scholarly journals Species Mixing Effects on Forest Productivity: A Case Study at Stand-, Species- and Tree-Level in the Netherlands

Forests ◽  
2018 ◽  
Vol 9 (11) ◽  
pp. 713 ◽  
Author(s):  
Huicui Lu ◽  
Godefridus Mohren ◽  
Miren del Río ◽  
Mart-Jan Schelhaas ◽  
Meike Bouwman ◽  
...  
Keyword(s):  
Forest Management ◽  
Soil Fertility ◽  
Basal Area ◽  
Tree Level ◽  
Stand Age ◽  
Forest Plot ◽  
Mixed Species ◽  
Mixed Stands ◽  
Individual Tree ◽  
Species Mixtures

Many monoculture forests have been converted to mixed-species forests in Europe over the last decades. The main reasons for this conversion were probably to increase productivity, including timber production, and enhance other ecosystem services, such as conservation of biodiversity and other nature values. This study was done by synthesizing results from studies carried out in Dutch mixed forests compared with monoculture stands and evaluating them in the perspective of the current theory. Then we explored possible mechanisms of higher productivity in mixed stands, in relation to the combination of species, stand age and soil fertility, and discussed possible consequences of forest management. The study covered five two-species mixtures and their corresponding monoculture stands from using long-term permanent forest plots over multiple decades as well as two inventories (around 2003 and 2013) across the entire Netherlands. These forest plot data were used together with empirical models at total stand level, species level and tree level. Overyielding in Douglas-fir–beech and pine–oak mixtures was maintained over time, probably owing to the intensive thinning and was achieved on the poorer soils. However, this overyielding was not always driven by fast-growing light-demanding species. On individual tree level, intra-specific competition was not necessarily stronger than inter-specific competition and this competitive reduction was less seen at lower soil fertility and dependent on species mixtures. Moreover, size-asymmetric competition for light was more associated with tree basal area growth than size-symmetric competition for soil resources. Overall, this study suggests a substantial potential of species mixing for increasing productivity and implies developing forest management strategies to convert monospecific forests to mixed-species forests that consider the complementarity in resource acquisition of tree species.

scholarly journals Species Mixing Regulation with Respect to Forest Ecosystem Service Provision

Forests ◽  
2018 ◽  
Vol 9 (10) ◽  
pp. 632 ◽  
Author(s):  
Fabian Schwaiger ◽  
Werner Poschenrieder ◽  
Peter Biber ◽  
Hans Pretzsch
Keyword(s):  
Forest Management ◽  
Groundwater Recharge ◽  
Ecosystem Service ◽  
Service Provision ◽  
Quantitative Methods ◽  
Basal Area ◽  
Stand Density ◽  
European Beech ◽  
Stand Age ◽  
Mixed Stands

The control and maintenance of species composition of mixed stands is a highly relevant objective of forest management in order to provide multifunctionality and climatic resilience. In contrast to this requirement there is, however, an evident lack of quantitative methods for mixture regulation. In this context, we propose an approach for the regulation of mixture proportions that has been implemented in a forest management model. The approach considers species-specific growth characteristics and takes into account the mixing effect on stand density. We present five exemplary simulations that apply the regulation. Each simulation maintains one of five desired species compositions. In these simulations, we consider the species European beech and Norway spruce under good site conditions, thus representing the most prominent mixed stands in Central Europe. Based on this model experiment, we analyze the potential benefit of controlled mixing regulation for achieving desired levels and combinations of ecosystem service provision, in particular productivity, diversity, and groundwater recharge. We found that a constant 50% basal area share of beech (equivalent growing space share of 80% to 70% depending on stand age) provided the most balanced supply of ecosystem services. Prominently, groundwater recharge considerably decreased when beech basal area shares were held below 50%. We discuss the ecological and practical implications of the regulation approach and different mixing shares.

Using tree rings to assess the potential of thinning to alleviate drought stress in a dry forest of western Canada

2021 ◽  
Author(s):  
David Montwé ◽  
Audrey Standish ◽  
Miriam Isaac-Renton ◽  
Jodi Axelson
Keyword(s):  
Tree Ring ◽  
Tree Rings ◽  
Basal Area ◽  
Dry Forest ◽  
Tree Level ◽  
Severe Drought ◽  
Western Canada ◽  
Drought Event ◽  
Individual Tree ◽  
Potential Benefits

<p>Increasing frequency of severe drought events under climate change is a major cause for concern for millions of hectares of forested land. One practical solution to improving forest resilience may be thinning. There may be several potential benefits, chief of which is that drought tolerance could be improved in the remaining trees due to lower competition for resources and increased precipitation throughfall. By improving resilience to drought, this may increase productivity of the remaining trees while lowering risks of mortality. Such potential benefits can effectively be quantified with data from statistically-sound, long-term field experiments, and tree rings provide a suitable avenue to compare treatments. We work with an experiment that applied different levels of tree retention to mature interior Douglas fir (<em>Pseudotsuga menziesii</em> var. <em>glauca</em>) in a dry ecosystem of western Canada. The treatments were applied in the winter of 2002/2003, coinciding with the aftermath of a severe natural drought event in 2002. We used tree-rings to quantify the extent to which thinning improves recovery and resilience of treated trees as compared to non-thinned controls. Tree-ring samples as well as height and diameter data were obtained from 83 trees from 8 treatment units of the randomized experimental design. Indicators for resilience to drought were calculated based on basal area increments. Thinning substantially increased basal area increments at the individual tree level, but more importantly, led to significantly higher recovery and resilience relative to the control. The results of this tree-ring analysis suggest that thinning may be a viable silvicultural intervention to counteract effects of severe drought events and to maintain tree cover.</p>

Individual-tree basal area growth models for jack pine and black spruce in northern Ontario

2004 ◽  
Vol 80 (3) ◽  
pp. 366-374 ◽  
Author(s):  
Lianjun Zhang ◽  
Changhui Peng ◽  
Qinglai Dang
Keyword(s):  
Tree Growth ◽  
Black Spruce ◽  
Basal Area ◽  
Jack Pine ◽  
Mixed Stands ◽  
Northern Ontario ◽  
Individual Tree ◽  
Basal Area Growth ◽  
Wide Range ◽  
Area Growth

Individual-tree models of five-year basal area growth were developed for jack pine (Pinus banksiana Lamb.) and black spruce (Picea mariana (Mill.) BSP) in northern Ontario. Tree growth data were collected from long-term permanent plots of pure and mixed stands of the two species. The models were fitted using mixed model methods due to correlated remeasurements of tree growth over time. Since the data covered a wide range of stand ages, stand conditions and tree sizes, serious heterogeneous variances existed in the data. Therefore, the coefficients of the final models were obtained using weighted regression techniques. The models for the two species were evaluated across 4-cm diameter classes using independent data. The results indicated (1) the models of jack pine and black spruce produced similar prediction errors and biases for intermediate-sized trees (12–28 cm in tree diameter), (2) both models yielded relatively large errors and biases for larger trees (> 28 cm) than those for smaller trees, and (3) the jack pine model produced much larger errors and biases for small-sized trees (< 12 cm) than did the black spruce model. Key words: mixed models, repeated measures, model validation

scholarly journals Estimating the Height and Basal Area at Individual Tree and Plot Levels in Canadian Subarctic Lichen Woodlands Using Stereo WorldView-3 Images

2019 ◽  
Vol 11 (3) ◽  
pp. 248 ◽  
Author(s):  
Benoît St-Onge ◽  
Simon Grandin
Keyword(s):  
Remote Sensing ◽  
Basal Area ◽  
Ground Cover ◽  
Structural Data ◽  
Tree Level ◽  
Individual Tree ◽  
Visual Contrast ◽  
The Relationship ◽  
Height Data

Lichen woodlands (LW) are sparse forests that cover extensive areas in remote subarctic regions where warming due to climate change is fastest. They are difficult to study in situ or with airborne remote sensing due to their remoteness. We have tested a method for measuring individual tree heights and predicting basal area at tree and plot levels using WorldView-3 stereo images. Manual stereo measurements of tree heights were performed on short trees (2–12 m) of a LW region of Canada with a residual standard error of ≈0.9 m compared to accurate field or UAV height data. The number of detected trees significantly underestimated field counts, especially in peatlands in which the visual contrast between trees and ground cover was low. The heights measured from the WorldView-3 images were used to predict the basal area at individual tree level and summed up at plot level. In the best conditions (high contrast between trees and ground cover), the relationship to field basal area had a R2 of 0.79. Accurate estimates of above ground biomass should therefore also be possible. This method could be used to calibrate an extensive remote sensing approach without in-situ measurements, e.g., by linking precise structural data to ICESAT-2 footprints.

scholarly journals Over- and Underyielding in Time and Space in Experiments with Mixed Stands of Scots Pine and Norway Spruce

Forests ◽  
2018 ◽  
Vol 9 (8) ◽  
pp. 495 ◽  
Author(s):  
Lars Drössler ◽  
Eric Agestam ◽  
Kamil Bielak ◽  
Małgorzata Dudzinska ◽  
Julia Koricheva ◽  
...  
Keyword(s):  
Norway Spruce ◽  
Scots Pine ◽  
Tree Species ◽  
Mixed Forest ◽  
Stand Age ◽  
Mixed Species ◽  
Mixed Stands ◽  
Mixing Effect ◽  
Individual Site ◽  
Study Sites

Pine-spruce forests are one of the commonest mixed forest types in Europe and both tree species are very important for wood supply. This study summarized nine European studies with Scots pine and Norway spruce where a mixed-species stand and both monocultures were located in an experimental set-up. Overyielding (where growth of a mixed stand was greater than the average of both monocultures) was relatively common and often ranged between 0% and 30%, but could also be negative at individual study sites. Each individual site demonstrated consistent patterns of the mixing effect over different measurement periods. Transgressive overyielding (where the mixed-species stand was more productive than either of the monocultures) was found at three study sites, while a monoculture was more productive on the other sites. Large variation between study sites indicated that the existing experiments do not fully represent the extensive region where this mixed pine-spruce forest can occur. Pooled increment data displayed a negative influence of latitude and stand age on the mixing effect of those tree species in forests younger than 70 years.

An individual-tree basal area growth model for black spruce in second-growth peatland stands

1999 ◽  
Vol 29 (5) ◽  
pp. 621-629 ◽  
Author(s):  
Hannu Hökkä ◽  
Arthur Groot
Keyword(s):  
Growth Model ◽  
Black Spruce ◽  
Basal Area ◽  
Random Parameter ◽  
Considerable Change ◽  
Tree Level ◽  
Individual Tree ◽  
Basal Area Growth ◽  
Second Growth ◽  
Area Growth

A basal area growth model was developed to predict the growth of individual trees in second-growth black spruce (Picea mariana (Mill.) BSP) stands on northeastern Ontario peatlands. The data were derived from stem analysis trees collected in 1985 and 1986 from stands harvested 47-68 years earlier. For a period starting from the date of data collection and going back to 10 years from the harvesting, tree basal area growth, diameters, and stand characteristics were retrospectively calculated at 5-year intervals. To estimate previous mortality, self-thinning relationships for black spruce were applied. In the model, 5-year basal area growth of a tree was expressed as a function of tree diameter, stand-level competition, tree-level competition, and peat thickness. There was considerable change in the growth-size relationship over time. A random parameter approach was applied in model construction to account for the spatial and temporal correlations of the observations. The proposed model explicitly incorporates factors normally included in a "random error" term and, therefore, should provide more sensitive tests of the contributions of the various factors to growth prediction. The estimated model showed only slight bias against the modeling data and the predicted stand basal area development was comparable with that given in other studies.

Modelling survival probability of individual trees in Norway spruce stands under different thinning regimes

2005 ◽  
Vol 35 (1) ◽  
pp. 113-121 ◽  
Author(s):  
Kjell Karlsson ◽  
Lennart Norell
Keyword(s):  
Norway Spruce ◽  
Goodness Of Fit ◽  
Basal Area ◽  
Site Index ◽  
Stand Age ◽  
Goodness Of Fit Test ◽  
Individual Tree ◽  
Thinning Intensity ◽  
Diameter At Breast Height ◽  
Breast Height

The probability that an individual tree will remain in even-aged Norway spruce (Picea abies (L.) Karst.) stands subjected to different thinning programmes was modelled, using data from a thinning experiment established in 25 localities in southern Sweden. A logistic regression approach was used to predict the probability and the Hosmer–Lemeshow goodness-of-fit test to evaluate the fit. Diameter at breast height (DBH), quadratic mean DBH, thinning intensity, thinning quotient, basal area, number of stems per hectare, stand age, number of thinnings, and site index were used as explanatory variables. Separate analyses for stands thinned from below, stands thinned from above, and unthinned stands were performed. The modelled probability graphs for trees not being removed, plotted against their diameter at breast height, had clear S-shapes for both unthinned stands and stands thinned from below. The graph for stands thinned from above was bell-shaped.

A new method for evaluating forest thinning: growth dominance in managed Pinus resinosa stands

2010 ◽  
Vol 40 (5) ◽  
pp. 843-849 ◽  
Author(s):  
John B. Bradford ◽  
Anthony W. D’Amato ◽  
Brian J. Palik ◽  
Shawn Fraver
Keyword(s):  
Tree Growth ◽  
Basal Area ◽  
Pinus Resinosa ◽  
Growth Patterns ◽  
Red Pine ◽  
New Method ◽  
Stand Age ◽  
Individual Tree ◽  
Individual Tree Growth

Growth dominance is a relatively new, simple, quantitative metric of within-stand individual tree growth patterns, and is defined as positive when larger trees in the stand display proportionally greater growth than smaller trees, and negative when smaller trees display proportionally greater growth than larger trees. We examined long-term silvicultural experiments in red pine ( Pinus resinosa Ait.) to characterize how stand age, thinning treatments (thinned from above, below, or both), and stocking levels (residual basal area) influence stand-level growth dominance through time. In stands thinned from below or from both above and below, growth dominance was not significantly different from zero at any age or stocking level. Growth dominance in stands thinned from above trended from negative at low stocking levels to positive at high stocking levels and was positive in young stands. Growth dominance in unthinned stands was positive and increased with age. These results suggest that growth dominance provides a useful tool for assessing the efficacy of thinning treatments designed to reduce competition between trees and promote high levels of productivity across a population, particularly among crop trees.

scholarly journals Spatial-Temporal Patterns of Spruce Budworm Defoliation within Plots in Québec

Forests ◽  
2019 ◽  
Vol 10 (3) ◽  
pp. 232 ◽  
Author(s):  
Mingke Li ◽  
David MacLean ◽  
Chris Hennigar ◽  
Jae Ogilvie
Keyword(s):  
Spatial Autocorrelation ◽  
Hot Spot ◽  
Basal Area ◽  
Temporal Patterns ◽  
Spruce Budworm ◽  
Balsam Fir ◽  
Tree Level ◽  
Cold Spot ◽  
Individual Tree ◽  
Tree Defoliation

We investigated the spatial-temporal patterns of spruce budworm (Choristoneura fumiferana (Clem.); SBW) defoliation within 57 plots over 5 years during the current SBW outbreak in Québec. Although spatial-temporal variability of SBW defoliation has been studied at several scales, the spatial dependence between individual defoliated trees within a plot has not been quantified, and effects of defoliation level of neighboring trees have not been addressed. We used spatial autocorrelation analyses to determine patterns of defoliation of trees (clustered, dispersed, or random) for plots and for individual trees. From 28% to 47% of plots had significantly clustered defoliation during the 5 years. Plots with clustered defoliation generally had higher mean defoliation per plot and higher deviation of defoliation. At the individual-tree-level, we determined ‘hot spot trees’ (highly defoliated trees surrounded by other highly defoliated trees) and ‘cold spot trees’ (lightly defoliated trees surrounded by other lightly defoliated trees) within each plot using local Getis-Ord Gi* analysis. Results revealed that 11 to 27 plots had hot spot trees and 27% to 64% of them had mean defoliation <25%, while plots with 75% to 100% defoliation had either cold spot trees or non-significant spots, which suggested that whether defoliation was high or low enough to be a hot or cold spot depended on the defoliation level of the entire plot. We fitted individual-tree balsam fir defoliation regression models as a function of plot and surrounding tree characteristics (using search radii of 3–5 m). The best model contained plot average balsam fir defoliation and subject tree basal area, and these two variables explained 80% of the variance, which was 2% to 5% higher than the variability explained by the neighboring tree defoliation, over the 3–5 m search radii tested. We concluded that plot-level defoliation and basal area were adequate for modeling individual tree defoliation, and although clustering of defoliation was evident, larger plots were needed to determine the optimum neighborhood radius for predicting defoliation on an individual. Spatial autocorrelation analysis can serve as an objective way to quantify such ecological patterns.

scholarly journals Generalized Nonlinear Mixed-Effects Individual Tree Diameter Increment Models for Beech Forests in Slovakia

Forests ◽  
2019 ◽  
Vol 10 (5) ◽  
pp. 451 ◽  
Author(s):  
Ram P. Sharma ◽  
Igor Štefančík ◽  
Zdeněk Vacek ◽  
Stanislav Vacek
Keyword(s):  
Basal Area ◽  
European Beech ◽  
Site Index ◽  
Development Stage ◽  
Mixed Effects ◽  
Site Quality ◽  
Stand Age ◽  
Sample Plot ◽  
Individual Tree ◽  
Diameter Increment

Individual tree growth and yield models precisely describe tree growth irrespective of stand complexity and are capable of simulating various silvicultural alternatives in the stands with diverse structure, species composition, and management history. We developed both age dependent and age independent diameter increment models using long-term research sample plot data collected from both monospecific and mixed stands of European beech (Fagus sylvatica L.) in the Slovak Republic. We used diameter at breast height (DBH) as a main predictor and other characteristics describing site quality (site index), stand development stage (dominant height and stand age), stand density or competition (ratio of individual tree DBH to quadratic mean diameter), species mixture (basal area proportion of a species of interest), and dummy variable describing stand management regimes as covariate predictors to develop the models. We evaluated eight versatile growth functions in the first stage using DBH as a single predictor and selected the most suitable one, i.e., Chapman-Richards function for further analysis through the inclusion of covariate predictors. We introduced the random components describing sample plot-level random effects and stochastic variations on the diameter increment, into the models through the mixed-effects modelling. The autocorrelation caused by hierarchical data-structure, which is assumed to be partially reduced by mixed-effects modelling, was removed through the inclusion of the parameter accounting for the autoregressive error-structures. The models described about two-third parts of a total variation in the diameter increment without significant trends in the residuals. Compared to the age independent mixed-effects model (conditional coefficient of determination, R c 2 = 0.6566; root mean square error, RMSE = 0.1196), the age dependent model described a significantly larger proportion of the variations in diameter increment ( R c 2 = 0.6796, RMSE = 0.1141). Diameter increment was significantly influenced differently by covariate predictors included into the models. Diameter increment decreased with the advancement of stand development stage (increased dominant height and stand age), increasing intraspecific competition (increased basal area proportion of European beech per sample plot), and diameter increment increased with increasing site quality (increased site index) and decreased competition (increased ratio of DBH to quadratic mean diameter). Our mixed-effects models, which can be easily localized with the random effects estimated from prior measurement of diameter increments of four randomly selected trees per sample plot, will provide high prediction accuracies. Our models may be used for simulating growth of European beech irrespective of its stand structural complexity, as these models have included various covariate variables describing both tree-and stand-level characteristics, thinning regimes, except the climate characteristics. Together with other forest models, our models will be used as inputs to the growth simulator to be developed in the future, which is important for decision-making in forestry.

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