The proportions of individuals of different kinds in two-type populations. A branching process problem arising in biology

1969 ◽  
Vol 6 (2) ◽  
pp. 249-260 ◽  
Author(s):  
P. Jagers

Consider an age-dependent branching process with two types of individuals. Suppose that individuals of one type beget children of both types, whereas those of the other type can only give birth to individuals of their own kind. This paper is a study of the relation between two random functions occurring in such processes starting from an ancestor of the first type, the two functions being the numbers of individuals of the two kinds. Under weak assumptions it is shown that the random proportion of individuals of one type converges as time passes, in mean square as well as almost surely to a non-random limit, easily determined in terms of the reproduction laws and life-length distributions of the process.

1969 ◽  
Vol 6 (02) ◽  
pp. 249-260 ◽  
Author(s):  
P. Jagers

Consider an age-dependent branching process with two types of individuals. Suppose that individuals of one type beget children of both types, whereas those of the other type can only give birth to individuals of their own kind. This paper is a study of the relation between two random functions occurring in such processes starting from an ancestor of the first type, the two functions being the numbers of individuals of the two kinds. Under weak assumptions it is shown that the random proportion of individuals of one type converges as time passes, in mean square as well as almost surely to a non-random limit, easily determined in terms of the reproduction laws and life-length distributions of the process.


1971 ◽  
Vol 8 (4) ◽  
pp. 655-667 ◽  
Author(s):  
M. L. Samuels

SummaryIn a standard age-dependent branching process, let Rn(t) denote the proportion of the population belonging to the nth generation at time t. It is shown that in the supercritical case the distribution {Rn(t); n = 0, 1, …} has asymptotically, for large t, a (non-random) normal form, and that the mean ΣnRn(t) is asymptotically linear in t. Further, it is found that, for large n, Rn(t) has the shape of a normal density function (of t).Two other random functions are also considered: (a) the proportion of the nth generation which is alive at time t, and (b) the proportion of the nth generation which has been born by time t. These functions are also found to have asymptotically a normal form, but with parameters different from those relevant for {Rn(t)}.For the critical and subcritical processes, analogous results hold with the random variables replaced by their expectations.


1971 ◽  
Vol 8 (04) ◽  
pp. 655-667 ◽  
Author(s):  
M. L. Samuels

Summary In a standard age-dependent branching process, let Rn (t) denote the proportion of the population belonging to the nth generation at time t. It is shown that in the supercritical case the distribution {Rn (t); n = 0, 1, …} has asymptotically, for large t, a (non-random) normal form, and that the mean ΣnRn (t) is asymptotically linear in t. Further, it is found that, for large n, Rn (t) has the shape of a normal density function (of t). Two other random functions are also considered: (a) the proportion of the nth generation which is alive at time t, and (b) the proportion of the nth generation which has been born by time t. These functions are also found to have asymptotically a normal form, but with parameters different from those relevant for {Rn (t)}. For the critical and subcritical processes, analogous results hold with the random variables replaced by their expectations.


1974 ◽  
Vol 6 (02) ◽  
pp. 291-308 ◽  
Author(s):  
Robert Fildes

In a branching process with variable lifetime, introduced by Fildes (1972) define Yjk (t) as the number of particles alive in generation k at time t when the initial particle is born in generation j. A limit law similar to that derived in the Bellman-Harris process is proved where it is shown that Yjk (t) suitably normalised converges in mean square to a random variable which is the limit random variable of Znm–n in the Galton-Watson process (m is the mean number of particles born).


1972 ◽  
Vol 4 (3) ◽  
pp. 453-474 ◽  
Author(s):  
Robert Fildes

A branching process with variable lifetime distribution is defined by a sequence of distribution functions {Gi(t)}, together with a probability generating function, h(s) = Σk∞= 0pksk. An ith generation particle lives a random length of time, determined by Gi(t). At the end of a particle's life it produces children, the number being determined by h(s). These offspring behave like the initial particle except they are (i + 1)th generation particles and have lifetime distribution Gi + 1 (t).Let Zi(t) be the number of particles alive at time t, the initial particle being born into the ith generation. Integral equations are derived for the moments of Zi(t) and it is shown that for some constants Ni, γ, a, Zi (t)/(Nitγ-1eαt) converges in mean square to a proper random variable.


1974 ◽  
Vol 6 (2) ◽  
pp. 291-308 ◽  
Author(s):  
Robert Fildes

In a branching process with variable lifetime, introduced by Fildes (1972) define Yjk(t) as the number of particles alive in generation k at time t when the initial particle is born in generation j. A limit law similar to that derived in the Bellman-Harris process is proved where it is shown that Yjk(t) suitably normalised converges in mean square to a random variable which is the limit random variable of Znm–n in the Galton-Watson process (m is the mean number of particles born).


1974 ◽  
Vol 11 (04) ◽  
pp. 678-686
Author(s):  
Edgar Z. Ganuza ◽  
S. D. Durham

Letting Z(t) be the number of objects alive at time t in a general supercritical age-dependent branching process generated by a single ancestor born at time 0, one achieves (Theorem 1) mean-square convergence of Z(t)/E[Z(t)] provided and , where N(t) is the number of offspring of the initial ancestor born by time t and α is the (positive) Malthusian parameter defined by . If the stronger conditions that (Theorem 2) and hold also, then the convergence is almost-sure. It is of interest that the embedded Galton-Watson process of successive generations need not have a finite mean for the conditions of the above theorems to hold. Similar results are obtained for the age-distribution as well.


1974 ◽  
Vol 11 (4) ◽  
pp. 678-686 ◽  
Author(s):  
Edgar Z. Ganuza ◽  
S. D. Durham

Letting Z(t) be the number of objects alive at time t in a general supercritical age-dependent branching process generated by a single ancestor born at time 0, one achieves (Theorem 1) mean-square convergence of Z(t)/E[Z(t)] provided and , where N(t) is the number of offspring of the initial ancestor born by time t and α is the (positive) Malthusian parameter defined by . If the stronger conditions that (Theorem 2) and hold also, then the convergence is almost-sure. It is of interest that the embedded Galton-Watson process of successive generations need not have a finite mean for the conditions of the above theorems to hold. Similar results are obtained for the age-distribution as well.


1972 ◽  
Vol 4 (03) ◽  
pp. 453-474 ◽  
Author(s):  
Robert Fildes

A branching process with variable lifetime distribution is defined by a sequence of distribution functions {G i (t)}, together with a probability generating function, h(s) = Σ k ∞= 0pks k . An ith generation particle lives a random length of time, determined by G i (t). At the end of a particle's life it produces children, the number being determined by h(s). These offspring behave like the initial particle except they are (i + 1)th generation particles and have lifetime distribution G i + 1 (t). Let Z i (t) be the number of particles alive at time t, the initial particle being born into the ith generation. Integral equations are derived for the moments of Z i (t) and it is shown that for some constants N i , γ, a, Z i (t)/(N i t γ-1 e αt ) converges in mean square to a proper random variable.


1973 ◽  
Vol 10 (03) ◽  
pp. 652-658
Author(s):  
J. Radcliffe

The mean square and almost sure convergence of W(t) = e–αt Z(t) is proved for a super-critical multitype age-dependent branching process allowing immigration at the epochs of a renewal process. It is shown that the Malthusian parameter, asymptotic frequencies of types and stationary age distributions are the same for the processes with and without immigration.


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