scholarly journals Radial Variation of Wood Anatomical Properties Determines the Demarcation of Juvenile-Mature Wood in Schima superba

Forests ◽  
2021 ◽  
Vol 12 (4) ◽  
pp. 512
Author(s):  
Yunpeng Wang ◽  
Rui Zhang ◽  
Zhichun Zhou

Schima superba is a precious broadleaf tree species that produces excellent timber. Analyzing the radial variation of wood anatomical properties within stems and determining the boundary between the juvenile and mature wood can provide a theoretical basis for the breeding and cultivation of S. superba. Herein, 30 54-year-old trees were used to analyze the radial variation of wood anatomical properties qualitatively. The correlation among wood properties was analyzed. On this basis, four types of model were used to predict the wood properties quantitatively, and the optimal segmentation model was used to determine the boundary between juvenile and mature wood. The results showed that the variation of the early stage (before the 20th year) was larger than that of the later stage (after the 20th year); moreover, the variation of the vessel features (9.56%) was greater than that of the fiber features (7.42%), the vessel lumen diameter (11.94%) and the fiber length (8.00%) had the most variation. There were three radial variation patterns in the wood properties. In the early stage, there was a significant positive correlation between the ring width and wood air-dry density (0.78) and wood basic density (0.89), and a significant negative correlation between the ring width and the fiber characters (−0.79~−0.93) and vessel characters (−0.64~−0.90). The growth models of Logistic and Richards were more suitable (R2 = 84.2%~95.2%) for the radial variation of wood anatomical properties than the nonlinear least square model (R2 = 67.7%~90.9%). The transition wood between juvenile and mature wood of S. superba was from the 11th to the 16th year based on aggregative indicators from the pith to bark at breast height. Finally, when taken together, the wood anatomical properties of S. superba had regular radial variation. Breeding programs need to consider both growth and wood anatomical properties in the early stage, but they can be neglected in the mature stage. The formation of mature wood of S. superba occurred after the 16th year.

IAWA Journal ◽  
2014 ◽  
Vol 35 (4) ◽  
pp. 363-384 ◽  
Author(s):  
Besma Bouslimi ◽  
Ahmed Koubaa ◽  
Yves Bergeron

The variability in wood properties of eastern white cedar (Thuja occidentalis L.) is relatively poorly known. Here we report the axial and the radial variation in selected anatomical properties, namely, ring width, wood density, and tracheid length and width. Forty-five trees were randomly sampled and felled from three selected sites in the Abitibi-Témiscamingue region, Quebec, Canada. Disks were systematically sampled at 0.5, 1.3, and 3 m stem height and at every 2 meters thereafter up to the tree top. Average ring density at breast height was 355 kg/m3 with a small difference between earlywood and latewood. The latewood proportion was uniform and constant within the tree at about 32%. The tracheids were fine and long, averaging 25.3 μm in width and 2.07 mm in length. The variation in wood density components between trees was highly significant. The cambial age effect on all measured properties was highly significant. Ring density decreased from a maximum near the pith to a minimum in the juvenile–mature wood transition zone and remained constant or decreased slightly thereafter. Annual ring width decreased from a maximum near the pith to a minimum at the 10th ring and increased thereafter. Tracheid length and width showed typical radial variation characterized by a steady increase from pith to bark. Within-tree axial variation was highly significant, but ring width showed more substantial changes. Changes in wood properties with height depend on cambial age and thus are implied since the proportion of juvenile wood in the stem increases from the base to the top.


Forests ◽  
2019 ◽  
Vol 10 (7) ◽  
pp. 589 ◽  
Author(s):  
Cyriac S. Mvolo ◽  
Ahmed Koubaa ◽  
Jean Beaulieu ◽  
Alain Cloutier ◽  
Maurice Defo ◽  
...  

We examined phenotypic relationships among radial growth-related, physical (i.e., related to wood density), and anatomical (i.e., related to tracheid dimensions) wood properties in white spruce (Picea glauca (Moench) Voss), in order to determine the strength and significance of their correlations. Additionally, principal component analysis (PCA) was used to establish if all of the properties must be measured and to determine the key properties that can be used as proxies for the other variables. Radial growth-related and physical properties were measured with an X-ray densitometer, while anatomical properties were measured with a Fiber Quality Analyzer. Fifteen wood properties (tracheid length (TL) and diameter (TD), earlywood tracheid length (ETL) and diameter (ETD), latewood tracheid length (LTL) and diameter (LTD), ring width (RW), ring area (RA), earlywood width (EWW), latewood width (LWW), latewood proportion (LWP), ring density (RD), intra-ring density variation, earlywood density (EWD), and latewood density (LWD)) were assessed. Relationships were evaluated at intra-ring and inter-ring levels in the juvenile wood (JW) and mature wood (MW) zones. Except for a few cases when mature tracheid diameter (TD) was involved, all intra-ring anatomical properties were highly and significantly correlated. Radial growth properties were correlated, with stronger relationships in MW compared to JW. Physical properties were often positively and significantly correlated in both JW and MW. A higher earlywood density coupled with a lower latewood density favored wood uniformity, i.e., the homogeneity of ring density within a growth ring. Managing plantations to suppress trees growth during JW formation, and enhancing radial growth when MW formation starts will favor overall wood quality. In order, RW-EWW-RA, TL-ETL-LTL, and RD-EWD-LWP are the three clusters that appeared in the three wood zones, the whole pith-to-bark radial section, the juvenile wood zone, and the mature wood zone.


IAWA Journal ◽  
2010 ◽  
Vol 31 (2) ◽  
pp. 169-178 ◽  
Author(s):  
Stergios Adamopoulos ◽  
Costas Passialis ◽  
Elias Voulgaridis

Cross-sectional discs were cut at breast height from 19 black locust trees aged 18–39 years of different clones and origin (Greece, Bulgaria and Hungarian clones J, U and NY). Ring width, latewood proportion and dry density were measured according to standard laboratory techniques in order to examine their relationships. Latewood proportion was found to increase (except for one tree from Bulgaria) with increasing ring width and logarithmic equations were calculated with relatively high correlation coefficients. A positive relation between dry density and latewood proportion was found in black locust of Bulgarian origin and in the Hungarian clone NY while in all other cases no meaningful relationship was observed. In mature wood, improved positive linear correlations were observed between dry density and latewood proportion with the exception of the Hungarian clone U. Regression analysis revealed low correlations between dry density and ring width and only black locust from Bulgaria had a high positive correlation. The positive correlations between dry density and ring width were slightly improved in mature wood with the exception of clone J from Hungary.


2012 ◽  
Vol 58 (No. 4) ◽  
pp. 145-151 ◽  
Author(s):  
A. Zeidler

The aim of this study was to investigate the wood density of Turkish hazel (Corylus colurna L.), an introduced species in the Czech Republic. Sample trees coming from a Czech site were tested for basic wood density and oven-dry density according to Czech national standards. Four sections were taken from each tree to evaluate the variability of wood density along the stem height and along the stem radius. The mean values for the basic density and the oven-dry density were 544 kg&middot;m<sup>&ndash;3 </sup>and 627 kg&middot;m<sup>&ndash;3</sup>, respectively. The results show that the highest density can be found in the bottom part of the tree, followed by a decline with the increasing height of the stem. For the radial direction from the pith to the bark, the pattern of density variability was more complex, reaching its highest value close to the cambium. Practically no correlation between annual ring width and density values was found out. &nbsp;


Forests ◽  
2021 ◽  
Vol 12 (7) ◽  
pp. 868
Author(s):  
Yao Xiao ◽  
Junhui Wang ◽  
Huiling Yun ◽  
Guijuan Yang ◽  
Jianwei Ma ◽  
...  

Catalpa bungei is an important timber tree. Improvements in growth and wood quality are important goals of C. bungei breeding, and it is necessary to understand the genetic parameters of specific target traits and genetic correlation between growth traits and wood properties for tree breeding. In this study, the genetic parameters of height, diameter at breast height (DBH) and wood properties were estimated and genetic and phenotypic correlations between growth traits and wood properties were evaluated in C. bungei. Finally, different selection scenarios were used to evaluate and select optimal clones. The results showed that there were significant differences in growth and wood properties among clones. The wood hardness (0.66–0.79), basic density (0.89), air-dried density (0.89) and compression strength parallel to the grain of wood (CSP) (0.84) had high repeatability. The variance component proportions indicated that the variation in wood properties came mainly from different genotypes (clones) rather than from different individuals of the same clone. The DBH showed a significant negative genetic correlation with the hardness of radial section (HRS) (−643), basic density (−0.531) and air-dry density (−0.495). This unfavorable relationship makes it difficult to improve growth and wood quality simultaneously in C. bungei. We selected the optimal clones under different scenarios, and we obtained 7.75–9.06% genetic gains for growth in the scenario in which height and DBH were the target traits. Genetic gains of 7.43–14.94% were obtained for wood properties by selecting optimal clones in the scenario in which wood properties were the target traits. Approximately 5% and 4% genetic gains were obtained for growth and wood properties, respectively, for the combined selection. This study provides new insights into the genetic improvement of wood quality in C. bungei.


Forests ◽  
2021 ◽  
Vol 12 (5) ◽  
pp. 634
Author(s):  
Szymon Bijak ◽  
Hubert Lachowicz

Black locust (Robinia pseudoacacia L.) is a non-native tree species that occupies a remarkable area in the forests of western Poland. It is mainly cultivated for the bee-keeping purposes as well as for its high quality wood. We investigated the impact of tree age and diameter on the selected structural, physical and mechanical attributes of wood of black locust that grows in conditions of mesic oligotrophic site. We analysed 200 samples originating from 18 trees that varied with age (38, 60 and 71 years old) and diameter (thin, medium and thick specimen selected according to Hartig’s method). Individual wood properties were determined along with corresponding European or Polish standards. Structural and mechanical attributes were determined for moisture of 12%. We found significant impact of tree age on tree-ring width, latewood proportion, density, oven-dry density, basic density, share of woody substance, porosity, as well as radial, tangential, longitudinal and volumetric shrinkage, compression strength parallel to grain, static bending, coefficient of compression strength parallel to grain and coefficient of static bending. The older the trees, the higher values of individual attributes were observed. In turn, the effect of tree diameter was less profound and no significant impact of that feature was found for latewood proportion, anisotropy and almost all of the shrinkage parameters. Thin trees exhibited the lowest values of the analysed parameters, while medium ones—the highest. In general, the highest technical quality of the investigated wood can be found in the youngest trees, whose wood characterises with the properties significantly exceeding native Polish tree species such as oak or beech.


IAWA Journal ◽  
2009 ◽  
Vol 30 (1) ◽  
pp. 49-58 ◽  
Author(s):  
Finto Antony ◽  
Lewis Jordan ◽  
Laurence R. Schimleck ◽  
Richard F. Daniels ◽  
Alexander Clark III

Mid-rotation fertilization is a common practice in the management of loblolly pine (Pinus taeda L.) plantations, typically providing large improvements in growth. However, concerns exist about the quality of wood produced following fertilization. The objective of this study was to develop an understanding of wood property changes following fertilization. Wood samples from a study involving four levels of fertilization applied to a thinned mid-rotation loblolly pine plantation located on the lower coastal plain of North Carolina were sampled. The study was laid out in a randomized complete block design involving four blocks and four levels of nitrogen fertilizer: Control-000, 112, 224 and 336 kg/ha, along with 28 kg/ha of phosphorus with all treatments. Thirty-two trees were felled and disks were cut at five heights from each tree. Wood properties including modulus of elasticity, air-dry density and tracheid anatomical properties were measured for each of the three post-fertilization annual growth rings using near infrared (NIR) spectra obtained from the radial face of strips cut from the disks. An analysis of variance was conducted on three-year basal area weighted average stiffness, air-dry density, and tracheid anatomical properties. A decrease in stiffness, air-dry density, tracheid wall thickness, and an increase in tracheid radial diameter were observed for the heaviest fertilizer treatment (336 kg/ha) compared to the control and 112 kg/ha of nitrogen. Microfibril angle (MFA), cell tangential diameter, and tracheid perimeter showed little change. Wood properties of trees receiving fertilizer rates of 112 and 224 kg/ha were not significantly affected.


2021 ◽  
Vol 67 (1) ◽  
Author(s):  
Togtokhbayar Erdene-Ochir ◽  
Futoshi Ishiguri ◽  
Ikumi Nezu ◽  
Bayasaa Tumenjargal ◽  
Bayartsetseg Baasan ◽  
...  

AbstractWood properties, such as annual ring width, wood fiber length, vessel element length, basic density, air-dry density, dynamic Young’s modulus, modulus of elasticity (MOE), modulus of rupture (MOR), absorbed energy in impact bending, compressive strength parallel to grain, and shearing strength, were investigated for wood from 10 naturally regenerated trees of Betula platyphylla Sukaczev in Mandal, Selenge, Mongolia. Mixed-effects models were used to evaluate the radial variations in the wood properties. The mean values of wood properties obtained in the present study were in almost the same range, with a few exceptions, as those reported by other researchers for other Betula species. The radial variations of wood properties in B. platyphylla were well-fitted to a nonlinear mixed-effects model (logarithmic formula); all examined wood properties increased from the pith and then became constant toward the bark side. The wood properties significantly differed between the core and outer wood. Basic density, air-dry density, and dynamic Young’s modulus were significantly correlated with MOE, MOR, and compressive strength. It is concluded that when the wood of B. platyphylla is utilized as raw materials for solid wood products, the differences between the core wood and outer wood should be considered. In addition, the selection of wood with higher strength properties can be achieved using the wood density and dynamic Young’s modulus as indicators.


2021 ◽  
Vol 52 (3) ◽  
pp. 774-782
Author(s):  
Abdulqader & et al.

Melia azedarach L.  was introduced into Kurdistan region and planted as an ornamental tree at the nurseries, parks and streets. This study investigated the quality of wood of 9 year old M. azedarach trees grown in Duhok province. Samples of wood from five trees were taken at breast height diameter; and their wood quality was studied in terms of morphological, physical and anatomical properties. Results revealed that the values of morphological properties were: the Heartwood percentage (69.01%), Sapwood percentage (12.93%), Bark percentage (8.06%) and Annual Ring Width (7.74mm). While, the values of the physical properties were: specific gravity (0.42), basic density of wood (0.36g/cm3), annual ring growth width (7.74cm), moisture content (44.46%), maximum moisture content (208.36%), volumetric shrinkage (13.18%), volumetric swelling (16.25%), fiber saturation point (36.34%), cell wall (28.05%) and porosity (71.94%). The values of anatomical properties were: fiber length (0.792 mm), fiber diameter at the mid- point of the fiber (17.75 μm), fiber double cell wall thickness (9.67 μm), fiber lumen width (8.07μm), runkel ratio (1.55), flexibility ratio (47.17), slenderness ratio (47.58), vessel length (241.57μm), vessel diameter at the mid- point of the vessel (169.09μm), and vessel lumen diameter (158.26μm). All properties showed a significant variation from pith to bark and between the trees. The results equip a basis for determining management planning opportune to production wood of M. azedarach plantation trees in Kurdistan region. Also M. azedarach is characterized by low strength resistance and perhaps low pulping production too, therefore according to its low wood quality for some properties, is not recommended for structural purposes. While anatomical properties, showed medium values of fiber dimensions which could be used for production of special type of paper


FLORESTA ◽  
2010 ◽  
Vol 40 (3) ◽  
Author(s):  
Israel Luiz de Lima ◽  
Beatriz Ribeiro Zanon ◽  
Eduardo Luiz Longui ◽  
Sandra Monteiro Borges Florsheim

Croton floribundus(capixingui) é encontrado nos estados de Minas Gerais, São Paulo e Paraná. Face à necessidade de novos conhecimentos das propriedades desta madeira, em condições de laboratório, objetivou-se estudar a variação radial da densidade básica e as dimensões celulares desta espécie, oriunda de população natural. De cinco árvores, com DAP médio de 20,35 cm, coletou-se um disco de 7 cm de espessura na região do DAP. Estas árvores estavam localizadas no Parque Estadual da Cantareira, do Instituto Florestal do Estado de São Paulo. De cada disco foram retiradas amostras representando as posições próximo à medula, região intermediária e próximo à casca, para o estudo da variação radial, das seguintes variáveis: densidade básica; comprimento, diâmetro e frequência de vasos; altura, largura e freqüência de raios; comprimento, diâmetro, espessura da parede e lume das fibras. Os resultados indicaram que ocorreu uma tendência de aumento no diâmetro de vasos e uma diminuição na frequência de raios no sentido medula-casca, e para as demais variáveis não ocorreu a mesma tendência.Palavras-chave: Croton floribundus; células da madeira; variação radial. AbstractRadial variation of the basic density and cellular dimensions of Croton floribundus. Croton floribundus(capixingui) is found in the States of Minas Gerais, São Paulo and Paraná. In view of the necessity of new knowledge of the wood properties in laboratory conditions, this work aimed to study the radial variation of the basic density and cellular dimensions of this species. Discs from five trees with average DBH of 20,35 cm, with thickness of 7 cm, were collected at DBH height. The trees came from the Cantareira State Park, property of the Forestry Institute of Sao Paulo State. From each disc, samples were removed, representing the positions pith, intermediate and bark, to study the radial variation of the following variables: basic density; length, diameter, and vessels frequency; length, width, and rays frequency; wall length, diameter, and thickness; and lumen diameter. The results indicated that occurred an increase tendency in the vessels diameter and reduction in the ray frequency from the pith to the bark; the other variables did not show this trend.Keywords:Croton floribundus; wood cells; pith-bark variation. 


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