Morphological relationships among subspecies of the Rufous-collared Sparrow, Zonotrichia capensis
The phenetic relationships among 22 described subspecies of Zonotrichia capensis, the Rufous-collared Sparrow, are investigated by analyses of variation in six external metric variables taken from over 1500 museum specimens. Both sexes show the same marked dichotomy between two groups of subspecies: pulacayensis, sanborni, antofagastae, chilensis, choraules, and australis versus all others. These six subspecies are from temperate high altitudes and high latitudes and occupy a geographically contiguous region in the southern Andes mountains and Patagonia. Within these two subgroups there is extensive overlap among individuals; finer division of the subspecies is difficult, and involves the association of subspecies from geographically disjunct regions. Thus the taxonomic hierarchy appears to be a poor representation of variation of the characters considered here. The association of geographically disjunct forms suggests that intraspecific variation in these characters reflects the local environment (via selection and (or) environmental effects) rather than historical (genealogical) relationships.