SYNOPSIS OF THE NEOTROPICAL GENUS OZAENA OLIVIER: CLASSIFICATION AND RECONSTRUCTED EVOLUTIONARY HISTORY (COLEOPTERA: CARABIDAE: OZAENINI)

1990 ◽  
Vol 122 (5) ◽  
pp. 779-815 ◽  
Author(s):  
George E. Ball ◽  
Danny Shpeley
Keyword(s):  
South America ◽  
Sister Group ◽  
Structural Features ◽  
Character Evolution ◽  
Type Locality ◽  
Sister Taxon ◽  
Fore Tibia ◽  
Sensilla Basiconica ◽  
Extant Species ◽  
And Shifts

AbstractDescriptions and illustrations of structural features of adults, a key, and chorological data provide the basis for characterizing the genus Ozaena Olivier and classifying the 10 included species, two of which are new: O. maxi (type locality: Iracubo, CAYENNE), and O. manu (type locality: Manu National Park, Madre de Dios Province, PERU). With a geographical range that extends from ca. 30°S to 30°N in the New World, all species of Ozaena occur in cis-Andean South America, only two ranging farther north: O. dentipes Olivier, to Panama; and O. lemoulti Bänninger to southern Arizona, USA. A reconstructed phylogeny postulates the following: the genus Platycerozaena Bänninger as the sister-group of Ozaena, together these genera comprising the Ozaena genus-group; within Ozaena, the O. dentipes group as sister-taxon of the O. lemoulti + convexa groups; within the O. dentipes group, O. maxi as sister-taxon of O. linearis Bänninger + dentipes; within the O. lemoulti group, the sequence O. ecuadorica Bänn. (O. elavata Bänn. [O. lemoulti + martinezi Ogueta]); and within the O. convexa group, O. grossa Bänn. as sister-taxon of O. convexa Bänn. + manu. Character evolution involves losses, gains, and shifts, in about equal numbers. Convergence is relatively rare, involving only three of 35 characters. The predominant mode of character evolution is simple, with few examples of sequential changes sustained through several speciation events. Basic modifications in evolution of the ground-plan of the Ozaena genus-group include reduction or loss (for example, tactile sense organs, antennal cleaner of the fore tibia, and adhesive vestiture of the male fore tarsi), and enhancement or gain (for example, scale-like sensilla basiconica on pronotum and elytra, concentration of sensilla basiconica in groups on the antennomeres, particularly antennomere 11). Within Ozaena, many modifications to the antennae and mouthparts indicate the importance of these organs to evolution of the group.Based on phylogenetic relationships and the distribution pattern of the extant species, a reconstructed geographical history of Ozaena postulates the following: first, a vicariance event in South America separating the ancestral stock of Ozaena into a northern vicar that gave rise to the O. dentipes group, and a southern vicar that gave rise to the O. lemoulti and convexa groups; followed by cycles of range expansion and contraction that allowed for differentiation of successive stocks of species. The temporal range for these speciation events is estimated to extend over a period of about 12 million years, or from the latter part of the Miocene epoch to the Recent. The most recent speciation events are estimated to be of Pliocene age.Based on lack of differentiation of South American and Middle American representatives of O. dentipes and O. lemoulti respectively, and extent of range in Middle America, these two species are postulated to be relatively recent invaders of the latter area, with O. dentipes being the more recent to arrive.

THE SPECIES OF EUCHEILOID PERICALINA: CLASSIFICATION AND EVOLUTIONARY CONSIDERATIONS (COLEOPTERA: CARABIDAE: LEBIINI)

1983 ◽  
Vol 115 (7) ◽  
pp. 743-806 ◽  
Author(s):  
G. E. Ball ◽  
D. Shpeley
Keyword(s):  
New Species ◽  
South America ◽  
New Genus ◽  
Habitat Type ◽  
Sister Group ◽  
Structural Features ◽  
The Body ◽  
Type Locality ◽  
Reproductive Structures ◽  
Middle America

AbstractThe endemic New World eucheiloid complex is distinguished from the other complexes (thyreopteroid, eurycoleoid, somotrichoid, and pericaloid) of pericaline Lebiini by securiform labial palpomere 3, serrate-setose margins of the pronotum, and flattened stylomere 2 of the ovipositor. The geographical range of eucheiloids extends from northern Argentina to southernmost United States (Brownsville, Texas). All taxa are believed to be arboreal. A key distinguishes among the three genera and 16 species, and each taxon is characterized in terms of structural features, habitat, and geographical distribution. The genera are:Hansus, new genus (generitype—H. reichardti, new species);InnaPutzeys (generitype—Inna punctataPutzeys =Polystichus boyeri(Solier)); andEucheilaDejean (generitype—Euchyla flavilabris(Dejean)). Seven new species and one new subspecies are described:Hansus reichardti(type locality—Guyana, EssequiboR.. Morrabali Ck.);Inna palpalis,I. atrata arbor, andI. inpa(type locality—Brazil, Amazonas, near Manaus);I. purpurea(type locality—Brazil, Amazonas, 60 km n. Manaus);I. splendens(type locality—Venezuela, Aragua, Rancho Grande);Eucheila adisi(type locality—Brazil, Amazonas, 60 km n. Manaus); andE. cordova(type locality—México, Veracruz, Cordova). The following new synonymies of specific names are proposed (valid names listed first):Inna costulataChaudoir, 1872 =I. granulataChaudoir, 1872;Inna boyeri(Solier, 1835) =I. punctataPutzeys, 1863, andI. texanaSchaeffer, 1910. A reconstructed phylogeny of the eucheiloids, based on analysis of 60 character states included in 33 characters, shows thatHansusis sister group of the ancestral stock ofInna+Eucheila. Evolutionary modifications have affected principally body size, head and mouthparts (feeding and associated adaptations), reproductive structures, and features of the body surface thought to be associated with avoidance of predators (concealment and flash coloration). Diversification may also have involved differentiation by habitat (type of forest occupied). The principal theater of evolution has been tropical South America, with incursions both southward and northward. Middle America has been invaded by five lineages, probably during Cenozoic time, from Middle Tertiary to the Quaternary. Of these lineages, three are represented in Middle America by endemic species:E. cordova,I. planipennisBates, andI. nevermanniLiebke. Two lineages are represented by species whose ranges extend northward from South America:I. costulataChaudoir andI. boyeri(Solier).

Diversity and Integration in Private International Law

2019 ◽  
Keyword(s):  
South America ◽  
International Law ◽  
Value Systems ◽  
Private International Law ◽  
Diverse Range ◽  
Cross Border ◽  
Wide Range ◽  
Depth Analysis ◽  
And Shifts ◽  
Substantive Law

This book opens a cross-regional dialogue and shifts the Eurocentric discussion on diversity and integration to a more inclusive engagement with South America in private international law issues. It promotes a contemporary vision of private international law as a discipline enabling legal interconnectivity, with the potential to transcend its disciplinary boundaries to further promote the reality of cross-border integration, with its focus on the ever-increasing cross-border mobility of individuals. Private international law embraces legal diversity and pluralism. Different legal traditions continue to meet, interact and integrate in different forms, at the national, regional and international levels. Different systems of substantive law couple with divergent systems of private international law (designed to accommodate the former in cross-border situations). This complex legal landscape impacts individuals and families in cross-border scenarios, and international commerce broadly conceived. Private international law methodologies and techniques offer means for the coordination of this constellation of legal orders and value systems in cross-border situations. Bringing together world-renowned academics and experienced private international lawyers from a wide range of jurisdictions in Europe and South America, this edited collection focuses on the connective capabilities of private international law in bridging and balancing legal diversity as a corollary for the development of integration. The book provides in-depth analysis of the role of private international law in dealing with legal diversity across a diverse range of topics and jurisdictions.

New details of the male terminalia of Diathoneura longipennis (Malloch, 1926) (Diptera, Drosophilidae)

2019 ◽  
Vol 40 (1) ◽  
pp. 45-50
Author(s):  
Carlos R. Vilela ◽  
Gerhard Bächli
Keyword(s):  
Costa Rica ◽  
South America ◽  
Late Stage ◽  
First Record ◽  
Type Locality

The male terminalia of a non-type Diathoneura longipennis (Malloch 1926) specimen,collected in Peru, were dissected and analyzed. The aedeagus and associated sclerites were found tohave been fixed at a late stage of protrusion, resulting in a different morphology, when compared to thetwo previous publications, which were fixed at earlier stages. Consequently, additional details of certainanatomical parts were observed and a better understanding of the aedeagus protruding process was attained.Diathoneura longipennis has been previously identified in San Mateo, Alajuela, Costa Rica (type locality)and Panama, and this specimen represents the first record of this species in South America.

Review of Eohiodon (Teleostei: Osteoglossomorpha) from western North America, with a phylogenetic reassessment of Hiodontidae

1997 ◽  
Vol 71 (6) ◽  
pp. 1109-1124 ◽  
Author(s):  
Li Guo-Qing ◽  
Mark V. H. Wilson ◽  
Lance Grande
Keyword(s):  
North America ◽  
Sister Group ◽  
Sensu Stricto ◽  
Valid Species ◽  
Western North America ◽  
Extant Species ◽  
The Family ◽  
Fossil Records ◽  
Fossil Genus ◽  
Freshwater Teleosts

Review of recently collected material of Eohiodon from North America suggests that there are two valid species, E. rosei (Hussakof) and E. woodroffi Wilson. Eohiodon falcatus Grande is identical to E. woodruffi in known skeletal features and nearly all meristic features and is treated as a junior synonym of the latter. The fossil genus Eohiodon Cavender differs from Hiodon Lesueur, which is known from both fossil and extant species, in numerous meristic and osteological features. The caudal skeleton in Eohiodon is nearly identical to that in Hiodon.The traditionally accepted Notopteroidei, containing Lycopteridae, Hiodontidae, and Notopteridae, is a polypheletic group. The Asian fossil family Lycopteridae is not more closely related to Hiodontidae than it is to other taxa in the Osteoglossomorpha, but is sister to all other Osteoglossomorpha. The Hiodontiformes sensu stricto, including only the family Hiodontidae, is the sister-group of the Osteoglossiformes. This family is not more closely related to notopterids than to other taxa in Osteoglossiformes. The Notopteridae are most closely related to the Mormyroidea; together they and the fossil family Ostariostomidae constitute the sister-group of the Osteoglossoidei.Fossil records of Hiodontiformes sensu stricto and Notopteroidei indicate a widespread pre-Neogene biogeographic range of these freshwater teleosts, suggesting that extinction must have been involved in the Cenozoic evolution of these two osteoglossomorph sublineages.

Checklist and taxonomic changes for Central and South American Philonthina (Coleoptera: Staphylinidae)

Zootaxa ◽  
2018 ◽  
Vol 4449 (1) ◽  
pp. 1 ◽  
Author(s):  
MARIANA CHANI-POSSE ◽  
ALFRED F. NEWTON ◽  
ASLAK KAPPEL HANSEN ◽  
ALEXEY SOLODOVNIKOV
Keyword(s):  
South America ◽  
Original Description ◽  
Junior Synonym ◽  
Type Locality ◽  
Type Material ◽  
New Combination ◽  
South American ◽  
New Combinations ◽  
Generic Composition ◽  
Taxonomic Changes

A checklist of all described species of Philonthina, a subtribe of the staphylinid tribe Staphylinini, known to occur in Central and South America (CASA) is presented. Included for each species, and for synonyms known from CASA, is a reference to the original description, type locality and type depository, and for each species the known distribution within and outside CASA. Type material was sought in the main European and American collections where it is deposited (BMNH, MNHUB, IRSNB and FMNH) and is summarized for all indigenous CASA species, with lectotypes designated for 16 names and confirmation of holotypes and prior designation of lectotypes when necessary. Based on recent phylogenetic work in Philonthina and our revision of types of CASA species of Philonthus Stephens, 1829 and Belonuchus Nordmann, 1837, some taxonomic changes are proposed. Thirty-one species of Philonthus are transferred to Belonuchus (16), Gabrius Stephens 1829 (14), and Bisnius Stephens 1829 (one) resulting in the following new combinations: B. abnormalis (Sharp 1885), B. celatus (Sharp 1885), B. corticalis (Sharp 1885), B. extremus (Sharp 1885), B. infimus (Sharp 1885), B. iteratus (Sharp 1887), B. latecinctus (Sharp 1885), B. lucilius (Sharp 1885), B. muticus (Sharp 1876), B. optatus (Sharp 1885), B. platypterus (Sharp 1885), B. rufiventris (Sharp 1887), B. rufocaudus (Sharp 1885), B. rufopygus (Sharp 1885), B. serraticornis (Sharp 1876), B. supernus (Herman 2001), G. approximans (Sharp 1885), G. armatipes (Sharp 1885), G. atricolor (Sharp 1885), G. championi (Sharp 1885), G. dampfi (Bernhauer 1929), G. elegans (Sharp 1885), G. forsterianus (Scheerpeltz 1960), G. misellus (Sharp 1885), G. nugax (Sharp 1885), G. ovaticeps (Sharp 1885), G. peruvianus (Bernhauer 1916), G. planulatus (Sharp 1885), G. rusticus (Sharp 1885), G. serpens (Sharp 1885) and Bi. subaeneipennis (Bernhauer 1916). Endeius nitidipennis Solier 1849 is transferred to Gabrius, resulting in the following new combination, G. nitidipennis (Solier 1849). Leptopeltus carchiensis Chani-Posse & Asenjo 2013 is proposed as junior synonym of Philonthus divisus Sharp 1891, which is transferred to Leptopeltus Bernhauer 1906 resulting in a new combination: Leptopeltus divisus (Sharp 1891). Belonuchus penetrans Silvestri 1946 is transferred to Pridonius Blackwelder 1952 as a new combination. Lectotypes are designated for Atopocentrum mirabile Bernhauer 1906, Philonthus armatipes Sharp 1885, Ph. atricolor Sharp 1885, Ph. championi Sharp 1885, Ph. misellus Sharp 1885, Ph. planulatus Sharp 1885, Ph. rusticus Sharp 1885, Ph. serpens Sharp 1885, Ph. abnormalis Sharp 1885, Ph. celatus Sharp 1885, Ph. infimus Sharp 1885, Ph. latecinctus Sharp 1885, Ph. muticus Sharp 1876, Ph. platypterus Sharp 1885, Ph. rufocaudus Sharp 1885 and Ph. rufopygus Sharp 1885. Of the 543 currently known species of Philonthina reported from CASA, at least 14 are believed to be adventive from elsewhere, 56 may occur naturally elsewhere, and 473 (87%) are evidently endemic to this region. Of the 31 genera represented by these described species, 20 (65%) are endemic to CASA. One genus, Gabronthus Tottenham 1955, is adventive. However, the actual philonthine fauna of CASA will undoubtedly be much larger, and the generic composition highly modified, when the fauna is fully explored and studied within a phylogenetical framework. 

The phylogenetic position of the tuatara, Sphenodon (Sphenodontida, Amniota), as indicated by cladistic analysis of the ultrastructure of spermatozoa

1992 ◽  
Vol 335 (1274) ◽  
pp. 207-219 ◽  
Keyword(s):  
Cladistic Analysis ◽  
Sister Group ◽  
Sister Taxon ◽  
Phylogenetic Position ◽  
Sister Group Relationship ◽  
Usual Concept ◽  
Cladistic Analyses ◽  
Relationship Of

Sphenodon has traditionally been regarded as a little changed survivor of the Permo-Triassic thecodont or eosuchian ‘stem reptiles’ but has alternatively been placed in the Lepidosauria as the plesiomorphic or even apomorphic sister-taxon of the squamates. A cladistic analysis of 16 characters from spermatozoal ultrastructure of Sphenodon and other amniotes unequivocally confirms its exceedingly primitive status. The analysis suggests that monotremes are the sister-group of birds; squamates form the sister-group of a bird + monotreme clade while the three sister-groups successively below the bird + monotreme + squa- mate assemblage are the caiman, the tuatara and the outgroup (turtles). The monotreme + bird couplet, supports the concept of the Haemothermia, but can only be regarded heuristically. The usual concept of mammals as a synapsid-derived outgroup of all other extant amniotes is not substantiated spermatologically. All cladistic analyses made, and a separate consideration of apomorphies, indicate that Sphenodon is spermatologically the most primitive amniote, excepting the Chelonia. It is advanced (apomorphic) for the amniotes in only two of the 16 spermatozoal characters considered. A close, sister-group relationship of Sphenodon with squamates is not endorsed.

A molecular phylogeny of the circum-Antarctic Opiliones family Neopilionidae

2021 ◽  
Author(s):  
Gonzalo Giribet ◽  
Kate Sheridan ◽  
Caitlin M. Baker ◽  
Christina J. Painting ◽  
Gregory I. Holwell ◽  
...  
Keyword(s):  
New Zealand ◽  
South America ◽  
Morphological Study ◽  
18S Rrna ◽  
New Caledonia ◽  
Sister Group ◽  
28S Rrna ◽  
South American ◽  
Australian Species ◽  
The Family

The Opiliones family Neopilionidae is restricted to the terranes of the former temperate Gondwana: South America, Africa, Australia, New Caledonia and New Zealand. Despite decades of morphological study of this unique fauna, it has been difficult reconciling the classic species of the group (some described over a century ago) with recent cladistic morphological work and previous molecular work. Here we attempted to investigate the pattern and timing of diversification of Neopilionidae by sampling across the distribution range of the family and sequencing three markers commonly used in Sanger-based approaches (18S rRNA, 28S rRNA and cytochrome-c oxidase subunit I). We recovered a well-supported and stable clade including Ballarra (an Australian ballarrine) and the Enantiobuninae from South America, Australia, New Caledonia and New Zealand, but excluding Vibone (a ballarrine from South Africa). We further found a division between West and East Gondwana, with the South American Thrasychirus/Thrasychiroides always being sister group to an Australian–Zealandian (i.e. Australia + New Zealand + New Caledonia) clade. Resolution of the Australian–Zealandian taxa was analysis-dependent, but some analyses found Martensopsalis, from New Caledonia, as the sister group to an Australian–New Zealand clade. Likewise, the species from New Zealand formed a clade in some analyses, but Mangatangi often came out as a separate lineage from the remaining species. However, the Australian taxa never constituted a monophyletic group, with Ballarra always segregating from the remaining Australian species, which in turn constituted 1–3 clades, depending on the analysis. Our results identify several generic inconsistencies, including the possibility of Thrasychiroides nested within Thrasychirus, Forsteropsalis being paraphyletic with respect to Pantopsalis, and multiple lineages of Megalopsalis in Australia. In addition, the New Zealand Megalopsalis need generic reassignment: Megalopsalis triascuta will require its own genus and M. turneri is here transferred to Forsteropsalis, as Forsteropsalis turneri (Marples, 1944), comb. nov.

Identity of Carex trichodes Steud. ex Boott and Carex lateriflora Phil. (subgen. Psyllophorae, sect. Junciformes, Cyperaceae) from Southern South America

Phytotaxa ◽  
2022 ◽  
Vol 530 (2) ◽  
pp. 230-236
Author(s):  
DIEGO N. PENNECKAMP
Keyword(s):  
South America ◽  
Type Locality ◽  
Type Material ◽  
Southern South America ◽  
In The Wild ◽  
Made In

The identity of Carex trichodes, a species only known from the collection of the type material made in 1854 is clarified after finding it in the wild in the vicinity of the type locality. It is concluded that C. trichodes was proposed based on stressed plants corresponding to the same taxon later described as C. lateriflora.

Systematics of Paraleucopis Malloch with proposal of Paraleucopidae, a new family of acalyptrate Diptera

Zootaxa ◽  
2019 ◽  
Vol 4668 (3) ◽  
pp. 301-328
Author(s):  
TERRY A. WHEELER ◽  
BRADLEY J. SINCLAIR
Keyword(s):  
New World ◽  
Current Knowledge ◽  
Sister Group ◽  
Type Locality ◽  
Western United States ◽  
Australian Species ◽  
New Family ◽  
The Family ◽  
Group Relationships ◽  
Northwestern Mexico

Paraleucopidae Wheeler fam. nov. is proposed for the previously unplaced New World genera Paraleucopis Malloch, Mallochianamyia Santos-Neto and Schizostomyia Malloch and undescribed Australian species. A key to genera of Paraleucopidae is provided. Paraleucopis is revised and includes nine species: P. auripes Wheeler & Sinclair sp. nov. (type locality: Andalgala, Argentina); P. bispinosa Wheeler & Sinclair sp. nov. (type locality: Socos, Coquimbo, Chile); P. boharti Wheeler & Sinclair sp. nov. (type locality: Andalgala, Argentina); P. boydensis Steyskal (type locality: nr. Palm Desert, California, USA); P. corvina Malloch (type species of genus; type locality: New Mexico, USA); P. mexicana Steyskal (type locality: Kino Bay, Mexico); P. nigra Wheeler & Sinclair sp. nov. (type locality: Portal, Arizona, USA); P. paraboydensis Wheeler & Sinclair sp. nov. (type locality: Willis Palms Oasis, California, USA); P. saguaro Wheeler & Sinclair sp. nov. (type locality: Usery Mtn Park, Arizona, USA). A key to the species of Paraleucopis is provided. The distribution of Paraleucopis is disjunct, with six species in the western United States and northwestern Mexico and three species in northern Chile and northern Argentina.                The sister group and superfamilial assignment of the Paraleucopidae cannot be established based on current knowledge although the family has affinities to some families of the Asteioinea sensu J.F. McAlpine. A well-supported hypothesis on the relationships of the families of the Acalyptratae will be required before the sister group relationships of Paraleucopidae can be determined. 

Taxonomic review of two fossil crocodylians from the Cenozoic of South America and its implications for the crocodylian fauna of the continent

Zootaxa ◽  
2019 ◽  
Vol 4656 (3) ◽  
pp. 475-486
Author(s):  
GIOVANNE M. CIDADE ◽  
DANIEL FORTIER ◽  
ASCANIO DANIEL RINCÓN ◽  
ANNIE SCHMALTZ HSIOU
Keyword(s):  
South America ◽  
Fossil Record ◽  
Middle Miocene ◽  
Taxonomic Status ◽  
Junior Synonym ◽  
Diverse Group ◽  
Taxonomic Review ◽  
Nomen Dubium ◽  
Extant Species ◽  
The World

The crocodylomorph fauna of the Cenozoic of South America is one of the richest and most diverse in the world. The most diverse group within that fauna is Alligatoroidea, with nearly all of its species belonging to the Caimaninae clade. Many of the fossil alligatoroid species from the Cenozoic of South America were proposed based on very incomplete remains, and as a result their validity requires revision. Two such species are Balanerodus logimus Langston, 1965, from the middle Miocene of Colombia and Peru, and Caiman venezuelensis Fortier & Rincón, 2012, from the Pliocene-Pleistocene of Venezuela. This study has performed a thorough review of the taxonomic status of these two alligatoroid species, concluding that B. logimus is a nomen dubium and that Ca. venezuelensis is a junior synonym of the extant species Ca. crocodilus. This review offers a significantly more accurate scenario for alligatoroid diversity in the Cenozoic of South America in different epochs such as the Miocene and Pleistocene. Additionally, the record of Ca. crocodilus from the Pleistocene of Venezuela is the first fossil record that can be assigned to this species. 

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