GROOTAERT

The genus Elaphropeza Macquart from the Oriental region is revised. In addition to the 79 known species (including seven new combinations of species previously placed within Drapetis Meigen) 51 new species are described and illustrated. The study is mainly based on freshly collected material in Singapore (544 samples, year cycle in eight stations and numerous hand captures). Smaller samples were available from Malaysia, Thailand and Cambodia. Of the 51 new species only 43 have been given a name: E. acantha sp. nov., E. asexa sp. nov., E. asiophila sp. nov., E. belumut sp. nov., E. benitotani sp. nov., E. bezzii sp. nov., E. bulohensis sp. nov., E. chekjawa sp. nov., E. combinata sp. nov., E. crassicercus sp. nov., E. darrenyeoi sp. nov., E. demeijerei sp. nov., E. equalis sp. nov., E. feminata sp. nov., E. flavicaput sp. nov., E. furca sp. nov., E. limosa sp. nov., E. luanae sp. nov., E. luteoides sp. nov., E. malayensis sp. nov., E. meieri sp. nov., E. melanderi sp. nov., E. modesta sp. nov., E. monacantha sp. nov., E. monospina sp. nov., E. murphyi sp. nov., E. neesoonensis sp. nov., E. ngi sp. nov., E. pauper sp. nov., E. pluriacantha sp. nov., E. riatanae sp. nov., E. sime sp. nov.,E. singaporensis sp. nov., E. singulata sp. nov., E. sivasothii sp. nov., E. spicata sp. nov., E. spiralis sp. nov., E. sylvicola sp. nov., E. temasek sp. nov., E. tiomanensis sp. nov., E. ubinensis sp. nov., E. yangi sp. nov., E. yeoi sp. nov. Eight species known only as females are briefly diagnosed but not named. Species (including type materials in most cases) described by C.R. Osten-Sacken, M. Bezzi, J.C.H. de Meijere, R. Senior-White, A.L. Melander and K.G.V. Smith were examined and re-described. Neotypes have been designated for E. calcarifera Bezzi, E. melanura Bezzi, E. formosae Bezzi and E. scutellaris Bezzi. Elaphropeza exul Osten-Sacken, 1882 is transferred as Crossopalpus exul (Osten-Sacken) comb. nov. Elaphropeza formosae Bezzi sensu Quate, 1960 re-described by L.W. Quate from Micronesia, is considered a new species and a new name is given: E. quatei sp. nov. All species of Elaphropeza are divided into two main species groups: E. biuncinata group and E. ephippiata group. Phylogenetic relationships within Elaphropeza are provisionally outlined. Some interesting morphological features and phenology of the species are discussed. Fourteen species have been found exclusively in mangroves. A key is given to the species of the Oriental Region (Indian subcontinent, Southeast Asia, Philippines and Taiwan, but excluding certain species of the Chinese mainland that proved to be different from the rest of the Oriental Region). A check list of the Oriental species is provided.


2007 ◽  
Vol 55 (3) ◽  
pp. 197 ◽  
Author(s):  
Paul M. Oliver ◽  
Mark N. Hutchinson ◽  
Steven J. B. Cooper

Diplodactylid geckos offer a model system for investigating the biogeographic history of Australia and adaptive radiations in the arid zone, but there is considerable uncertainty in the systematics of several key genera. We used sequence data from mitochondrial DNA to carry out a comprehensive analysis of phylogenetic relationships of geckos in the genus Diplodactylus. Parsimony and Bayesian analyses were highly concordant and allocated all species to one of two monophyletic clades, one comprising the species placed in the vittatus and conspicillatus species groups, the other comprising species placed in the stenodactylus and steindachneri species groups, plus D. byrnei, formerly in the vittatus group. The distinctness of these two clades is supported by external morphology of the digits, body and limb proportions, and osteology of the bones in the orbital region, and we use these characters to formally define the two clades as genera. We revive and expand the genus Lucasium for D. byrnei, D. steindachneri and the stenodactylus group, with the other species staying in a redefined Diplodactylus. The monotypic Rhynchoedura is distinct from Lucasium, although the Bayesian mtDNA analysis (but not parsimony) gives some support for a sister-group relationship between Lucasium and Rhynchoedura. Molecular data suggest that each of these clades represents a distinct radiation into semiarid and arid terrestrial habitats during the mid-Tertiary, well before the hypothesised Pliocene onset of major aridification.


2007 ◽  
Vol 21 (1) ◽  
pp. 73 ◽  
Author(s):  
Jie Wu ◽  
Hong-Zhang Zhou

The genus Priochirus Sharp (Coleoptera : Staphylinidae) is a rove beetle group with great variation in cephalic structures, especially the teeth on the frontal margin of head. In this paper, phylogenetic relationships among ten subgenera of the genus Priochirus sensu lato, and of Priochirus and other genera of the tribe Leptochirini are analysed. A cladistic analysis was conducted based on 62 characters of adult morphology. Results indicate that the genus Priochirus is not a monophyletic group and is divided into two lineages (called here Lineages I and II respectively). The subgenus Euleptarthrus Jakobson, which included three species-groups (japonicus, longicornis and malayanus), is shown to be polyphyletic. Lineage I corresponds with the Leiochirus + (Exochirus + Syncampsochirus) + (Eutriacanthus + (Plastus + (japonicus-group of Euleptarthrus + (Barychirus + Stigmatochirus)))) clade and forms a sister group to the genus Thoracochirus Bernhauer. Lineage II corresponds with the Priochirus, s. str. + (Cephalomerus + (longicornis- and malayanus-group of the Euleptarthrus)) clade. In addition, the analysis reveals sister relationships between representatives of Lineage I and the genus Thoracochirus. On the basis of phylogenetic inference, the genus Plastus Bernhauer, stat. nov. is proposed to include the subgenera of Lineage I and a new subgenus Sinumandibulus, subgen. nov., which is erected for the japonicus-group of the former subgenus Euleptarthrus. The genus Priochirus Sharp sensu novo is redefined to include the subgenera of Lineage II. Seven new species are described from China: Plastus (Sinumandibulus) curvaticornis, sp. nov. from Sichuan; Plastus (Sinumandibulus) recticornis, sp. nov. from Hubei; Priochirus (Euleptarthrus) amblyodontus, sp. nov. from Hunan; Priochirus (Euleptarthrus) baoxingensis, sp. nov. from Sichuan, Priochirus (Euleptarthrus) elongates, sp. nov. from Guizhou, Priochirus (Euleptarthrus) parvicornis, sp. nov. from Fujian and Priochirus (Euleptarthrus) oxygonus, sp. nov. from Hainan.


2014 ◽  
Vol 46 (3) ◽  
pp. 441-453 ◽  
Author(s):  
Christian PRINTZEN

AbstractThe lichen genus Biatora comprises inconspicuous crustose lichens that are typically found on organic substrata such as tree bark, bryophytes and detritus. During the last 20 years many new species have been added to the genus making its delimitation more and more difficult. The infrageneric relationships of the 42 species have never been investigated thoroughly. Using DNA sequences from three gene loci (ITS, RPB2, mrSSU) and 59 OTUs, an attempt was made to reconstruct the phylogenetic relationships of Biatora and its infrageneric groups. Cliostomum appears to be the closest relative of Biatora. The position of Mycobilimbia in the Lecania-clade is confirmed. Phylogenetic relationships within Biatora are poorly supported, but six different species groups that are also phenotypically distinguished are more or less well supported: the vernalis-, meiocarpa-, hertelii-, ocelliformis-, beckhausii- and rufidula- groups. The analysis also confirms the presence of several undescribed taxa. Biatora subduplex as currently circumscribed appears to be heterogeneous, as does B. helvola. Based on the phylogeny, the distributional range of B. alaskana is extended to Japan. The new combinations Biatora ementiens (Nyl.) Printzen and Biatora hemipolia (Nyl.) S. Ekman & Printzen are made and both names are typified.


Behaviour ◽  
2014 ◽  
Vol 151 (9) ◽  
pp. 1313-1334 ◽  
Author(s):  
Martha L. Tobias ◽  
Jeremy Korsh ◽  
Darcy B. Kelley

In anurans, male clasps can elicit release calls from either sex. Male release calls have been observed in many anuran genera and this vocal response is thus highly conserved. Female release calls, however, are not as prevalent, suggesting that evolutionary trajectories for anuran release calls differ by sex. We analyzed male and female release calls in all available species of Xenopus, a fully aquatic African genus. Phylogenetic relationships in this genus include three species groups, two of which are clades and one of which is characterized by a reticulated phylogeny due in part to hybridizations between species with different ploidy levels (Evans et al., 2004; Evans, 2008). In all species, males produce release calls when clasped by another male. Females in the reticulated group do not produce release calls, but females in the rest of the genus do. Release calls consist of click trains of variable durations and inter-call intervals. In both sexes, inter-click interval divides the genus into groups with different click rates and these groups are phylogenetically related. In general, inter-click interval is shorter in male than in female release calls. Across species and sexes, release calls are characterized by a single, low (∼1000 Hz) dominant frequency. In X. laevis Congo and X. borealis, clasp duration is longer for male–female than for male–male pairs and clasp duration is correlated with the number, but not the duration, of release calls in male–male pairs. We discuss evolutionary scenarios for release call traits as well as the sex difference in occurrence.


2011 ◽  
Vol 59 (5) ◽  
pp. 344
Author(s):  
Paul M. Oliver ◽  
Mark N. Hutchinson ◽  
Steven J. B. Cooper

Diplodactylid geckos offer a model system for investigating the biogeographic history of Australia and adaptive radiations in the arid zone, but there is considerable uncertainty in the systematics of several key genera. We used sequence data from mitochondrial DNA to carry out a comprehensive analysis of phylogenetic relationships of geckos in the genus Diplodactylus. Parsimony and Bayesian analyses were highly concordant and allocated all species to one of two monophyletic clades, one comprising the species placed in the vittatus and conspicillatus species groups, the other comprising species placed in the stenodactylus and steindachneri species groups, plus D. byrnei, formerly in the vittatus group. The distinctness of these two clades is supported by external morphology of the digits, body and limb proportions, and osteology of the bones in the orbital region, and we use these characters to formally define the two clades as genera. We revive and expand the genus Lucasium for D. byrnei, D. steindachneri and the stenodactylus group, with the other species staying in a redefined Diplodactylus. The monotypic Rhynchoedura is distinct from Lucasium, although the Bayesian mtDNA analysis (but not parsimony) gives some support for a sister-group relationship between Lucasium and Rhynchoedura. Molecular data suggest that each of these clades represents a distinct radiation into semiarid and arid terrestrial habitats during the mid-Tertiary, well before the hypothesised Pliocene onset of major aridification.


1980 ◽  
Vol 11 (3) ◽  
pp. 313-342 ◽  
Author(s):  
Leif Lyneborg

AbstractNeotabuda Kröber (1931) is a widespread genus of therevids in southern Africa. Its possible monophyly is discussed, and a sister-group relationship with the Palaearctic genus Salentia A. Costa is motivated. Twenty species are recognized, only five of which were described previously. All species are keyed, described and figured. They are arranged in two species-groups, each with two subgroups, and the phylogenetic relationships of these are discussed. The fifteen new species are: incrassaia (Cape Prov.), latifrons (Cape Prov.), longestylata (Cape Prov.), lanigera (Cape Prov.), pilosa (Cape Prov.), subpilosa (Cape Prov.), turneri (Cape Prov.), multisetosa (Cape Prov.), longicornis (Cape Prov.), tomentosa (Namibia), major (Namibia), natalensis (Natal Prov.), diversicornis (Natal Prov.), truncata (Mozambique, Natal Prov.), and nigropilusa (Cape Prov.). The five previously described species are (in their original combination): Thereva anthracina Loew, 1858; Orthactia nigra Kröber, 1912; Pachygenia nitida Kröber, 1912; Actorthia capensis Kröber, 1931; and Neotabuda ater Kröber, 1931. Pachygenia Kröber, 1912 and Neotabuda Kröber, 1931 are placed in new synonymy, but Pachygenia Kröber cannot be used because of its preoccupation by Pachygenia Motschulsky (1874).


1994 ◽  
Vol 126 (4) ◽  
pp. 1007-1059 ◽  
Author(s):  
Bradley J. Sinclair

AbstractThere are 16 species of Trichoclinocera Collin recognized from the Nearctic region, including eight new species. Four of six species-groups of Trichoclinocera are recognized in the Nearctic region, with the following included species: (1) T. comata group — T. cascadensis sp.nov. (type-locality Starvation Creek, Oregon, U.S.A.), T. comata (Melander) comb.nov., and T. rupestris sp.nov. (type-locality Newton Creek, Wyoming, U.S.A.); (2) T. dolicheretma group — T. dolicheretma (Melander); (3) T. lapponica group — T. lapponica (Ringdahl) comb.nov.; (4) T. longipes group — T. agilis sp.nov. (type-locality Fall Creek Falls, Tennessee, U.S.A.), T. ctenistes (Melander), T. dasycoxa sp.nov. (type-locality Ottawa, Ontario, Canada), T. falcata sp.nov. (type-locality Gatineau Park, Quebec, Canada), T. fumosa (Vaillant) comb.nov., T. hamifera (Melander), T. longipes (Walker). T. minor (Melander), T. ozarkensis sp.nov. (type-locality Haw Creek Falls, Arkansas, U.S.A.), and T. pectinifemur sp.nov. (type-locality Phoenixville, Pennsylvania, U.S.A.). The following new synonyms are proposed: T. brunnipennis (Melander) = T. dolicheretma; and Seguyella saltans (Vaillant) = T. hamifera. Lectotypes are designated for T. comata, T. ctenistes, T. dolicheretma, T. brunnipennis, T. hamifera, T. lapponica, and T. stackelbergi Collin. Seguyella Vaillant and Acanthoclinocera Saigusa are newly synonymized with Trichoclinocera and all species formerly placed in these genera are transferred to Trichoclinocera, except Acanthoclinocera glaucescens (Brunetti) which is transferred as Wiedemannia glaucescens (Brunetti) comb.nov.This revision includes a key to Nearctic species, a description of each species, and maps of species distributions. Homologies of male terminalia of Clinocerinae are summarized and the first descriptions of the immature stages of species of Trichoclinocera also are provided. A world list of described species of Trichoclinocera is provided and all species are assigned to one of six species-groups, except T. cummingi sp.nov. (type-locality Robber’s Cave, Oklahoma. U.S.A.), T. fluviatilis (Brunetti) comb.nov., T. fuscipennis Saigusa, and T. maewa Smith, which remain unplaced. The monophyly of Trichoclinocera is justified and phylogenetic relationships to the remaining Clinocerinae and among species-groups are discussed.


2006 ◽  
Vol 37 (2) ◽  
pp. 213-226 ◽  
Author(s):  
Ljubiša Stanisavljević ◽  
Olivera Petrović-Obradović ◽  
Marija Milutinović ◽  
Snežana Tomanović ◽  
Željko Tomanović ◽  
...  

AbstractPhylogenetic relationships among four genera and 25 species of the tribe Praini (Braconidae: Aphidiinae) were investigated on the basis of 15 characters (13 morphological and two life history characters) using parsimony analysis. Areopraon Mackauer and Pseudopraon Starý are basal within the tribe. Both genera include species in which the external and internal types of pupation exist simultaneously [for example, Pseudopraon mindariphagum Starý, Areopraon lepelleyi (Waterston) and A. pilosum Mackauer)]. It can therefore be asserted that the external type of pupation behavior in the Praini probably originated in the polymorphic pupation behavior of ancient genera such as Pseudopraon and Areopraon. The genera Areopraon, Pseudopraon, and Dyscritulus Hincks form a monophylum separate from Praon. Areopraon is not retrieved as monophyletic. Species of the genus Praon Haliday possess conical apical spines on the top of the ovipositor sheath as an apomorphic character state. Within the genus Praon, we recognized the following closely related species groups: Parapraon + Praon dorsale-yomenae; and Praon rosaecola. Two Praini species are redescribed. Mesopraon Starý is a new synonym of Areopraon. Areopraon silvestre (Starý) and Areopraon helleni (Starý) are new combinations. A key to the recent Praini genera is given.


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