Phylogeny of Palaearctic Pharyngodonidae parasite species of Testudinidae: a morphological approach

2003 ◽  
Vol 81 (11) ◽  
pp. 1885-1893 ◽  
Author(s):  
Salah Bouamer ◽  
Serge Morand

The phylogenetic relationships of 23 oxyurid species from five genera (21 parasite species of the Palaearctic Testudinidae, 1 parasite species of Uromastix acanthinurus Bell, 1825 from Algeria, and 1 parasite species of Cteno sa ura pectinata (Wiegmann, 1834) from Mexico) were investigated using 30 morphological characters obtained from species descriptions. The nonweighted analysis produced one shortest tree. All species of the ingroup form a monophyletic group and the oxyurid species of Testudinidae form a monophyletic group. The type species of the genus Alaeuris Thapar, 1925 is the basal member of the species parasitizing Testudinidae. The analysis confirms the monophyly of the genus Thaparia Ortlepp, 1933, whereas the genera Mehdiella Seurat, 1918 and Tachygonetria Wedl, 1862 are considered paraphyletic groups. The large diversification in the genus Tachygonetria is linked to their position in the host caecum. The ancestral state is in the paramucous and the derived state is in the centre of the caecum. This suggests that recent speciation in the group occurs in the centre of the caecum.

Zootaxa ◽  
2002 ◽  
Vol 87 (1) ◽  
pp. 1 ◽  
Author(s):  
FRANK LOUIS CARLE ◽  
KARL M. KJER

Phylogenetic analysis was performed on a set of 242 morphological characters. The taxon sample included 31 Libellula, and representative species from selected libeluline tribes, from all libellulid subfamilies, from all libelluloid families, from all anisopteran superfamilies, and Epiophlebia. Corduliinae was shown to be paraphyletic even among genera characterized by a well developed anal loop bisector. Sympetrini was found to be polyphyletic with Crocothemis the sister group to Libellulini. The traditional placement of Trameini, far from Libellulini is in doubt, because it is here placed as the sister group to Crocothemis + Libellulini. Kennedy’s phylogeny of Libellula was largely corroborated, with the following exceptions: the subgenera Libellula, Eolibellula, and Syntetrum form a monophyletic group which is the sister group to a clade including Belonia, Holotania, Neotetrum, and Eotainia subgenus nov. [type species Mesothemis composita Hagen]; and Eurothemis is determined to be the sister group of Ladona instead of Neotetrum. In addition we confirm Belonia to be monophyletic, and find Platetrum + Plathemis to form a monophyletic group, sister to Ladona + Eurothemis; these four subgenera together form the sister group to Libellula sensu stricto (s.s.).


Phytotaxa ◽  
2015 ◽  
Vol 201 (3) ◽  
pp. 197
Author(s):  
Rafael Lira ◽  
Victoria Sosa ◽  
Talitha Legaspi ◽  
Patricia Dávila

We examine the limits and phylogenetic relationships of Ibervillea and Tumamoca belonging to tribe Coniandreae in the Cucurbitaceae. These taxa are found in xeric areas from southern United States to Guatemala. There has been no previous phylogenetic studies considering all their taxa together, just partially. Furthermore, we include as well species of Dieterlea, another similar and sympatric genus which recognition is under debate, formerly considered as a synonym of Ibervillea. Using molecular and morphological characters we performed molecular and total evidence parsimony and Bayesian analyses. Our results confirm that species in Ibervillea and Dieterlea are part of a monophyletic group, supporting the integration of both genera as proposed in previous phylogenetic and taxonomic studies. By examining all the species of the three genera, our results are the first to suggest that Tumamoca is also part of this monophyletic group. Therefore we propose that the species of Ibervillea, Dieterlea, and one species of Tumamoca should be included into the same genus. According to the priority rule, Ibervillea is the name to be used. Tumamoca macdougalli needs to be transferred to Ibervillea and the combination is made here. We suggest that the second species in Tumamoca, T. mucronata, known by a single specimen that is lost, does not deserve recognition.


Zootaxa ◽  
2008 ◽  
Vol 1926 (1) ◽  
pp. 41-52 ◽  
Author(s):  
TERRY A. WHEELER ◽  
JULIA J. MLYNAREK

Agrophaspidium gen. n. is described for six species: Agrophaspidium flavimana sp. n. (type locality: La Selva, Costa Rica); Agrophaspidium hastatum Duda, 1930, comb. n. (type locality: Suiza de Turrialba, Costa Rica); Agrophaspidium minutum sp. n. (type locality: Gatun Lake, near Colón, Panama); Agrophaspidium monticola sp. n. (type species of genus; type locality: Monteverde, Costa Rica); Agrophaspidium pollinosum sp. n. (type locality: Portachuelo Pass, Venezuela); and Agrophaspidium psilotum sp. n. (type locality: Guanacaste National Park, Costa Rica). Agrophaspidium is distinguished from other Oscinellinae by the combination of an anteromedial-posterolateral arrangement of the vertical bristles, a dorsally flattened scutellum with four elongate marginal projections bearing stout bristles, and characters of the male genitalia (enlarged epandrium, branched surstylus, enlarged and modified cercus). Although the phylogenetic relationships of species within Agrophaspidium are resolved by morphological characters, the generic relationships to other Oscinellinae are unresolved.


Zootaxa ◽  
2010 ◽  
Vol 2553 (1) ◽  
pp. 35 ◽  
Author(s):  
MARJOLAINE GIROUX ◽  
TERRY A. WHEELER

Sarcophaga (Bulbostyla) subgen. nov. is described as a new subgenus of Sarcophaga Meigen to accommodate some species previously assigned to the subgenus S. (Neobellieria) Blanchard. Sarcophaga (Bulbostyla) contains nine species: S. airosalis sp. nov., S. cadyi sp. nov. (type species), S. cuautla sp. nov., S. fattigina sp. nov., S. ironalis sp. nov., S. semimarginalis Hall, S. sternalis (Reinhard), S. subdiscalis Aldrich and S. yorkii Parker. All species are described and illustrated and a key to the species is provided. The species within the subgenus are morphologically uniform externally and are distinguished mostly on male genitalic characters. Phylogenetic relationships within Bulbostyla are unresolved based on morphological characters and will require consideration of additional characters, such as molecular sequence data. The genus-group taxon Robackina Lopes is removed from synonymy with the subgenus Sarcophaga (Neobellieria) and reinstated as a valid subgenus of Sarcophaga (stat. nov.) to accommodate the single New World species Sarcophaga triplasia Wulp. A lectotype is designated for S. triplasia. The subgenus and species are redescribed and illustrated.


Zootaxa ◽  
2007 ◽  
Vol 1401 (1) ◽  
pp. 33 ◽  
Author(s):  
REBECCA L. HUNTER

A phylogenetic analysis using morphological characters was done on the Antarctic ophiuroid genus Ophiurolepis Matsumoto, 1915. This genus is one of the more abundant and ecologically dominant ophiuroid genera in the Antarctic and surrounding Southern Ocean. Maximum parsimony was used to infer phylogenetic relationships. Although strongly supported nodes were not recovered for most groupings within Ophiurolepis, this first attempt at a phylogeny revealed the presence of three tentative clades. Two of the three Ophiurolepis clades included species currently assigned to other genera, but closely allied with Ophiurolepis in the taxonomic literature. This indicates that Ophiurolepis as currently defined is not a monophyletic group. Additional forms of data, namely molecular, are needed to more definitively resolve relationships within this group.


Author(s):  
Guillermo E Terán ◽  
Mauricio F Benitez ◽  
J Marcos Mirande

Abstract The freshwater fish genus Astyanax is one of the most diverse among the Characidae. The genus is defined by a combination of character states that are widely distributed in Characidae. In addition, the genus has the broadest geographical distribution in the family, being found in a great variety of environments of the Neotropical region. Although phylogenetic relationships were treated only partially, many authors agree that the genus is not monophyletic. In this contribution, we study the phylogenetic relationships of Astyanax in the context of the family Characidae, by combining morphological and molecular data. A total of 520 morphological characters, nine molecular markers and 608 taxa are analysed, of which 98 belong to Astyanax. According to our results, Astyanax is not monophyletic. We recovered species attributed to Astyanax in different subfamilies: Gymnocharacinae (including the type species), Stevardiinae and Tetragonopterinae. Among the species recovered in Gymnocharacinae, most (including the type species, the resurrected Psalidodon, and the new genus Andromakhe gen. nov.) were recovered in Gymnocharacini, while the remaining ones were recovered in Probolodini (transferred to Deuterodon or the new genus Makunaima gen. nov.).


2006 ◽  
Vol 84 (12) ◽  
pp. 1753-1770 ◽  
Author(s):  
José L. Martínez-y-Pérez ◽  
Teresa Mejía-Saulés ◽  
Victoria Sosa

Luziola is a small aquatic genus consisting of nine American species and belonging to Poaceae (subfamily Ehrhartoideae, tribe Oryzeae). We surveyed 160 morphological characters and performed phylogenetic analyses to determine relationships within the genus and between it and its closest relatives, and to deduce the evolution of a number of characters in relation to the aquatic habit. Our results confirm that Luziola is a monophyletic group and that Zizaniopsis is its sister group among taxa examined but revealed only a few clades within Luziola, the best supported of which consists of Luziola fragilis Swallen, Luziola fluitans (Michx.) Terrell & H. Rob., and Luziola peruviana J.F. Gmel. We used this phylogenetic evidence to reconstruct the evolution of several characters hypothesized to be correlated to an aquatic life style, including reduction or absence of glumes, papillae arranged in horizontal rows on the lemmas, dumb-bell shaped silica bodies, and a change from hermaphrodite to unisexual florets.


Zootaxa ◽  
2018 ◽  
Vol 4471 (1) ◽  
pp. 1
Author(s):  
JULIA J. MLYNAREK ◽  
TERRY A. WHEELER

The phylogenetic relationships of the chloropid tribe Elachipterini were analysed. Sixty-eight exemplar species and seven outgroup species were included in a cladistic analysis based on 76 morphological characters of adult specimens in order to test existing, non-phylogenetic, classifications of the tribe. Nine genera are recognized in the Elachipterini: Allomedeia Mlynarek & Wheeler, Alombus Becker, Anatrichus Loew, Ceratobarys Coquillett, Disciphus Becker, Elachiptera Macquart, Goniaspis Duda, Melanochaeta Bezzi and Sepsidoscinis Hendel. Myrmecosepsis Kertesz is synonymised with Anatrichus, and Togeciphus Nishijima and Cyrtomomyia Becker are synonymised with Elachiptera. Ceratobarys is removed from synonymy with Elachiptera and all Neotropical species and two Nearctic species previously assigned to Elachiptera are transferred to Ceratobarys. Melanochaeta is a valid genus; the type species Melanochaeta capreolus clusters with other species of Melanochaeta and not Oscinella. New combinations include Anatrichus hystrix (Kertesz, 1914) (Myrmecosepsis); Anatrichus taprobane (Andersson, 1977) (Myrmecosepsis), Ceratobarys attenuata (Adams, 1908) (Elachiptera); Ceratobarys cultrata (Wheeler & Forrest, 2002) (Elachiptera); Ceratobarys flavida (Williston, 1896) (Elachiptera); Ceratobarys melinifrons (Mlynarek & Wheeler, 2008) (Elachiptera); Ceratobarys fucosa (Mlynarek & Wheeler, 2008) (Elachiptera); Ceratobarys queposana (Mlynarek & Wheeler, 2008) (Elachiptera); Ceratobarys rubida (Becker, 1912) (Elachiptera); Ceratobarys sacculicornis (Enderlein, 1911) (Elachiptera); Ceratobarys willistoni (Sabrosky, 1948)  (Elachiptera), Elachiptera ensifer (Sabrosky, 1951) (Cyrtomomyia); Elachiptera ericius (Kanmiya, 1983) (Togeciphus); Elachiptera katoi (Nishijima, 1955) (Togeciphus); Elachiptera maculinervis (Becker, 1910) (Cyrtomomyia); Elachiptera punctulata (Becker, 1912) (Cyrtomomyia); Elachiptera subelongata (Kanmiya, 1983) (Disciphus); Elachiptera truncatus (Liu & Yang, 2012) (Togeciphus); Elachiptera tuberculata (Adams, 1905) (Cyrtomomyia) and all the species that were placed in Lasiochaeta are returned to Melanochaeta. A key to genera of the tribe Elachipterini is provided and diagnoses are provided for all genera. The tribe is divided into two geographically distinct clades: the Anatrichus clade includes the Old World tropical genera Allomedeia, Alombus, Anatrichus, Disciphus and Sepsidoscinis; the Elachiptera clade includes the primarily Neotropical genera Goniaspis and Ceratobarys and the widespread, but primarily Holarctic, genera Elachiptera and Melanochaeta. 


2001 ◽  
Vol 58 (2) ◽  
pp. 183-200 ◽  
Author(s):  
G. M. PLUNKETT

Phylogenetic relationships involving the angiosperm order Apiales (Apiaceae and Araliaceae) are troublesome at nearly every taxonomic level and have eluded several generations of botanists. Because of difficulties in interpreting and polarizing morphological character states at deeper phylogenetic levels, most studies in Apiales have focused on relationships between the two families and among/within the apialean genera. In the present study, however, recent contributions from molecular analyses are reviewed and combined using a ‘supertree’ approach to test traditional hypotheses of relationships involving Apiales, and to re-evaluate assumptions of character-state evolution in the order. Results from this study confirm that Apiales form a monophyletic group with Pittosporaceae (along with Griselinia G. Forst., Melanophylla Baker, Torricellia DC. and Aralidium Miq.), and should be transferred out of subclass Rosidae (away from both Cornales and Sapindales) to the Asteridae (in a position close to Asterales and Dipsacales). These findings are also supported by several lines of morphological, anatomical, and phytochemical evidence, and provide a more satisfactory framework for interpreting relationships and character-state evolution within the major clades of Apiales.


Zootaxa ◽  
2007 ◽  
Vol 1575 (1) ◽  
pp. 35-45 ◽  
Author(s):  
KÔJI SASAKAWA

The supraspecific taxonomy of the coastal carabid Bembidion (‘Cillenus’) complex is controversial because a single ‘synapomorphy,’ a large head with long mandibles, may be the result of functional convergence. This study examined the phylogeny of B. (‘Cillenus’) complex based on morphological characters to revise the complicated taxonomy of this group. All type species of the genera concerned were included, and membranous parts of the genitalia (male endophallus and female vagina and spermatheca) were examined for the first time. Because all species are treated under the genus Bembidion Latreille (s. l.), Bembidion umi nom. nov. is proposed as a new name for Sakagutia marina Uéno. In the resultant tree, reciprocal monophyly of most traditional taxa was not supported, whereas a monophyletic group of the following seven species was relatively well supported: B. seticorne Lindroth, B. yokohamae (Bates), B. sumaoi Morita, B. aestuarii (Uéno & Habu), B. tokunoshimanum (Nakane), B. tsutsuii (Uéno), and B. yakushimanum sp. nov. Based on these results, the subgenus Desarmatocillenus Netolitzky (type species: B. yokohamae) was redefined, and two subgenera were synonymised with it: Novicillenus Uéno & Habu (type species: B. aestuarii) syn. nov., and Corallicillenus Uéno (type species: B. tsutsuii) syn. nov.. Bembidion (Desarmatocillenus) yakushimanum sp. nov. is described from Japan.


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