Dynamics of dead wood in old-growth hemlock–hardwood forests of northern Wisconsin and northern Michigan

1994 ◽  
Vol 24 (8) ◽  
pp. 1672-1683 ◽  
Author(s):  
Lucy E. Tyrrell ◽  
Thomas R. Crow
Keyword(s):  
Tree Mortality ◽  
Structural Integrity ◽  
Dead Wood ◽  
Woody Debris ◽  
Basal Area ◽  
Exponential Model ◽  
Decay Rates ◽  
Permanent Plots ◽  
Old Growth ◽  
Northern Michigan

We studied the dynamics of coarse woody debris (logs and snags) in old-growth forests by estimating rates of tree mortality, snag change, and log decay in hemlock–hardwood stands located in northern Wisconsin and northern Michigan. To estimate mortality and snag changes, we recensused live trees and dead snags in permanent plots in 15 stands. We also recorded recent mortality along transects, and noted category of mortality (standing death, breakage, or uprooting) for gap-maker trees and logs in 25 stands. Decay rates were estimated from a simple exponential model of wood density from log sections against the age of the wood since tree death, and from ages of trees growing on decaying "nurse" logs. From data obtained in permanent plots, annual tree mortality averaged 0.9% original basal area, 0.9% original live tree density, and 4.8 trees/ha. Of the three categories of tree mortality, standing death accounted for 62% of all mortality, while breakage represented 25%, and uprooting, 13%. Based on origin of logs, species composition affected category of mortality. Eastern hemlock (Tsugacanadensis (L.) Carr.) was significantly more prone to uprooting while yellow birch (Betulaalleghaniensis Britton) and paper birch (Betulapapyrifera Marsh.) were less prone to uprooting than expected if independence of species and categories of mortality were assumed (χ2 = 216.5, df = 14, p < 0.001). Changes in snags (fragmentation to shorter snags or collapse at the base) occurred for <7% of the original snags annually. Fragmentation occurred for 2.1 snags/(ha × year), and collapse for 1.3 snags/(ha × year). We estimated that it takes nearly 200 years for hemlock logs to lose structural integrity and become partially incorporated into the soil. At >350 years, the two oldest hemlock–hardwood stands had accumulated volumes of logs >65 m3/ha distributed among all decay classes, and appeared to be at a dead wood equilibrium in which rates of log production from mortality balance rates of wood loss by decay.

Input and decay of coarse woody debris in coniferous stands in western Oregon and Washington

1982 ◽  
Vol 12 (1) ◽  
pp. 18-28 ◽  
Author(s):  
Phillip Sollins
Keyword(s):  
Tree Mortality ◽  
Large Diameter ◽  
Mineral Soil ◽  
Small Diameter ◽  
Decay Rates ◽  
Root Turnover ◽  
Permanent Plots ◽  
Old Growth ◽  
Salvage Logging ◽  
Standing Dead

At 10 locations in Oregon and Washington, tree mortality resulted in dry-matter transfer of 1.5–4.5 Mg•ha−1•year−1 of boles and branches to the forest floor and 0.3–1.3 Mg•ha−1•year−1 of large-diameter roots directly to the mineral soil. The first value is about the same as that reported for leaf fall in similar stands; the second value generally is smaller than that reported for fine root turnover. Results are based on measurements by the U.S. Forest Service spanning 16–46 years and areas as large as 42 ha. Values based on intervals < 10 years were highly variable and potentially misleading.At an old-growth Douglas-fir stand in Washington, fallen boles accounted for 81 Mg/ha, standing dead for 54 Mg/ha. Density of fallen boles averaged from 0.14 to 0.27 g/cm3 depending on decay state. Values were lower than some previously reported because (1) our sample included small-diameter fallen boles that tend to decay rapidly, and (2) we measured density with techniques that minimized compaction and shrinkage.The decay rate at the old-growth stand, calculated indirectly by dividing bole mortality (megagrams per hectare per year) by the amount (megagrams per hectare) of fallen and standing dead woody material, was 0.028 year−1. This rate, three to five times those previously calculated directly from change in density alone, was almost identical to values calculated elsewhere from change in both volume and density. Decay rates based on change in density alone include only respired and leached material and exclude the large amount of material lost in fragmentation. This study shows the value of permanent plots, undisturbed by salvage logging, for retrospective studies of decomposition, nutrient cycling, and productivity.

Coarse woody debris in old-growth temperate beech (Nothofagus) forests of New Zealand

1994 ◽  
Vol 24 (10) ◽  
pp. 1989-1996 ◽  
Author(s):  
Glenn H. Stewart ◽  
Larry E. Burrows
Keyword(s):  
New Zealand ◽  
Coarse Woody Debris ◽  
Tree Mortality ◽  
Structural Integrity ◽  
Woody Debris ◽  
Hardwood Forests ◽  
Old Growth ◽  
Carbon And Nitrogen ◽  
Decay Sequence ◽  
Nothofagus Forests

The volume, biomass, and carbon and nitrogen content of coarse woody debris were measured on three 1-ha reference plots in old-growth Nothofagusfusca (Hook. f.) Oerst.–Nothofagusmenziesii (Hook. f.) Oerst. forest on the South Island of New Zealand. Two decay sequences for logs and one for standing dead trees (snags) were recognised from two-way indicator species analysis (TWINSPAN) of up to 30 variables related to physical characteristics and structural integrity. Wood volume (up to 800 m3•ha−1) and biomass were high (up to 300 Mg•ha−1), and the inside-out decay sequence from heartwood to sapwood was unusual compared with that of other temperate hardwood forests. Coarse woody debris represented significant carbon and nitrogen pools, with ca. 150 Mg•ha−1 and 370 kg•ha−1, respectively, in one stand. The coarse woody debris component of these broad-leaved evergreen hardwood forests was much higher than that reported for other temperate hardwood forests and approaches that of many northern hemisphere conifer forests. The large coarse woody debris pools are discussed in relation to live stand biomass, natural disturbances and tree mortality, and decomposition processes.

Selected Nova Scotia old-growth forests: Age, ecology, structure, scoring

2003 ◽  
Vol 79 (3) ◽  
pp. 632-644 ◽  
Author(s):  
Bruce J Stewart ◽  
Peter D Neily ◽  
Eugene J Quigley ◽  
Lawrence K Benjamin
Keyword(s):  
Age Structure ◽  
Coarse Woody Debris ◽  
Sugar Maple ◽  
Dead Wood ◽  
Woody Debris ◽  
Large Diameter ◽  
Basal Area ◽  
Red Spruce ◽  
White Pine ◽  
Old Growth

A study of four old-growth stands in Nova Scotia was conducted to document the ecological characteristics of these currently rare Acadian forest ecosystems. Stands were selected to represent the two dominant climax forest types, hemlock–red spruce–eastern white pine, and sugar maple–yellow birch–beech. Data include measurements of age structure, species composition, diameter distribution, basal area, height, coarse woody debris, snags, vertical structure, and canopy condition. All stands were determined to be uneven-aged. Old-growth reference ages calculated for the stands ranged from 164 to 214 years. All stands displayed broad diameter distributions that had peak basal area representation in the 40- to 50-cm diameter classes. Volumes of dead wood ranged from 111 to 148 m3/ha in the softwood stands and from 63 to 83m3/ha in the hardwood stands. Dead wood consisted of approximately one-third snags and two thirds downed coarse woody debris. Measurements from the stands were used to evaluate Nova Scotia's recently developed Old Forest Scoring System. Six stand attributes were rated for a maximum score of 100: stand age, primal value, number of large-diameter trees, length of large-diameter dead wood, canopy structure, and understorey structure. Based on the age attribute, three of the four stands were classed as Mature Old Growth and one was very close, indicating that all are in the shifting mosaic stage of late forest succession. The scores for all stands were relatively high, ranging from 75 to 85, as would be expected from some of the best old-growth stands in the province. Key words: old growth, climax, primal, late succession, uneven-aged, scoring, coarse woody debris, age structure, diameter, Acadian forest, northern hardwood, red spruce, eastern hemlock, white pine, sugar maple, yellow birch, American beech

Relating jack pine budworm damage to stand inventory variables in northern Michigan

1996 ◽  
Vol 26 (12) ◽  
pp. 2180-2190 ◽  
Author(s):  
Deborah G. McCullough ◽  
Lyle J. Buss ◽  
Larry D. Marshall ◽  
Jari Kouki
Keyword(s):  
Tree Mortality ◽  
Basal Area ◽  
Site Index ◽  
The United States ◽  
Jack Pine ◽  
Inventory Data ◽  
Upper Peninsula ◽  
Jack Pine Budworm ◽  
Lake States ◽  
Northern Michigan

Stand-level mortality and top kill from a 1991–1993 jack pine budworm (Choristoneurapinuspinus Freeman) outbreak were surveyed annually in the Raco Plains area of the Hiawatha National Forest in Michigan's Upper Peninsula from 1992 to 1994. Defoliation was visually estimated and percentage of trees killed or top killed was determined in 104 stands. In 1994, tree mortality attributable to the outbreak averaged 8% and 17% of trees had dead tops. Current stand inventory data, including age, site index, basal area, and size, were acquired from the Hiawatha Forest. Stands were grouped on the basis of inventory variables used for jack pine management in the Lake States region of the United States. Differences in tree mortality and top kill between groups, and associations between tree mortality and inventory variables, were evaluated. Tree mortality was greater in overmature stands and in overstocked stands, but stand size had little effect. Contrary to expectations, mortality was lower on poor sites with low site index values than on better sites with higher site index values. Mortality was not related to abundance of open-grown, full-canopied wolf trees or to abundance of trees infected with pine gall rust (Endocronartiumharknessii (J.P. Moore) Y. Hiratsuka (=Peridermiumharknessii J.P. Moore)). Amount of top kill was related to defoliation severity and was higher in overmature and understocked stands. Top kill was not strongly associated with amount of tree mortality or with inventory variables.

Overstory litter inputs and nutrient returns in an old-growth temperate forest ecosystem, Olympic National Park, Washington

2002 ◽  
Vol 32 (4) ◽  
pp. 742-750 ◽  
Author(s):  
Robert L Edmonds ◽  
Georgia LD Murray
Keyword(s):  
National Park ◽  
Woody Debris ◽  
Basal Area ◽  
Tsuga Heterophylla ◽  
Old Growth ◽  
Olympic National Park ◽  
Shade Tolerant Species ◽  
Woody Litter ◽  
Fine Wood ◽  
Temperate Forest Ecosystem

Overstory litterfall rates and nutrient returns were determined in an old-growth temperate rainforest watershed in the Hoh River valley, Olympic National Park, Washington. Litter was sorted into green needles, senescent needles, fine wood, reproductive, and miscellaneous litter (mostly arboreal lichens and mosses). Understory and coarse woody debris inputs were not determined. Total annual overstory litterfall averaged 3594 kg·ha–1 and varied among the six plant communities in the watershed. There was a trend for litterfall to be higher in the upper watershed; elevations ranged from 180 to 850 m. Needles provided the greatest amount of litterfall (60%) with woody litter and other material averaging 18 and 22%, respectively. Highest senescent needle litterfall occurred from July to October, but highest woody litterfall was from January to April. Green needles provided only 3% of annual needle litterfall. Green and senescent needle litterfall were related to western hemlock (Tsuga heterophylla (Raf.) Sarg.) basal area, suggesting that this shade-tolerant species was the greatest contributor to needle litterfall. The following quantities (kg·ha–1) of nutrients were returned to the forest floor annually: Ca, 26.8; N, 24.6; K, 4.0; Mg, 3.0; P, 2.9; Mn, 1.7; and Na, 1.2.

scholarly journals Acumulación De Restos Lignino-Celulósicos (Necromasa) En Un Bosque Templado De Antiguo Crecimiento Dominado Por Nothofagus En El Centro- Sur De Chile

2016 ◽  
Vol 12 (9) ◽  
pp. 69
Author(s):  
Carlos Belezaca Pinargote ◽  
Darwin Salvatierra Piloso ◽  
Diana Delgado Campusano ◽  
Roberto Godoy Bórquez ◽  
Eduardo Valenzuela Flores ◽  
...  
Keyword(s):  
Coarse Woody Debris ◽  
National Park ◽  
Woody Debris ◽  
Basal Area ◽  
Stem Volume ◽  
Decomposition Level ◽  
Old Growth ◽  
Central Chile ◽  
Forest Density ◽  
South Central

Coarse woody debris (CWD) are the necromass in wooded environments and comply with various ecosystems functions, such as seedling nursery, habitat other organisms, store carbon (C) and nutrients, etc.. Volume, necromass, and decay states of CWD dead in an old-growth temperate (OGTF) forest in Puyehue National Park, South-Central Chile were evaluated. In 10 plots of 900 m2 CWD was quantified (≥ 10 cm diameter), whose necromass classified using a scale of five categories/stages of decay, necromass (1 = lowest and 5 = highest degradation). The average forest density was 299 trees ha-1, 112 m2 ha-1 of basal area, and 2.395 m3 of stem volume. The upper arboreal stratum was dominated by N. betuloides. The greatest amount of CWD belonged to N. betuloides (95,2%), where logs (52,7%) and branches (35%), plant structures were present in greater numbers. The bulk of necromass found in advanced states of decomposition (level 4 and 5) with 56,5% and 34,1%, respectively. It was determined that the CWD volume was 632 m3 ha-1, representing a necromass of 321.5 Mg ha- 1. These results demonstrate that the old-growth temperate forests of southern Chile are significant reserves of coarse woody debris, which contributes to the biogeochemistry of these complex and remote ecosystems.

scholarly journals Bryophyte Conservation in Managed Boreal Landscapes: Fourteen-Year Impacts of Partial Cuts on Epixylic Bryophytes

2021 ◽  
Vol 4 ◽  
Author(s):  
Jeffrey Opoku-Nyame ◽  
Alain Leduc ◽  
Nicole J. Fenton
Keyword(s):  
Species Composition ◽  
Coarse Woody Debris ◽  
Black Spruce ◽  
Woody Debris ◽  
Permanent Plots ◽  
Old Growth ◽  
Clear Cut ◽  
Old Growth Forest ◽  
Old Growth Forests ◽  
Over Time

Clear cut harvest simplifies and eliminates old growth forest structure, negatively impacting biodiversity. Partial cut harvest has been hypothesized (1) to have less impact on biodiversity than clear cut harvest, and (2) to encourage old growth forest structures. Long-term studies are required to test this hypothesis as most studies are conducted soon after harvest. Using epixylic bryophytes as indicators, this study addresses this knowledge gap. Fourteen years after harvest, we examined changes in epixylic bryophyte community composition richness and traits, and their microhabitats (coarse woody debris characteristics and microclimate) along an unharvested, partial cuts and clear cuts harvest treatment in 30 permanent plots established in the boreal black spruce (Picea mariana) forests of northwestern Quebec, Canada. Our results were compared to those of an initial post-harvest study (year 5) and to a chronosequence of old growth forests to examine species changes over time and the similarity of bryophyte communities in partial cut and old growth forests. Coarse woody debris (CWD) volume by decay class varied among harvest treatments with partial cuts and clear cuts recording lower volumes of early decay CWD. The epixylic community was richer in partial cuts than in mature unharvested forests and clear cuts. In addition, species richness and overall abundance doubled in partial and clear cuts between years 5 and 14. Species composition also differed among treatments between years 5 and 14. Furthermore, conditions in partial cut stands supported small, drought sensitive, and old growth confined species that are threatened by conditions in clear cut stands. Lastly, over time, species composition in partial cuts became more similar to old growth forests. Partial cuts reduced harvest impacts by continuing to provide favorable microhabitat conditions that support epixylic bryophytes. Also, partial cut harvest has the potential to encourage old growth species assemblages, which has been a major concern for biodiversity conservation in managed forest landscapes. Our findings support the promotion of partial cut harvest as an effective strategy to achieve species and habitat conservation goals.

scholarly journals The importance of coarse woody debris in dynamic phases exposure in the beech (Fagus orientalis L.) stands of Hyrcanian forests

2019 ◽  
Vol 65 (No. 10) ◽  
pp. 408-422
Author(s):  
Vahid Etemad ◽  
Mohsen Javanmiri pour ◽  
Zeinab Foladi
Keyword(s):  
Coarse Woody Debris ◽  
Dead Wood ◽  
Woody Debris ◽  
Average Frequency ◽  
Permanent Plots ◽  
Evolutionary Stage ◽  
Gap Formation ◽  
Dead Trees ◽  
Significant Difference ◽  
Natural Stands

In a natural forest, phases of different dynamics are gradually replaced to create sustainability in the stands. Coarse woody debris is among the most significant structural elements of natural stands that perform an influential position in the identification of dynamic phases. Therefore, the focus of this study is on dead wood conditioning as one of the major structural components in determining the various dynamic phases in the northern forests of Iran as part of the temperate forests. For this study, compartment 326 of Gorazbon District was considered as one of the control parcels of Kheyroud Forest. In this parcel, 25 one-hectare sample plots were selected as permanent plots for a long-term forest structure and succession studies. The coarse woody debris by 100% sampling method was measured. The results showed that there are 8 main phases in this area (gap formation, understorey initiation, stem exclusion, volume accumulation, volume degradation, multiple, lighting, old-growth). The extensive forest area (52%) is located in the understorey initiation and stem exclusion phases. The results also showed that the total average volume of snags and logs was 41.5 m<sup>3</sup>·ha<sup>–1</sup>. Furthermore, the mean dead wood volume in decay classes 1, 2, 3 and 4 was 10.33, 12.22, 9.15 and 83.9 m<sup>3</sup>·ha<sup>–1</sup>, respectively. The average frequency of dead trees in the diameter classes smaller than 25 cm, 25–50 cm and in the diameter class more than 50 cm is 25.79, 6.93, and 4.88. The significance analysis results obtained by ANOVA test showed that there is a significant difference between volume, snag and log stock and the shape of dead wood in various dynamic phases. Therefore, in general, dead wood in the forest differs according to habitat, evolutionary stage (dynamic phases), standing volume and species diversity of the tree species.

Stream channel configuration, landform, and riparian forest structure in the Cascade Mountains, Washington

2000 ◽  
Vol 57 (4) ◽  
pp. 699-707 ◽  
Author(s):  
Byron W Rot ◽  
Robert J Naiman ◽  
Robert E Bilby
Keyword(s):  
Plant Community ◽  
Riparian Forest ◽  
Woody Debris ◽  
Basal Area ◽  
Old Growth ◽  
Tree Diameter ◽  
Riparian Tree ◽  
Channel Configuration ◽  
Riparian Plant ◽  
Channel Type

The hierarchical relationship of five key elements, valley constraint, riparian landform, riparian plant community, channel type, and channel configuration, are described for 21 sites in mature to old-growth riparian forests of the western Cascades Mountains, Washington, U.S.A. Channel type (bedrock, plane-bed, and forced pool-riffle) was closely related to channel configuration (especially large woody debris (LWD) volume, density, and LWD-formed pools) at the smallest spatial scale and valley constraint at the largest. Valley constraint significantly influenced off-channel habitat (r2 = 0.71) and LWD volume within forced pool-riffle channels (r2 = 0.58). Riparian plant community composition was differentiated by four landform classes: three alluvial landforms based on height above the channel and one based on hillslope. Just above the active channel, floodplain landforms contained more deciduous stems than conifer and greater conifer basal area than deciduous. Conifers dominated other landforms. The diameter of in-channel LWD increased with the age of the riparian forest (r2 = 0.34). In old-growth forests, LWD diameter was equivalent to or greater than the average riparian tree diameter for all sites. In younger forests, the mixed relationship between LWD and riparian tree diameter may reflect a combination of LWD input from the previous old-growth stand and LWD input from the existing stand.

scholarly journals Recovery of Forest Structure Following Large-Scale Windthrows in the Northwestern Amazon

Forests ◽  
2021 ◽  
Vol 12 (6) ◽  
pp. 667
Author(s):  
J. David Urquiza Muñoz ◽  
Daniel Magnabosco Marra ◽  
Robinson I. Negrón-Juarez ◽  
Rodil Tello-Espinoza ◽  
Waldemar Alegría-Muñoz ◽  
...  
Keyword(s):  
Forest Structure ◽  
Large Scale ◽  
Tree Mortality ◽  
Basal Area ◽  
Forest Recovery ◽  
Old Growth ◽  
Study Region ◽  
Central Amazon ◽  
Old Growth Forest ◽  
Study Sites

The dynamics of forest recovery after windthrows (i.e., broken or uprooted trees by wind) are poorly understood in tropical forests. The Northwestern Amazon (NWA) is characterized by a higher occurrence of windthrows, greater rainfall, and higher annual tree mortality rates (~2%) than the Central Amazon (CA). We combined forest inventory data from three sites in the Iquitos region of Peru, with recovery periods spanning 2, 12, and 22 years following windthrow events. Study sites and sampling areas were selected by assessing the windthrow severity using remote sensing. At each site, we recorded all trees with a diameter at breast height (DBH) ≥ 10 cm along transects, capturing the range of windthrow severity from old-growth to highly disturbed (mortality > 60%) forest. Across all damage classes, tree density and basal area recovered to >90% of the old-growth values after 20 years. Aboveground biomass (AGB) in old-growth forest was 380 (±156) Mg ha−1. In extremely disturbed areas, AGB was still reduced to 163 (±68) Mg ha−1 after 2 years and 323 (± 139) Mg ha−1 after 12 years. This recovery rate is ~50% faster than that reported for Central Amazon forests. The faster recovery of forest structure in our study region may be a function of its higher productivity and adaptability to more frequent and severe windthrows. These varying rates of recovery highlight the importance of extreme wind and rainfall on shaping gradients of forest structure in the Amazon, and the different vulnerabilities of these forests to natural disturbances whose severity and frequency are being altered by climate change.

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