Forest turnover and the dynamics of bole wood litter in subalpine balsam fir forest
A chronosequence of three stands of balsam fir was sampled in 1974 and 1982; during these 8 years, recruitment was absent so mortality alone accounted for an 18–30% decrease in live tree density. In a mature 78-year-old stand, the mass of bole wood on the forest floor was 1.4 kg•m−2 compared with an estimated aboveground live and dead bole biomass of 17.2 kg•m−2. During 5 years of repetitive sampling, annual bole input to the forest floor was episodic and variable in time and space, ranging from 3 to 365 g•m−2•year−1. A mass balance model was used to characterize the changes in wood litter on the forest floor. If most of the live trees die within a short period of time, bole input would occur in a pulse event and cause a peak in wood litter mass, which would then decline over time (and with stand maturation) as decomposition prevails. The assumption of steady-state conditions for wood litter is not valid; rather the mass of wood litter will wax and wane through time. Over a landscape, spatial patterns in the abundance of wood litter reflect a stand's history; old mature stands would have little wood litter while young regenerating stands would have large amounts. A maximum value for wood litter would be found in a stand located immediately behind a fir wave. Natural disturbances from wind and avalanches lead to contrasting patterns with high and low wood litter values, respectively. About 41% of forest turnover in the balsam fir zone is initiated from natural disturbance and fir waves.