scholarly journals Winged fruits of rutaceous affinity from the Eocene of western North America

2020 ◽  
Vol 76 (2) ◽  
pp. 211-216
Author(s):  
Steven R. Manchester ◽  
Kory A. Disney ◽  
Kasey K. Pham

A new kind of fin-winged fruit is recognized from lacustrine shales of the early Eocene Tepee Trail Formation of northwestern Wyoming and from the middle Eocene Clarno Formation of central Oregon, USA. The fruits are obovate with five thick lateral wings, borne on a thick pedicel and bearing scars of hypogynous perianth and disk. The fruit surface is covered with small circular dots interpreted as glands. This combination of characters leads us to infer affinities with the Rutaceae, although no identical modern genus is known. We establish the new genus and species, Quinquala obovata.

2012 ◽  
Vol 86 (6) ◽  
pp. 973-978 ◽  
Author(s):  
Gregg F. Gunnell

Uintasoricines are diminutive plesiadapiforms that are found in the latest Paleocene through middle Eocene, predominantly in North America. They are not a diverse group but individual species may be locally abundant and they are a persistent element of the plesiadapiform radiation in North America surviving over a span of approximately 16 million years. Recent field work in southern Wyoming at South Pass has led to the discovery of a new genus and species of uintasoricine. The new form is smaller in tooth dimensions compared to other known uintasoricines, being slightly smaller thanUintasorex montezumicusfrom California. Both the newly described taxon andU. montezumicusare among the smallest plesiadapiforms yet known with body weights estimated to be 20 to 25 g. The sediments of the Cathedral Bluffs Tongue of the Wasatch Formation at South Pass contain a unique upland fauna—the presence of a distinctive uintasoricine in this assemblage adds further evidence to support the notion that this upland environment was a biodiversity hotspot during the latest early Eocene.


2019 ◽  
Vol 151 (6) ◽  
pp. 783-816
Author(s):  
S. Bruce Archibald ◽  
Robert A. Cannings

AbstractWe describe the first dragonflies (Odonata: Anisoptera) from the early Eocene Okanagan Highlands of far-western North America from nine fossils. Six are assigned to five species in four new, named genera of Aeshnidae: Antiquiala snyderaenew genus and species, Idemlinea versatilisnew genus and species, Ypshna brownleeinew genus and species, Ypshna latipennatanew genus and species, and Eoshna thompsonensisnew genus and species; we treat one as Aeshnidae genus A, species A; one is assigned to Gomphidae: Auroradraco eosnew genus and species; and we treat a ninth, fragmentary fossil of unknown family affinity as Anisoptera indeterminate genus A, species A, which represents a seventh genus and eighth species. The dominance of Aeshnidae is consistent with other Paleocene and Eocene fossil localities. Auroradraco eos is the only fossil Gomphidae in the roughly 66-million-year gap between occurrences in mid-Cretaceous Burmese amber and the early Oligocene of France. Ypshna appears close to Parabaissaeshna ejerslevense from the early Eocene Fur Formation of Denmark; this is not surprising given Holarctic intercontinental connections at this time and a growing list of insect taxa shared between the Okanagan Highlands and the Fur Formation.


Zootaxa ◽  
2021 ◽  
Vol 4966 (3) ◽  
pp. 392-400
Author(s):  
S. BRUCE ARCHIBALD ◽  
ROBERT A. CANNINGS

We describe Republica weatbrooki, a new genus and species of damselfly (Odonata, Zygoptera, Euphaeidae, Eodichromatinae) from the early Eocene (Ypresian) fossil locality at Republic, Washington, U.S.A. Its single specimen is the sole damselfly known from the Okanagan Highlands series of localities in far-western North America. 


Zootaxa ◽  
2009 ◽  
Vol 2063 (1) ◽  
pp. 55-63 ◽  
Author(s):  
VLADIMIR N. MAKARKIN ◽  
S. BRUCE ARCHIBALD

A new genus and species Allorapisma chuorum gen. sp. nov. is described from the Early Eocene locality at Republic, Washington, U.S.A. The forewing venation of Allorapisma is most similar to that of the genus Principiala Makarkin & Menon from the Early Cretaceous of Brazil and Britain. A new, informal suprageneric taxon consisting of these genera is proposed, the Principiala group. The habitats of extant and fossil Ithonidae are briefly discussed.


Zootaxa ◽  
2021 ◽  
Vol 4934 (1) ◽  
pp. 1-133
Author(s):  
S. BRUCE ARCHIBALD ◽  
ROBERT A. CANNINGS ◽  
ROBERT J. ERICKSON ◽  
SETH M. BYBEE ◽  
ROLF W. MATHEWES

We describe the Cephalozygoptera, a new, extinct suborder of Odonata, composed of the families Dysagrionidae and Sieblosiidae, previously assigned to the Zygoptera, and possibly the Whetwhetaksidae n. fam. The Cephalozygoptera is close to the Zygoptera, but differs most notably by distinctive head morphology. It includes 59 to 64 species in at least 19 genera and one genus-level parataxon. One species is known from the Early Cretaceous (Congqingia rhora Zhang), possibly three from the Paleocene, and the rest from the early Eocene through late Miocene. We describe new taxa from the Ypresian Okanagan Highlands of British Columbia, Canada and Washington, United States of America: 16 new species of Dysagrionidae of the existing genus Dysagrion (D. pruettae); the new genera Okanagrion (O. threadgillae, O. hobani, O. beardi, O. lochmum, O. angustum, O. dorrellae, O. liquetoalatum, O. worleyae, all new species); Okanopteryx (O. jeppesenorum, O. fraseri, O. macabeensis, all new species); Stenodiafanus (S. westersidei, new species); the new genus-level parataxon Dysagrionites (D. delinei new species, D. sp. A, D. sp. B, both new); and one new genus and species of the new family Whetwhetaksidae (Whetwhetaksa millerae). 


2019 ◽  
Vol 151 (04) ◽  
pp. 521-530 ◽  
Author(s):  
Vladimir N. Makarkin ◽  
S. Bruce Archibald ◽  
James E. Jepson

AbstractOne new genus of Inocelliidae (Raphidioptera) with one new species and one undetermined specimen is described from the Eocene of North America: Paraksenocellia borealis new genus, new species from the early Eocene (Ypresian) Okanagan Highlands shale at Driftwood Canyon, British Columbia, Canada (a forewing), and Paraksenocellia species from the middle Eocene (Lutetian) of the Coal Creek Member of the Kishenehn Formation, northwestern Montana, United States of America (a hind wing). These are the oldest records of the family. The new genus possesses many character states that are rare in Inocelliidae, e.g., a very long pterostigma extending to ScP in both the forewings and hind wings; the forewing subcostal space has three crossveins; the forewing and hind wing AA1 are deeply forked; the crossvein between CuA and CuP is located far distad the crossvein 1r-m. Paraksenocellia is confidently a member of the Inocelliidae, as it possesses a proximal shift of the basal crossvein 1r-m (connecting R and M) in the forewing and the loss of the basal crossvein 1r-m in the hind wing, both apomorphies of the family. It shares some character states with the Mesozoic Mesoraphidiidae, which we consider to be mostly stem-group plesiomorphies.


1986 ◽  
Vol 60 (3) ◽  
pp. 758-762 ◽  
Author(s):  
Hans-Dieter Sues

Dinnebitodon amarali, a new genus and species of Tritylodontidae, is described from the Early Jurassic Kayenta Formation of northeastern Arizona. It is distinguished from Kayentatherium wellesi by the possession of three (rather than one) upper incisors and upper postcanine teeth with two (rather than three) lingual cusps.


1991 ◽  
Vol 123 (5) ◽  
pp. 1135-1147 ◽  
Author(s):  
Valerie M. Behan-Pelletier ◽  
Nickolai A. Ryabinin

AbstractA new genus and species, Sacculozetes filosus, is described from riparian grassland in the Soviet Far East, and a new species, Guatemalozetes danos, is described from shortgrass and alpine prairie in western North America, both on the basis of adults. The genus Guatemalozetes Mahunka is redefined; this is the first record of this genus from temperate North America. A close relationship is indicated between Sacculozetes and Lamellobates Hammer, Paralamellobates Bhaduri and Raychaudhuri, and Hypozetes Balogh. The uncertain family placement of Sacculozetes and Guatemalozetes within the Ceratozetoidea is discussed. Adults of both genera exhibit character states of the Mycobatidae, but until immatures are discovered, they are considered unplaced genera in the superfamily.


2016 ◽  
Vol 56 (2) ◽  
pp. 247-305 ◽  
Author(s):  
Friðgeir Grímsson ◽  
Guido W. Grimm ◽  
Reinhard Zetter ◽  
Thomas Denk

Abstract Modern lineages of the beech family, Fagaceae, one of the most important north-temperate families of woody flowering plants, have been traced back to the early Eocene. In contrast, molecular differentiation patterns indicate that the Fagus lineage, Fagoideae, with a single modern genus, evolved much earlier than the remaining lineages within Fagaceae (Trigonobalanoideae, Castaneoideae, Quercoideae). The minimum age for this primary split in the Fagaceae has been estimated as 80 ± 20 Ma (i.e. Late Cretaceous) in recently published, time-calibrated phylogenetic trees including all Fagales. Here, we report fagaceous fossils from the Campanian of Wyoming (82-81 Ma; Eagle Formation [Fm]), the Danian of western Greenland (64-62 Ma; Agatdal Fm), and the middle Eocene of British Columbia (ca 48 Ma; Princeton Chert), and compare them to the Fagaceae diversity of the recently studied middle Eocene Hareøen Fm of western Greenland (42-40 Ma). The studied assemblages confirm that the Fagus lineage (= Fagoideae) and the remainder of modern Fagaceae were diverged by the middle Late Cretaceous, together with the extinct Fagaceae lineage(s) of Eotrigonobalanus and the newly recognised genus Paraquercus, a unique pollen morph with similarities to both Eotrigonobalanus and Quercus. The new records push back the origin of (modern) Fagus by 10 Ma and that of the earliest Fagoideae by 30 Ma. The earliest Fagoideae pollen from the Campanian of North America differs from its single modern genus Fagus by its markedly thicker pollen wall, a feature also seen in fossil and extant Castaneoideae. This suggests that a thick type 1 foot layer is also the plesiomorphic feature in Fagoideae although not seen in any of its living representatives. The Danian Fagus pollen of Greenland differs in size from those of modern species but is highly similar to that of the western North American early Eocene F. langevinii, the oldest known beech so far. Together with the Quercus pollen record, absent in the Campanian and Danian formations but represented by several types by the middle Eocene, this confirms recent dating estimates focussing on the genera Fagus and Quercus, while rejecting estimates from all-Fagales-dated trees as too young. The basic Castaneoideae pollen type, still found in species of all five extant genera of this putatively paraphyletic subfamily, represents the ancestral pollen type of most (modern) Fagaceae (Trigonobalanoideae, Castaneoideae, Quercoideae).


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