Coarse woody debris decomposition documented over 65 years on southern Vancouver Island

1998 ◽  
Vol 28 (5) ◽  
pp. 788-793 ◽  
Author(s):  
Jeff N Stone ◽  
Andy MacKinnon ◽  
John V Parminter ◽  
Ken P Lertzman
Keyword(s):  
Decay Rate ◽  
Pacific Northwest ◽  
Rate Constant ◽  
Coarse Woody Debris ◽  
Woody Debris ◽  
Vancouver Island ◽  
Douglas Fir ◽  
Permanent Plots ◽  
Decay Rate Constant ◽  
Exponential Decay Rate

In 1929-1930, Stig Schenström and J.D. Curtis established an experiment to study thinning dynamics of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) on southern Vancouver Island, British Columbia. As a subcomponent of the experiment, the coarse woody debris (CWD) from the previous stand were mapped and the decay condition classified on five permanent plots. These scaled drawings and classifications were updated in 1945-1947 and 1995-1996. This unique 65-year period of CWD observation confirms that observations of CWD volume loss on Vancouver Island are similar to elsewhere in the Pacific Northwest. The simple exponential decay rate constant was 0.022 ·year-1 based on volume of primarily Douglas-fir on the five plots. The decay rate constant by large-end diameter was 0.067 ·year-1 for logs <= 20 cm, 0.056 ·year-1 for 21-40 cm, 0.021 ·year-1 for 41-80 cm, and 0.012 ·year-1 for logs > 80 cm.

Coarse woody debris in mixed-conifer forests, Sequoia National Park, California

1987 ◽  
Vol 17 (10) ◽  
pp. 1265-1272 ◽  
Author(s):  
Mark E. Harmon ◽  
Kermit Cromack Jr. ◽  
Bradley G. Smith
Keyword(s):  
Decay Rate ◽  
National Park ◽  
Woody Debris ◽  
Permanent Plots ◽  
Nitrogen Storage ◽  
Decay Rate Constant ◽  
Stand Volume ◽  
Sequoia National Park ◽  
Mixed Conifer Forests ◽  
Class Information

The decay rate of Abiesconcolor (Gord. & Glend.) Lindl. logs and cover, mass, and volume of logs and snags in six midelevational forest stands of Sequoia National Park, California, are reported. Based on a chronosequence, Abiesconcolor boles have a decay rate-constant of 0.05 year−1 and a half-life of 14 years. A decay classification system was developed for Abiesconcolor, Calocedrusdecurrens (Torr.) Florin, Pinusjeffreyi Grev. & Balf., and Pinuslambertiana Dougl. logs. Dimensions taken from maps of six permanent plots were combined with decay-class information to estimate volume, mass, and projected cover of logs and snags. Total mass ranged from 29 Mg ha−1 in a Pinusjeffreyi forest to 400 Mg ha−1 in a Sequoiadendrongiganteum (Lindl.) Buchh. dominated stand. Volume, projected cover, and nitrogen storage exhibited patterns similar to mass, ranging from 84 to 1160 m3 ha−1, 3.1 to 9.3%, and 41 to 449 kg ha−1, respectively.

Determination of the decay rate constant for hepatocytes immobilized in alginate microcapsules

2010 ◽  
Vol 27 (1) ◽  
pp. 86-93 ◽  
Author(s):  
Aldo Leal-Egaña ◽  
Aránzazu Díaz-Cuenca ◽  
Augustinus Bader
Keyword(s):  
Decay Rate ◽  
Rate Constant ◽  
Decay Rate Constant

Coarse Woody Debris in Douglas-Fir Forests of Western Oregon and Washington

Ecology ◽  
1988 ◽  
Vol 69 (6) ◽  
pp. 1689-1702 ◽  
Author(s):  
Thomas A. Spies ◽  
Jerry F. Franklin ◽  
Ted B. Thomas
Keyword(s):  
Coarse Woody Debris ◽  
Woody Debris ◽  
Douglas Fir

Managing for Mature Habitat in Production Forests of Western Oregon and Washington

1996 ◽  
Vol 10 (2) ◽  
pp. 422-428 ◽  
Author(s):  
Elizabeth C. Cole
Keyword(s):  
Pacific Northwest ◽  
Coarse Woody Debris ◽  
Management Practices ◽  
Woody Debris ◽  
Wildlife Habitat ◽  
Low Grade ◽  
Habitat Features ◽  
Canopy Opening ◽  
Tolerant Species ◽  
Shade Tolerant Species

Standard timber management practices in the Pacific Northwest result in stands which often vary from unmanaged stands in structure and composition. Forest and wildlife managers have identified a deficit of stands in the mature (> 100-yr-old) age class that contain certain desirable wildlife habitat features. Techniques are being developed that would increase the likelihood that managed stands can produce these characteristics. The key desirable components in these stands include large (> 75 cm diam breast height) conifer trees, snags, coarse woody debris, and understory structure, including regeneration. Vegetation management techniques can facilitate development of these components within stands. Thinning the overstory, underplanting shade-tolerant species, and creating snags and coarse woody debris can be accomplished within a production forest. Maintaining shade-intolerant species requires a higher level of disturbance and canopy opening than needed for shade-tolerant species. Treatments which remove competition from shrubs and herbaceous plants may be necessary to insure growth and survival of understory regeneration. Injection of different herbicides into low-grade conifers may yield different types of snags in comparison to girdling or topping. Although much of the understory may be eliminated during future thinnings and final harvest, some of the structure will remain and could be carried over into the next rotation along with snags and large coarse woody debris. These treatments are expected to enhance mature habitats in present and future cycles with minimum impact on yield.

scholarly journals Coarse woody debris in the old-growth forests of British Columbia

2003 ◽  
Vol 11 (S1) ◽  
pp. S135-S157 ◽  
Author(s):  
M C Feller
Keyword(s):  
British Columbia ◽  
Coarse Woody Debris ◽  
Functional Role ◽  
Animal Species ◽  
Woody Debris ◽  
Douglas Fir ◽  
Forest Age ◽  
Old Growth ◽  
Old Growth Forests

This paper synthesizes data extracted from the literature and data collected in various studies by the author on the quantity, characteristics, and functional importance of coarse woody debris (CWD) in the old-growth forests of British Columbia (B.C.). There is little agreement in the literature about the minimum diameter of CWD or the number of decay classes recognized. In western North America, five decay classes are commonly used, but recent studies suggest fewer decay classes are preferable. Comparisons among decay classes and biogeoclimatic zones and subzones in B.C. reveal that quantities and volumes are greatest (up to approximately 60 kg/m2 and approximately 1800 m3/ha, respectively), and CWD persists the longest (sometimes in excess of 1000 years) in the Coastal Western Hemlock (CWH) biogeoclimatic zone. The quantity and ground cover of CWD increase with forest productivity. Persistence of CWD has varied from less than 100 to over 800 years in two coastal (CWH and Mountain Hemlock (MH)) and three interior (Interior Douglas-fir (IDF), Interior Cedar–Hemlock (ICH), and Engelmann Spruce – Subalpine Fir (ESSF)) biogeoclimatic zones. Trends in CWD quantity with forest age in managed coastal B.C. forests suggest a U-shaped curve, with greater quantities occurring in recent cutovers than in old-growth forests, and lowest quantities occurring in middle-aged forests. This may be the normal trend in CWD with forest age, with departures from this trend resulting from disturbance- or environment-specific factors. Relatively large amounts of data exist on the characteristics of CWD in the CWH, IDF, ICH, ESSF, and Boreal White and Black Spruce (BWBS) biogeoclimatic zones, but such data for the Coastal Douglas-fir, Sub-Boreal Pine–Spruce, Sub-Boreal Spruce (SBS), and Spruce–Willow–Birch biogeoclimatic zones appear relatively sparse. There have been few studies of the functional role of CWD in B.C. forests, but those studies that have been completed indicate that CWD is an important habitat component for some plant and animal species. A total of 169 plant species, including >95% of all lichens and liverworts, were found to grow on CWD in old-growth forests in the CWH, MH, IDF, ICH, and ESSF biogeoclimatic zones. One third of these species were restricted to CWD. Studies in several biogeoclimatic zones have found that CWD provided preferred habitat for and was associated with higher populations of some small animal species, such as shrews, some voles, and some salamanders, in old-growth forests, but the effects varied with species and biogeoclimatic zone. The nutrient cycling role of CWD is not yet well known, but it currently appears to be relatively insignificant in B.C. old-growth forests. Although it has been considered that CWD could increase mineral soil acidification and eluviation, no evidence for this was found in a study of the CWH, MH, IDF, ICH, ESSF, BWBS, and SBS biogeoclimatic zones. Future studies of the functional role of CWD should consider both scale (square metre vs. hectare) and temporal (changes in CWD with forest age) issues, as studies including these are sparse and both may be important. Key words: biogeoclimatic zones, British Columbia, coarse woody debris, old-growth forests.

Coarse woody debris and forest floor respiration in an old-growth coniferous forest on the Olympic Peninsula, Washington, USA

1994 ◽  
Vol 24 (9) ◽  
pp. 1811-1817 ◽  
Author(s):  
James L. Marra ◽  
Robert L. Edmonds
Keyword(s):  
Coarse Woody Debris ◽  
Forest Floor ◽  
Woody Debris ◽  
Large Diameter ◽  
Small Diameter ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Respiration Rates ◽  
Decay Class ◽  
Class 1

Carbon dioxide evolution rates for downed logs (coarse woody debris) and the forest floor were measured in a temperate, old-growth rain forest in Olympic National Park, Washington, using the soda lime trap method. Measurements were taken every 4 weeks from October 22, 1991, to November 19, 1992. Respiration rates for Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) and western hemlock (Tsugaheterophylla (Raf.) Sarg.), logs were determined for decay classes 1–2, 3, and 5 in two diameter classes. Overall, western hemlock logs respired at a rate 35% higher (4.37 g CO2•m−2•day−1) than Douglas-fir logs (3.23 g CO2•m−2•day−1). Respiration rates for decay class 1–2 logs of both species were similar to decay class 5 logs (4.46 and 4.07 g CO2•m−2•day−1, respectively), but decay class 3 logs respired at a lower rate (3.23 g CO2•m−2•day−1). Seasonal patterns of respiration rates occurred, particularly for decay class 1 and 2 western hemlock logs where monthly averages ranged from a low of 2.67 g CO2•m−2•day−1 in February 1992 to a high of 8.30 g CO2•m−2•day−1 in September 1992. Rates for decay class 1–2 western hemlock logs were greater than those from the forest floor, which ranged from 3.42 to 7.13 g CO2•m−2•day−1. Respiration rates were depressed in late July and August compared with fall and spring owing to the summer drought characteristic of the Pacific Northwest. Large-diameter western hemlock logs in decay class 1–2 had higher respiration rates than small-diameter logs, whereas large-diameter decay class 3 western hemlock logs had lower respiration rates than small-diameter logs.

scholarly journals Bryophyte Conservation in Managed Boreal Landscapes: Fourteen-Year Impacts of Partial Cuts on Epixylic Bryophytes

2021 ◽  
Vol 4 ◽  
Author(s):  
Jeffrey Opoku-Nyame ◽  
Alain Leduc ◽  
Nicole J. Fenton
Keyword(s):  
Species Composition ◽  
Coarse Woody Debris ◽  
Black Spruce ◽  
Woody Debris ◽  
Permanent Plots ◽  
Old Growth ◽  
Clear Cut ◽  
Old Growth Forest ◽  
Old Growth Forests ◽  
Over Time

Clear cut harvest simplifies and eliminates old growth forest structure, negatively impacting biodiversity. Partial cut harvest has been hypothesized (1) to have less impact on biodiversity than clear cut harvest, and (2) to encourage old growth forest structures. Long-term studies are required to test this hypothesis as most studies are conducted soon after harvest. Using epixylic bryophytes as indicators, this study addresses this knowledge gap. Fourteen years after harvest, we examined changes in epixylic bryophyte community composition richness and traits, and their microhabitats (coarse woody debris characteristics and microclimate) along an unharvested, partial cuts and clear cuts harvest treatment in 30 permanent plots established in the boreal black spruce (Picea mariana) forests of northwestern Quebec, Canada. Our results were compared to those of an initial post-harvest study (year 5) and to a chronosequence of old growth forests to examine species changes over time and the similarity of bryophyte communities in partial cut and old growth forests. Coarse woody debris (CWD) volume by decay class varied among harvest treatments with partial cuts and clear cuts recording lower volumes of early decay CWD. The epixylic community was richer in partial cuts than in mature unharvested forests and clear cuts. In addition, species richness and overall abundance doubled in partial and clear cuts between years 5 and 14. Species composition also differed among treatments between years 5 and 14. Furthermore, conditions in partial cut stands supported small, drought sensitive, and old growth confined species that are threatened by conditions in clear cut stands. Lastly, over time, species composition in partial cuts became more similar to old growth forests. Partial cuts reduced harvest impacts by continuing to provide favorable microhabitat conditions that support epixylic bryophytes. Also, partial cut harvest has the potential to encourage old growth species assemblages, which has been a major concern for biodiversity conservation in managed forest landscapes. Our findings support the promotion of partial cut harvest as an effective strategy to achieve species and habitat conservation goals.

scholarly journals The importance of coarse woody debris in dynamic phases exposure in the beech (Fagus orientalis L.) stands of Hyrcanian forests

2019 ◽  
Vol 65 (No. 10) ◽  
pp. 408-422
Author(s):  
Vahid Etemad ◽  
Mohsen Javanmiri pour ◽  
Zeinab Foladi
Keyword(s):  
Coarse Woody Debris ◽  
Dead Wood ◽  
Woody Debris ◽  
Average Frequency ◽  
Permanent Plots ◽  
Evolutionary Stage ◽  
Gap Formation ◽  
Dead Trees ◽  
Significant Difference ◽  
Natural Stands

In a natural forest, phases of different dynamics are gradually replaced to create sustainability in the stands. Coarse woody debris is among the most significant structural elements of natural stands that perform an influential position in the identification of dynamic phases. Therefore, the focus of this study is on dead wood conditioning as one of the major structural components in determining the various dynamic phases in the northern forests of Iran as part of the temperate forests. For this study, compartment 326 of Gorazbon District was considered as one of the control parcels of Kheyroud Forest. In this parcel, 25 one-hectare sample plots were selected as permanent plots for a long-term forest structure and succession studies. The coarse woody debris by 100% sampling method was measured. The results showed that there are 8 main phases in this area (gap formation, understorey initiation, stem exclusion, volume accumulation, volume degradation, multiple, lighting, old-growth). The extensive forest area (52%) is located in the understorey initiation and stem exclusion phases. The results also showed that the total average volume of snags and logs was 41.5 m<sup>3</sup>·ha<sup>–1</sup>. Furthermore, the mean dead wood volume in decay classes 1, 2, 3 and 4 was 10.33, 12.22, 9.15 and 83.9 m<sup>3</sup>·ha<sup>–1</sup>, respectively. The average frequency of dead trees in the diameter classes smaller than 25 cm, 25–50 cm and in the diameter class more than 50 cm is 25.79, 6.93, and 4.88. The significance analysis results obtained by ANOVA test showed that there is a significant difference between volume, snag and log stock and the shape of dead wood in various dynamic phases. Therefore, in general, dead wood in the forest differs according to habitat, evolutionary stage (dynamic phases), standing volume and species diversity of the tree species.

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