Response of northwestern chipmunks (Tamias amoenus) to variable habitat structure in young lodgepole pine forest
This study was designed to test the hypothesis that large-scale habitat alteration by stand thinning over a range of densities would increase the abundance and related population dynamics of northwestern chipmunks (Tamias amoenus) in young lodgepole pine (Pinus contorta) forest. Replicate study areas were located near Penticton, Kamloops, and Prince George in south-central British Columbia, Canada. Each study area had three stands thinned to densities of approximately 500 (low), 1000 (medium), and 2000 (high) stems/ha, with an unthinned young pine stand and an old-growth pine stand for comparison. Chipmunk populations were sampled intensively in thinned stands from 1989 to 1991 and in the unthinned and old-growth stands from 1990 to 1991. Habitat structure was sampled in all stands in 1990. For herbs and shrubs, the crown volume index values were similar among stands; for trees, this index was lowest for the low-density stands, with the index for all three thinned stands being lower than that for the unthinned stands. Species diversity and the structural diversity of vegetation were similar among stands. The abundance of chipmunks was significantly higher in low-density than in high-density thinned stands at Penticton (1.3-1.9 times higher) and Prince George (2.4-3.8 times higher) but not at Kamloops. Chipmunks were less abundant in old-growth stands than in the other four treatment stands. Breeding performance and recruitment followed the same pattern as abundance. Chipmunk survival was generally similar among stands. There were heavier chipmunks in the low-density stands in some years at Penticton. Northwestern chipmunks appear to prefer "open" habitats generated by heavy thinning of young lodgepole pine stands. This result was achieved in three different forest ecological zones and may enhance the overall forest ecosystem.