scholarly journals Cost of Locomotion: Unsteady Medusan Swimming

1985 ◽  
Vol 119 (1) ◽  
pp. 149-164 ◽  
Author(s):  
THOMAS L. DANIEL

The influence of unsteady (time varying) motion on the energetics of swimming was investigated with measurements and theoretical estimates of the specific cost of locomotion for two species of hydrozoan medusae: Gonionemus vertens L. Agassiz and Stomotoca atra L. Agassiz. These species, both about 1 g, provide a broad range of swimming speeds for which locomotor energetics can be explored. The cost of locomotion (a dimensionless ratio defined as the rate of energy consumption divided by the product of an animal's weight and speed) was estimated from the oxygen consumption rate of medusae tethered to a force platform. Swimming beat frequency, as monitored by the force platform, was correlated with velocity obtained from ciné-films of freely swimming medusae. The specific cost of locomotion was 6.3 (dimensionless), nearly one order of magnitude greater than the extrapolated cost of locomotion for a vertebrate swimmer of equivalent body mass. The great magnitude of this cost is attributed to two aspects of the periodic pulsatile pattern of locomotion by these cnidarians: (1) the energy expenditure for periodic accelerations of the animal's mass and some mass of fluid about it and (2) the energy dissipated in bell deformations and recovery strokes. Nearly 25% of the augmented cost is attributed to the energy required to overcome an unsteady flow force, that is the force required to accelerate fluid about the animal. Such a high cost of locomotion is apparently a general consequence of swimming with a discontinuous production of thrust. The mechanics of discontinuous swimming are explored by measuring the hydrodynamic coefficients associated with unsteady flows (added-mass coefficients) for models of medusae. The results suggest that the effects of vortex formation and shedding may significantly increase the magnitude of the forces produced by or resisting unsteady animal locomotion.

2011 ◽  
Vol 14 (2) ◽  
Author(s):  
Thomas G Koch

Current estimates of obesity costs ignore the impact of future weight loss and gain, and may either over or underestimate economic consequences of weight loss. In light of this, I construct static and dynamic measures of medical costs associated with body mass index (BMI), to be balanced against the cost of one-time interventions. This study finds that ignoring the implications of weight loss and gain over time overstates the medical-cost savings of such interventions by an order of magnitude. When the relationship between spending and age is allowed to vary, weight-loss attempts appear to be cost-effective starting and ending with middle age. Some interventions recently proven to decrease weight may also be cost-effective.


2011 ◽  
Vol 279 (1729) ◽  
pp. 640-644 ◽  
Author(s):  
Graham N. Askew ◽  
Federico Formenti ◽  
Alberto E. Minetti

In Medieval Europe, soldiers wore steel plate armour for protection during warfare. Armour design reflected a trade-off between protection and mobility it offered the wearer. By the fifteenth century, a typical suit of field armour weighed between 30 and 50 kg and was distributed over the entire body. How much wearing armour affected Medieval soldiers' locomotor energetics and biomechanics is unknown. We investigated the mechanics and the energetic cost of locomotion in armour, and determined the effects on physical performance. We found that the net cost of locomotion ( C met ) during armoured walking and running is much more energetically expensive than unloaded locomotion. C met for locomotion in armour was 2.1–2.3 times higher for walking, and 1.9 times higher for running when compared with C met for unloaded locomotion at the same speed. An important component of the increased energy use results from the extra force that must be generated to support the additional mass. However, the energetic cost of locomotion in armour was also much higher than equivalent trunk loading. This additional cost is mostly explained by the increased energy required to swing the limbs and impaired breathing. Our findings can predict age-associated decline in Medieval soldiers' physical performance, and have potential implications in understanding the outcomes of past European military battles.


2011 ◽  
Vol 480-481 ◽  
pp. 1197-1200
Author(s):  
Feng Liu ◽  
Jun Min Wang

In recent years, in order to reduce building energy consumption in China, and vigorously promote the development of ecological construction, the paper studies disadvantages of the cost of traditional project management applications in the ecological construction projects, exploring specific cost management adapt to the ecological construction project.


1988 ◽  
Vol 138 (1) ◽  
pp. 301-318 ◽  
Author(s):  
N. C. Heglund ◽  
C. R. Taylor

In this study we investigate how speed and stride frequency change with body size. We use this information to define ‘equivalent speeds’ for animals of different size and to explore the factors underlying the six-fold difference in mass-specific energy cost of locomotion between mouse- and horse-sized animals at these speeds. Speeds and stride frequencies within a trot and a gallop were measured on a treadmill in 16 species of wild and domestic quadrupeds, ranging in body size from 30 g mice to 200 kg horses. We found that the minimum, preferred and maximum sustained speeds within a trot and a gallop all change in the same rather dramatic manner with body size, differing by nine-fold between mice and horses (i.e. all three speeds scale with about the 0.2 power of body mass). Although the absolute speeds differ greatly, the maximum sustainable speed was about 2.6-fold greater than the minimum within a trot, and 2.1-fold greater within a gallop. The frequencies used to sustain the equivalent speeds (with the exception of the minimum trotting speed) scale with about the same factor, the −0.15 power of body mass. Combining this speed and frequency data with previously published data on the energetic cost of locomotion, we find that the mass-specific energetic cost of locomotion is almost directly proportional to the stride frequency used to sustain a constant speed at all the equivalent speeds within a trot and a gallop, except for the minimum trotting speed (where it changes by a factor of two over the size range of animals studied). Thus the energy cost per kilogram per stride at five of the six equivalent speeds is about the same for all animals, independent of body size, but increases with speed: 5.0 J kg-1 stride-1 at the preferred trotting speed; 5.3 J kg-1 stride-1 at the trot-gallop transition speed; 7.5 J kg-1 stride-1 at the preferred galloping speed; and 9.4 J kg-1 stride-1 at the maximum sustained galloping speed. The cost of locomotion is determined primarily by the cost of activating muscles and of generating a unit of force for a unit of time. Our data show that both these costs increase directly with the stride frequency used at equivalent speeds by different-sized animals. The increase in cost per stride with muscles (necessitating higher muscle forces for the same ground reaction force) as stride length increases both in the trot and in the gallop.


1995 ◽  
Vol 198 (3) ◽  
pp. 629-632 ◽  
Author(s):  
V A Langman ◽  
T J Roberts ◽  
J Black ◽  
G M Maloiy ◽  
N C Heglund ◽  
...  

Large animals have a much better fuel economy than small ones, both when they rest and when they run. At rest, each gram of tissue of the largest land animal, the African elephant, consumes metabolic energy at 1/20 the rate of a mouse; using existing allometric relationships, we calculate that it should be able to carry 1 g of its tissue (or a load) for 1 km at 1/40 the cost for a mouse. These relationships between energetics and size are so consistent that they have been characterized as biological laws. The elephant has massive legs and lumbers along awkwardly, suggesting that it might expend more energy to move about than other animals. We find, however, that its energetic cost of locomotion is predicted remarkably well by the allometric relationships and is the lowest recorded for any living land animal.


1999 ◽  
Vol 86 (5) ◽  
pp. 1657-1662 ◽  
Author(s):  
Young-Hui Chang ◽  
Rodger Kram

Previous studies have suggested that generating vertical force on the ground to support body weight (BWt) is the major determinant of the metabolic cost of running. Because horizontal forces exerted on the ground are often an order of magnitude smaller than vertical forces, some have reasoned that they have negligible cost. Using applied horizontal forces (AHF; negative is impeding, positive is aiding) equal to −6, −3, 0, +3, +6, +9, +12, and +15% of BWt, we estimated the cost of generating horizontal forces while subjects were running at 3.3 m/s. We measured rates of oxygen consumption (V˙o 2) for eight subjects. We then used a force-measuring treadmill to measure ground reaction forces from another eight subjects. With an AHF of −6% BWt,V˙o 2 increased 30% compared with normal running, presumably because of the extra work involved. With an AHF of +15% BWt, the subjects exerted ∼70% less propulsive impulse and exhibited a 33% reduction inV˙o 2. Our data suggest that generating horizontal propulsive forces constitutes more than one-third of the total metabolic cost of normal running.


2009 ◽  
Vol 13 (2) ◽  
Author(s):  
Richard C. Hicks ◽  
Keith Wright

Implementations of inference engine systems invoke many costs, including the cost of the inference engine itself, the cost of integrating the inference engine, and the cost of specialized personnel needed to create and maintain the system. These costs make a very high return on investment a criterion for incorporating these systems into the corporate portfolio of applications and technologies. Recently, the No Inference Engine Theory (NIET) [8] has been developed for creating procedural propositional logic rule-based systems. The NIET systems are implemented in traditional procedural languages such as C++ and do not need an inference engine or proprietary languages, thus eliminating the cost of the inference engine, the cost of integrating the system, and the cost for knowledge of a proprietary language. In addition, these procedural systems are an order of magnitude faster [8] than inference systems and maintain linear performance. For problems using propositional logic, the procedural systems described in this paper offer dramatically lower costs, higher performance, and ease of integration. Lowering the external costs and eliminating the need for specialized skills should make NIET systems more profitable and lead to the wider use of propositional logic systems in business.


Author(s):  
Eric Timmons ◽  
Brian C. Williams

State estimation methods based on hybrid discrete and continuous state models have emerged as a method of precisely computing belief states for real world systems, however they have difficulty scaling to systems with more than a handful of components. Classical, consistency based diagnosis methods scale to this level by combining best-first enumeration and conflict-directed search. While best-first methods have been developed for hybrid estimation, conflict-directed methods have thus far been elusive as conflicts summarize constraint violations, but probabilistic hybrid estimation is relatively unconstrained. In this paper we present an approach (A*BC) that unifies best-first enumeration and conflict-directed search in relatively unconstrained problems through the concept of "bounding" conflicts, an extension of conflicts that represent tighter bounds on the cost of regions of the search space. Experiments show that an A*BC powered state estimator produces estimates up to an order of magnitude faster than the current state of the art, particularly on large systems.


2021 ◽  
Author(s):  
Robert Godin ◽  
James R. Durrant

The energy cost of lifetime gain in solar energy conversion systems is determined from a breadth of technologies. The cost of 87 meV per order of magnitude lifetime improvement is strikingly close to the 59 meV determined from a simple kinetic model.


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