Variation in the choanal structure of Recent turtles
The choanal regions of all the living genera and almost all of the living forms of turtles are described, based on a sample of over 4000 specimens. In all families of cryptodires except the cheloniids and derrnochelyids there are usually ridges or flaps along the lateral margins of the choanae. In chelydrids, most kinosternids, some platystemids, some emydids, and some trionychids there is a papilla or bulge extending medially into the choana from this flap or ridge, and in dermatemydids, carettochelyids, and some trionychids there may be several or many such papillae, Cheloniids have one, or in Chelonia many, papillae along the lateral choanal margin. Other forms lack these papillae. Dermochelyids and pleurodires have no special structures along the lateral choanal margins. In almost all cases there is some and often there is considerable variation in these structures.The possible phylogenetic and taxonomic implications of the findings are discussed. In some groups such as the genera of emydids, the choanal structure may prove to be taxonomically useful, but in other cases it does not seem helpful. For example, in the trionychids, the intra-specific variation is so great that it largely obscures the interspecific differences.