Litter production and nutrient resorption in western red cedar and western hemlock forests on northern Vancouver Island, British Columbia

1995 ◽  
Vol 25 (11) ◽  
pp. 1850-1857 ◽  
Author(s):  
Rodney J. Keenan ◽  
Cindy E. Prescott ◽  
J.P. (Hamish) Kimmins
Keyword(s):  
Nutrient Availability ◽  
Forest Floor ◽  
Forest Type ◽  
Vancouver Island ◽  
The Other ◽  
Western Hemlock ◽  
Litter Fall ◽  
Red Cedar ◽  
N Concentration ◽  
Nutrient Use

Fine litter fall and concentrations of N and P in green foliage and foliar litter were measured in three species over 1 year in two forest types at three sites on northern Vancouver Island to explore the hypothesis that differences in nutrient use and cycling between the dominant tree species on each forest type contribute to differences in forest floor nutrient availability. Total annual aboveground fine litter fall was significantly higher in second-growth, windstorm-derived 85-year-old stands of western hemlock (Tsugaheterophylla (Raf.) Sarg.) and amabilis fir (Abiesamabilis (Dougl.) Forbes) forests (4137 kg•ha−1) than in adjacent old-growth forests of western red cedar (Thujaplicata Donn) and western hemlock (3094 kg•ha−1) occurring on similar sites. Cedar had significantly lower N concentration in green foliage (9.3 mg•g−1) and litter (4.3 mg•g−1) than the other species in each forest type. Hemlock had a higher litter N concentration in the hemlock–amabilis fir type (8.3 mg•g−1) than in the cedar–hemlock type (6.4 mg•g−1). Cedar resorbed a significantly higher percentage of N during leaf senescence (76%), than hemlock in the cedar–hemlock type (64%), hemlock in the hemlock–amabilis fir type (51%), or amabilis fir (18%). Nitrogen-use efficiency (litter-fall mass/litter N) was considerably higher in cedar (235 kg litter/kg N) than in the other species in either forest type (90–156 kg litter/kg N). These results suggest that differences within and between species in the two types in nutrient use and the amount of nutrients cycling through the litter fall and internal redistribution pathways are contributing to lower rates of nutrient cycling and forest floor nutrient availability in the cedar–hemlock type.

Mass and nutrient content of woody debris and forest floor in western red cedar and western hemlock forests on northern Vancouver Island

1993 ◽  
Vol 23 (6) ◽  
pp. 1052-1059 ◽  
Author(s):  
Rodney J. Keenan ◽  
Cindy E. Prescott ◽  
J.P. Hamish Kimmins
Keyword(s):  
Forest Floor ◽  
Forest Type ◽  
Woody Debris ◽  
Nutrient Content ◽  
Vancouver Island ◽  
Western Hemlock ◽  
N Availability ◽  
Red Cedar ◽  
The Mean ◽  
Western Red Cedar

Biomass and C, N, P, and K contents of woody debris and the forest floor were surveyed in adjacent stands of old-growth western red cedar (Thujaplicata Donn)–western hemlock (Tsugaheterophylla (Raf.) Sarg.) (CH type), and 85-year-old, windstorm-derived, second-growth western hemlock–amabilis fir (Abiesamabilis (Dougl.) Forbes) (HA type) at three sites on northern Vancouver Island. Carbon concentrations were relatively constant across all detrital categories (mean = 556.8 mg/g); concentrations of N and P generally increased, and K generally decreased, with increasing degree of decomposition. The mean mass of woody debris was 363 Mg/ha in the CH and 226 Mg/ha in the HA type. The mean forest floor mass was 280 Mg/ha in the CH and 211 Mg/ha in the HA stands. Approximately 60% of the forest floor mass in each forest type was decaying wood. Dead woody material above and within the forest floor represented a significant store of biomass and nutrients in both forest types, containing 82% of the aboveground detrital biomass, 51–59% of the N, and 58–61% of the detrital P. Forest floors in the CH and HA types contained similar total quantities of N, suggesting that the lower N availability in CH forests is not caused by greater immobilization in detritus. The large accumulation of forest floor and woody debris in this region is attributed to slow decomposition in the cool, wet climate, high rates of detrital input following windstorms, and the large size and decay resistance of western red cedar boles.

Availability of N and P in the forest floors of adjacent stands of western red cedar–western hemlock and western hemlock–amabilis fir on northern Vancouver Island

1993 ◽  
Vol 23 (4) ◽  
pp. 605-610 ◽  
Author(s):  
C. E. Prescott ◽  
M.A. McDonald ◽  
G.F. Weetman
Keyword(s):  
Forest Floor ◽  
Forest Type ◽  
Western Hemlock ◽  
Red Cedar ◽  
Floor Layer ◽  
Second Growth ◽  
Extractable P ◽  
Western Red Cedar ◽  
Forest Floors ◽  
Floor Material

Availability of N and P was compared in the forest floors of old-growth forests of western red cedar (Thujaplicata Donn)and western hemlock (Tsugaheterophylla (Raf.) Sarg.) (CH forests), and second-growth forests of western hemlock and amabilis fir (Abiesamabilis (Dougl.) Forbes) (HA forests) of windthrow origin. Five samples of each forest floor layer (litter, fermentation (woody and nonwoody), and humus (woody and nonwoody)) were collected from three forests of each type (CH and HA). All layers of CH forest floors had smaller concentrations of total and extractable N and mineralized less N during 40-day aerobic incubations in the laboratory. Total and extractable P was lower in the litter layer of CH forest floors. Seedlings of western red cedar, Sitka spruce (Piceasitchensis (Bong.) Carr.), western hemlock, and amabilis fir grown from seed in forest floor material from CH forests grew more slowly and took up less N and P than did seedlings grown in HA forest floor material. The low supply of N and P in CH forest floors may contribute to the nutrient supply problems encountered by regenerating trees on cutovers of this forest type.

Responses of western hemlock, Pacific silver fir, and western red cedar plantations on northern Vancouver Island to applications of sewage sludge and inorganic fertilizer

1993 ◽  
Vol 23 (9) ◽  
pp. 1815-1820 ◽  
Author(s):  
G.F. Weetman ◽  
M.A. McDonald ◽  
C.E. Prescott ◽  
J.P. Kimmins
Keyword(s):  
Sewage Sludge ◽  
Vancouver Island ◽  
Western Hemlock ◽  
Municipal Sewage ◽  
Municipal Sewage Sludge ◽  
Phosphorus Fertilization ◽  
Silver Fir ◽  
Nitrogen And Phosphorus ◽  
Red Cedar ◽  
Western Red Cedar

A field experiment was designed to determine whether or not municipal sewage sludge would be effective for fertilization for chlorotic and checked plantations of western hemlock (Tsugaheterophylla (Raf.) Sarg.), Pacific silver fir (Abiesamabilis (Dougl.) Forbes), and western red cedar (Thujaplicata Donn ex. D. Don) already shown to be responsive to conventional nitrogen and phosphorus fertilization. Sewage sludge was applied at an estimated rate of 500 kg N/ha and 133 kg P/ha and ammonium nitrate and triple superphosphate were applied at 225 kg N/ha and 75 kg P/ha to plots planted 8 years earlier on a cutover of old-growth cedar–hemlock forest. Current-year leader growth and foliar vector analyses showed that the trees responded to both treatments during the first growing season with a doubling or tripling of growth rates and improved nutrition. The apparent problem of insufficient sulphur following nitrogen and phosphorus fertilization was not seen in trees treated with sludge. There were no apparent problems in micronutrient supply in these plantations. The extensive area of checked plantations on northern Vancouver Island present an opportunity for the disposal of sewage sludge.

Root Grafts and their Silvicultural Implications

1972 ◽  
Vol 2 (2) ◽  
pp. 111-120 ◽  
Author(s):  
S. Eis
Keyword(s):  
Root System ◽  
Vancouver Island ◽  
Douglas Fir ◽  
Western Hemlock ◽  
Slow Recovery ◽  
Red Cedar ◽  
Western Red Cedar ◽  
Root Grafts

Many trees in stands of Douglas fir, western hemlock and western red cedar on Vancouver Island were joined by functional grafts. In a partially cut stand, 45% of the stumps showed evidence of continued growth and half of these (23%) were still growing vigorously more than 22 years after logging. On experimentally detopped trees, growth extended several meters up the bole. Dominant trees usually supported the growth of the root system and lower boles of grafted suppressed trees.Translocation through grafts may partially explain the frequent stagnation and slow recovery of stands after thinning from above, and may be involved in the usually rapid increase of growth after thinning from below. It is probably a contributing factor in establishing dominance and determining mortality in overtopped trees. In species that graft freely, the use of silvicides in spacing and thinning treatments should be restricted to young stands before grafts are established.

Comparison of the chemical properties of forest floors, decaying wood, and fine roots in three ecosystems on Vancouver Island

1981 ◽  
Vol 11 (2) ◽  
pp. 216-218 ◽  
Author(s):  
H. J. Quesnel ◽  
L M. Lavkulich
Keyword(s):  
Forest Floor ◽  
Fine Roots ◽  
Chemical Properties ◽  
Nutrient Status ◽  
Vancouver Island ◽  
Loss On Ignition ◽  
Elemental Concentrations ◽  
Red Cedar ◽  
Forest Floors ◽  
Decaying Wood

Elemental concentrations were measured and compared for LF horizons, H horizons, decaying wood, and fine (< 2 mm) roots of three ecosystems on northern Vancouver Island. The principal tree species of these ecosystems were western hemlock (Tsugaheterophylla (Raf.) Sarg.), amabilis fir (Abiesamabilis (Dougl.) Forbes), and western red cedar (Thujaplicata Donn.). The H horizons had greater Mg and Na values than the LF horizons, while the opposite result was found for K and loss on ignition (LOI). The decaying wood represents a significant accumulation of nutrient-deficient biomass that could immobilize N. The decomposing fine roots will temporarily immobilize N while possibly increasing the concentration of elements such as Fe, Al, and Mn. These materials should be separated from forest floor samples in order to represent more accurately the nutrient status of forest floor horizons.

Photosynthetic photon flux density, red:far-red ratio, and minimum light requirement for survival of Gaultheriashallon in western red cedar–western hemlock stands in coastal British Columbia

1989 ◽  
Vol 19 (11) ◽  
pp. 1470-1477 ◽  
Author(s):  
Christian Messier ◽  
Terry W. Honer ◽  
James P. Kimmins
Keyword(s):  
Forest Floor ◽  
Photosynthetic Photon Flux Density ◽  
Western Hemlock ◽  
Photon Flux ◽  
Photon Flux Density ◽  
Photosynthetic Photon Flux ◽  
Light Requirement ◽  
Red Cedar ◽  
Sky Conditions ◽  
Overcast Sky

Measurements of photosynthetic photon flux density (PPFD; global, diffuse, and direct: 400–700 nm) and red:far-red ratio (660:730 nm) were made above and below an understory of salal (Gaultheriashallon Pursh) within three western red cedar (Thujaplicata Donn)–western hemlock (Tsugaheterophylla (Raf.) Sarg.) stands on the northern end of Vancouver Island, British Columbia. These stands were chosen to represent a wide range of canopy closure and understory salal cover. The measurements were repeated under both clear and overcast sky conditions. Within each stand, the percent cover, height, number of stems per hectare, and leaf morphology of the salal were measured. As expected, there was a decrease in both the salal cover and in the global PPFD and red:far-red ratio measured directly above salal as the percent tree cover increased. Salal leaf thickness and specific leaf weight decreased, whereas its leaf area increased, as global PPFD and red:far-red ratio decreased. The results indicate that the minimum light requirement for salal survival is between 3.9 and 11.1 μmol•m−2•s−1 (1.2 to 3.3% of global PPFD measured in an adjacent clearing) and 4.5 and 27.2 μmol•m−2•s−1 (0.3 to 1.8% of global PPFD measured in the clearing) under overcast and clear sky conditions, respectively. The global PPFD and red:far-red ratio measured 30 cm above the forest floor (beneath the salal) did not vary widely among the three stands. At this level, global PPFD values were very low, ranging from 0.15 to 0.35% and 0.58 to 0.75% of the global PPFD measured in the clearing under clear and overcast sky conditions, respectively. Red:far-red ratios at the forest floor level varied from 0.13 to 0.25 and 0.40 to 0.58 under clear and overcast sky conditions, respectively. The silvicultural implications of the results are discussed.

scholarly journals Study Productivity And Decomposition Litterfall In Sibolangit Forest, Deli Serdang To Support Field Trip Plantation Ecology

2016 ◽  
Vol 1 (3) ◽  
pp. 57
Author(s):  
Tonggo Sinaga
Keyword(s):  
Soil Moisture ◽  
Forest Ecosystem ◽  
Rain Forest ◽  
Forest Floor ◽  
The Other ◽  
Litter Fall ◽  
Productivity Decomposition ◽  
Nutrients Cycling ◽  
Fallen Leaves ◽  
High Topography

The research was conducted in mountain rain forest of Sibolangit, Deli Serdang, Northen Sumatera. The objectives of this research are to know litter fall productivity, decomposition and litterfall nutrients. Litterfall is the fallen leaves, twigs, flowers and fruits to forest-floor, which one of the input of nutrients to anorganic sril. This input is one of the nutrients cycling to forest ecosystem and the other form atmosfer and a lot of animals. Based on available data on 20 traps, litterfall productivity 5,91 gr/m2/week or 3073,2 kg/ha/year in high topography. Litterfall productivity 4,92gr/m2/week or 2558,4 kg/ha/year in low topography. Litterfall decomposition (lossed of weight) about 25,48% in 30 days ( 1 month ) for meranti ( Shorea sp ) and leaves of Ficus sp had lossed of weight about 8,09% in 30 days ( 1 month ). Ration C/N (Carbon-Nitrogen) litterfall and soil moisture made differrent litterfall decomposition (lossed of weight)   Keywords : Productivity, Decomposition, Litterfall, Productivity.

Nitrogen mineralization and decomposition in forest floors in adjacent plantations of western red cedar, western hemlock, and Douglas-fir

1994 ◽  
Vol 24 (12) ◽  
pp. 2424-2431 ◽  
Author(s):  
C.E. Prescott ◽  
C.M. Preston
Keyword(s):  
Mass Loss ◽  
N Mineralization ◽  
Forest Floor ◽  
Douglas Fir ◽  
Western Hemlock ◽  
N Availability ◽  
Net N Mineralization ◽  
Red Cedar ◽  
Western Red Cedar ◽  
Forest Floors

To determine if western red cedar (Thujaplicata Donn) litter contributes to low N availability in cedar–hemlock forests, we measured concentrations of N and rates of net N mineralization in forest floors from single-species plantations of cedar, western hemlock (Tsugaheterophylla (Raf.) Sarg.), and Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) on the same site in coastal British Columbia. Concentrations of total and extractable N and rates of net N mineralization during laboratory incubations were lowest in the cedar forest floor and highest in Douglas-fir. Less C was mineralized in the cedar forest floor during incubation, and the amount of N mineralized per unit C was least in cedar. Rates of mass loss of foliar litter of the three species were similar during the first 50 weeks of a 70-week laboratory incubation, but cedar lost mass more quickly during the final 20 weeks. Rates of net N mineralization in the forest floors were significantly correlated with the initial percent N, C/N, % Klason lignin, and lignin/N of foliar litter. Foliar litter of cedar had lower concentrations of N and greater proportions of alkyl C (based on 13C NMR spectroscopy) than Douglas-fir litter. These characteristics of cedar litter may contribute to low N availability in cedar–hemlock forest floors. Concentrations of alkyl C (waxes and cutin) may be better than lignin for predicting rates of mass loss and N mineralization from litter.

Vertical fine root distributions of western redcedar, western hemlock, and salal in old-growth cedar–hemlock forests on northern Vancouver Island

2002 ◽  
Vol 32 (7) ◽  
pp. 1208-1216 ◽  
Author(s):  
Jennifer N Bennett ◽  
Ben Andrew ◽  
Cindy E Prescott
Keyword(s):  
Forest Floor ◽  
Fine Roots ◽  
Fine Root ◽  
Mineral Soil ◽  
Vancouver Island ◽  
Western Hemlock ◽  
Old Growth ◽  
Total N ◽  
Western Redcedar ◽  
Vertical Distributions

The vertical distributions of fine roots of western hemlock (Tsuga heterophylla (Raf.) Sarg.) western redcedar (Thuja plicata Donn ex D. Don), and salal (Gaultheria shallon Pursh) were characterized in old-growth cedar–hemlock forests on northern Vancouver Island. Total biomasses of cedar, hemlock, and salal roots in the forest floor and upper mineral soil were 817, 620, and 187 g·m–2, respectively. Hemlock and salal fine roots were concentrated in the upper forest floor, while cedar fine roots were evenly distributed through the profile. Salal and hemlock fine root densities (g·m–3) in the forest floor and mineral soil were positively correlated, as were salal and cedar root biomass distributions (g·m–2). Only salal and hemlock root densities were significantly correlated with N concentrations. Hemlock root densities were negatively correlated with total N, and salal root densities were negatively correlated with total N and soluble organic N. Based on fine root densities, hemlock and salal probably compete for resources in the upper forest floor, whereas cedar accesses resources in the lower organic and mineral soil horizons. The differences in the vertical distributions of cedar, hemlock, and salal fine roots may partly explain the co-occurrence and different productivities of the three species in cedar-hemlock forests.

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