The effects of experimental fires on black spruce forest floors in interior Alaska

1983 ◽  
Vol 13 (5) ◽  
pp. 879-893 ◽  
Author(s):  
C. T. Dyrness ◽  
Rodney A. Norum
Keyword(s):  
Forest Floor ◽  
Black Spruce ◽  
Mineral Soil ◽  
Available P ◽  
Small Scale ◽  
Total N ◽  
Burned Areas ◽  
Forest Floors ◽  
Surface Mineral ◽  
Total P

Seven units (about 2 ha each) of black spruce – feather moss forest were experimentally burned over a range of fuel moisture conditions during the summer of 1978. Surface woody fuels were sparse and the principal carrier fuel was the forest floor (largely mosses and their decomposition products). Forest floors after burning comprised a small-scale mosaic of unburned, scorched, lightly burned, moderately burned, and heavily burned (organic materials entirely consumed) conditions. Percentage of the unit area in the moderately and heavily burned condition ranged from 11.2 to 77.2% and percent decrease in forest-floor thickness varied from 27.4 to 63.1% in the seven units. Forest-floor consumption was most closely correlated with the moisture content of lower moss (01 horizon) and lower duff layers (022 horizon) at the time of burning. For the first 3 years after fire, biomass production was greater on heavily burned than on lightly burned sites (58 vs. 37 g/m2 on an annual basis). Heavily burned sites were completely dominated by the invading species Epilobiumangustifolium L., Ceratodonpurpureus (Hedw.) Brid., and Marchantiapolymorpha L., whereas lightly burned plots were occupied by sprouting species such as Calamagrostiscanadensis (Michx.) Beauv., Vacciniumuliginosum L., and Ledumgroenlandicum Oeder. Soil pH and amounts of total P and available P in the forest floor increased significantly as a result of burning; and in all cases, increases reached a maximum in moderately and heavily burned areas. Total N in the forest floor increased significantly in moderately burned, but decreased slightly in heavily burned areas. Total N and total P showed smaller increases in the surface mineral soil as a result of burning. Supplies of available P in the mineral soil increased almost 4-fold in moderately burned and over 16-fold in heavily burned areas.

The effect of wildfire on soil chemistry in four forest types in interior Alaska

1989 ◽  
Vol 19 (11) ◽  
pp. 1389-1396 ◽  
Author(s):  
C. T. Dyrness ◽  
K. Van Cleve ◽  
J. D. Levison
Keyword(s):  
Forest Floor ◽  
Soil Chemistry ◽  
Black Spruce ◽  
Mineral Soil ◽  
White Spruce ◽  
Available P ◽  
Ph Effects ◽  
Quaking Aspen ◽  
Forest Types ◽  
Surface Mineral

Soil chemical properties were studied after a wildfire in stands of white spruce (Piceaglauca (Moench) Voss), black spruce (Piceamariana (Mill.) B.S.P.), paper birch (Betulapapyrifera Marsh.), and quaking aspen (Populustremuloides Michx.). Samples of the forest floor and surface 5 cm of mineral soil were collected from burned sites and unburned controls and analyzed soon after the fire. With the exception of soil pH, effects of the fire on soil chemistry differed among the four forest types. Generally, amounts of exchangeable K, Ca, and Mg did not appreciably increase in the forest floor and surface mineral soil except in heavily burned areas in white spruce and black spruce. Fire reduced amounts of N by about 50% in white spruce, aspen, and birch forest floors. In black spruce, quantities of N were slightly higher in heavily burned locations. Forest floor C:N ratios were substantially lower in heavily burned locations in white spruce and black spruce than in unburned controls. Burning did not have a marked influence on supplies of available P in the forest floor, except in heavily burned black spruce, where average amounts were 12.50 g/m2 versus only 0.46 g/m2 in the control. Burning caused more moderate gains in available P in surface mineral soils under aspen and white spruce. We concluded that fire caused marked short-term changes in soil chemistry in the four forest types. How long these changes will persist is unknown.

Litterfall and soil characteristics in canopy gaps occupied by vine maple in a coastal western hemlock forest

1997 ◽  
Vol 77 (4) ◽  
pp. 703-711 ◽  
Author(s):  
Aynslie E. Ogden ◽  
Margaret G. Schmidt
Keyword(s):  
Litter Decomposition ◽  
Forest Floor ◽  
Mineral Soil ◽  
Canopy Gaps ◽  
Plant Interactions ◽  
Total N ◽  
Hardwood Tree ◽  
Forest Floors ◽  
Surface Mineral ◽  
Weak Tendency

In some low-elevation coastal British Columbia forests, canopy gaps can be occupied by the hardwood tree species, vine maple (Acer circinatum). The objective of this study was to determine how vine maple gaps influence litterfall, litter decomposition, and forest floor and mineral soil properties. Measurements were made on six vine maple gaps paired with six conifer canopy plots. Vine maple gaps had significantly less conifer litterfall during the autumn, higher pH, and higher concentrations of Ca, Mg and K in the forest floor, thinner forest floors, and a weak tendency for lower C/N ratios, higher pH values and higher total N concentrations in the surface mineral soil. Vine maple litter was found to decompose significantly faster than conifer litter and to have higher concentrations of N, P, Ca, Mg, K, Fe and Zn. Decomposition rates of vine maple litter and of conifer litter did not differ significantly between vine maple gap and conifer canopy plots. Larger vine maple clones had significantly thicker forest floors with higher concentrations of Ca, and higher N concentrations and lower C/N ratios in the surface mineral soil than gaps with smaller vine maple clones. The results indicate that vine maple gaps may improve the nutritional status of the sites that they occupy within conifer forests. Key words: Litterfall, litter decomposition, soil-plant interactions, vine maple, canopy openings, canopy gaps

Bioassay of forest floor nitrogen supply for plant growth

1986 ◽  
Vol 16 (6) ◽  
pp. 1320-1326 ◽  
Author(s):  
K. Van Cleve ◽  
O. W. Heal ◽  
D. Roberts
Keyword(s):  
Forest Floor ◽  
Black Spruce ◽  
Nitrogen Concentration ◽  
N Uptake ◽  
Mineral Soil ◽  
Forest Types ◽  
N Supply ◽  
Paper Birch ◽  
Forest Floors ◽  
P Supply

Using a bioassay approach, this paper considers the nitrogen-supplying power of forest floors from examples of the major forest types in interior Alaska. Yield and net N uptake by paper birch seedlings grown in standardized mixtures of quartz sand and forest floor organic matter, and separate incubation estimates of N mineralization and nitrification for the forest floors, were employed to evaluate potential N supply. Black spruce and floodplain white spruce forest floors supplied only one-fifth the amount of N taken up by seedlings growing in other forest floors. Incubation estimates showed these forest floors yielded 4 and 15 times less extractable N, respectively, than the more fertile birch forest floors. In comparison with earlier estimates of P supply from these same forest floors, the upland types showed greater deficiency of N whereas floodplain types showed greater deficiency of P in control of seedling yield. The latter condition is attributed to the highly calcareous nature of the floodplain mineral soil, the consequent potential for P fixation, and hence greater potential deficiency of the element compared with N in mineralizing forest floors. Nitrogen concentration of the forest floors was the best predictor of bioassay response.

Different nitrogen sources for fertilizing western hemlock in western Washington

1984 ◽  
Vol 14 (2) ◽  
pp. 155-162 ◽  
Author(s):  
M. A. Radwan ◽  
D. S. DeBell ◽  
S. R. Webster ◽  
S. P. Gessel
Keyword(s):  
Forest Floor ◽  
Mineral Soil ◽  
Volume Growth ◽  
Basal Area ◽  
N Fertilization ◽  
Height Growth ◽  
Growth Responses ◽  
Total N ◽  
Area Growth ◽  
Western Washington

Effects of different sources of fertilizer N on selected chemical characteristics of soils and foliage, and on growth of western hemlock (Tsugaheterophylla (Raf.) Sarg.) were compared at three different sites in western Washington. Treatments were the following: untreated control (O), ammonium nitrate (AN), ammonium sulfate (AS), calcium nitrate (CN), urea (U), and urea – ammonium sulfate (US). Fertilizers were applied in the spring (April–May) at 224 kg N/ha. Forest floor and mineral soil, to a depth of 5 cm, and foliage were sampled periodically for 2 years. Height and diameter of selected trees were measured periodically for 4 years. Results are reported mostly for two sites, one in the Cascade Range and one in the coastal zone in western Washington. The pH of forest floor and mineral soil varied by treatment, and the two urea fertilizers caused substantial initial rise. Effects on soil and foliar nutrients varied by fertilizer, sampling date, and location. In general, all fertilizers increased NH4 N, N03 N, and total N in the forest floor and mineral soil, and total N in the foliage. Also, with some exceptions, especially with foliar P in the Cascade site, fertilization reduced foliar content of important nutrients. At the Cascade site, 4-year growth responses in height, basal area, and volume averaged over all fertilizers were 30, 34, and 32%, respectively. AN, AS, CN, and urea resulted in height growth significantly (P < 0.20) higher than that of the control. Significant basal area growth and volume-growth responses were produced by AN, CN, and US. No significant height-growth response to any fertilizer occurred in the coastal stand; basal area growth and volume-growth responses averaged 27 and 21%, respectively, and best response occurred with urea. These results suggest that the low and inconsistent response of hemlock to N fertilization cannot be improved by applying some N fertilizer other than urea. Factors limiting response to N fertilization may be associated with availability of native N and other nutrients or other characteristics of hemlock sites and stands.

Precipitation nutrients in the open and under two forests in Minnesota

1977 ◽  
Vol 7 (1) ◽  
pp. 112-119 ◽  
Author(s):  
Elon S. Verry ◽  
D. R. Timmons
Keyword(s):  
Forest Floor ◽  
Black Spruce ◽  
Inorganic Nitrogen ◽  
Annual Precipitation ◽  
Nutrient Inputs ◽  
Nutrient Input ◽  
Forest Canopies ◽  
North Central ◽  
Annual Amount ◽  
Total P

Concentrations of N, P, K, Ca, Mg, and Na were measured in rain and snow in the open, and in throughfall and stemflow under black spruce and aspen forests in north-central Minnesota. Concentrations of total P in rain and black spruce throughfall were inversely related to storm size. Annual precipitation nutrient inputs to the forest floor were calculated for each site. In general, nutrient contributions from snow were less than 10% of the annual nutrient input from precipitation at each site, and differences in snow nutrient input between sites were minimal. Rainfall nutrient input differed significantly between sites. Rain and snow passing through both forest canopies were enriched with nutrients except inorganic nitrogen. Total annual nutrients added to the forest floor under the black spruce stand averaged 1.7 times that added in the open; the annual amount added under the aspen stand averaged 5.2 times that added in the open.

Effets du scarifiage sur les propriétés du sol et l'ensemencement naturel dans une pessière noire à mousses de la forêt boréale québécoise

1996 ◽  
Vol 26 (1) ◽  
pp. 72-86 ◽  
Author(s):  
Marcel Prévost
Keyword(s):  
Organic Matter ◽  
Black Spruce ◽  
Block Design ◽  
Mineral Soil ◽  
Organic Matter Content ◽  
Control Treatment ◽  
Winter Period ◽  
Soil Warming ◽  
Total N ◽  
Humus Layer

Two types of scarification (cone and disk) were applied at two intensities (simple and double passes), in a randomized complete block design, established alongside buffer stands of spruce protective of water courses, that provided a natural seed source. Treatment effects on seedbed evolution, natural seeding of black spruce (Piceamariana (Mill.) BSP), competing vegetation, and soil physical and chemical properties were examined over a 3-year period. In situ nitrogen mineralization was also studied, using the buried-bag method. All scarification treatments created a surface horizon (0–10 cm) with 80% less organic matter content than the control treatment. However, treatments tended to loosen the exposed deep layers, creating microsites whose compactness appeared adequate for root development (1.07–1.22 Mg/m3). The organic matter loss mainly decreased exchangeable K and Mg in the surface 20 cm of scarified microsites. Scarification had little impact on total N of sampled profiles and clearcutting did not increase N mineralization with regard to the forest, during the first year after disturbance. The weak soil warming and the stability of temperatures under the unscarified humus suggest that clearcutting did not significantly enhanced microbial activity on the site. However, removal of the insulating humus layer allowed a significant summer soil warming in the furrows. Despite this, scarified microsites were characterized by N immobilization during the first growing season after treatment. However, net N production was positive during the winter period, presumably because of a N-flux phenomenon. Scarification improved black spruce regeneration by natural seeding. Three years after treatment, stocking levels reached 40 to 51% in the scarified sectors while they reached 31% in the controls, this gap being mainly attributed to the second germination year. The difference can be explained by the improved receptivity of bare mineral soil, well-decomposed humus, and mixed mineral–organic seedbeds that covered 12–20% of the scarified areas immediately after treatment. Generally, results indicate that microsites created by a light scarification are as receptive as microsites created by a severe perturbation. Finally, every scarification treatment efficiently controlled the ericaceous shrub cover during the first 3 years after treatment.

Productivity and nutrient cycling in taiga forest ecosystems

1983 ◽  
Vol 13 (5) ◽  
pp. 747-766 ◽  
Author(s):  
Keith Van Cleve ◽  
Lola Oliver ◽  
Robert Schlentner ◽  
Leslie A. Viereck ◽  
C. T. Dyrness
Keyword(s):  
Nutrient Cycling ◽  
Soil Temperature ◽  
Forest Floor ◽  
Black Spruce ◽  
Lignin Content ◽  
Mineral Soil ◽  
Forest Types ◽  
Interior Alaska ◽  
Taiga Forest ◽  
Tree Production

This paper considers the productivity and nutrient cycling in examples of the major forest types in interior Alaska. These ecosystem properties are examined from the standpoint of the control exerted over them by soil temperature and forest-floor chemistry. We conclude that black spruce Piceamariana (Mill.) B.S.P. occupies the coldest, wettest sites which support tree growth in interior Alaska. Average seasonal heat sums (1132 ± 32 degree days (DD)) for all other forest types were significantly higher than those encountered for black spruce (640 ± 40 DD). In addition, black spruce ecosystems display the highest average seasonal forest-floor and mineral-soil moisture contents. Forest-floor chemistry interacts with soil temperature in black spruce to produce the most decay-resistant organic matter. In black spruce the material is characterized by the highest lignin content and widest C/N (44) and C/P (404) ratios. Across the range of forest types examined in this study, soil temperature is strongly related to net annual aboveground tree production and the annual tree requirement for N, P, K, Ca, and Mg. Forest floor C/N and C/P ratios are strongly related to annual tree N and P requirement and the C/N ratio to annual tree production. In all cases these controls act to produce, in black spruce, the smallest accumulation of tree biomass, standing crop of elements, annual production, and element requirement in aboveground tree components.

Organic matter and nitrogen content of the forest floor in even-aged northern hardwoods

1984 ◽  
Vol 14 (6) ◽  
pp. 763-767 ◽  
Author(s):  
C. Anthony Federer
Keyword(s):  
Organic Matter ◽  
Nitrogen Content ◽  
Bulk Density ◽  
Forest Floor ◽  
Mineral Soil ◽  
Organic Content ◽  
Stand Age ◽  
Clear Cutting ◽  
Mature Forest ◽  
Forest Floors

Organic content of the forest floor decreases for several years after clear-cutting, and then slowly recovers. Thickness, bulk density, organic matter, and nitrogen content of forest floors were measured for 13 northern hardwood stands in the White Mountains of New Hampshire. Stands ranged from 1 to about 100 years in age. Forest-floor thickness varied significantly with stand age, but bulk density, organic fraction, and nitrogen fraction were independent of age. Total organic content of the forest floor agreed very well with data from Covington's (W. W. Covington 1981. Ecology, 62: 41–48) study of the same area. Both studies indicated that mature forest floors have about 80 Mg organic matter•ha−1 and 1.9 Mg nitrogen•ha−1. Within 10 or 15 years after cutting, the organic matter content of the floor decreases to 50 Mg•ha−1, and its nitrogen content to 1.1 Mg•ha−1. The question whether the decrease is rapid and the minimum broad and flat, or if the decrease is gradual and the minimum sharp, cannot be answered. The subsequent increase to levels reached in mature forest requires about 50 years. Some of the initial decrease in organic matter and nitrogen content of the forest floor may be caused by organic decomposition and nitrogen leaching, but mechanical and chemical mixing of floor into mineral soil, during and after the harvest operation, may also be important. The difference is vital with respect to maintenance of long-term productivity.

Bioassay of forest floor phosphorus supply for plant growth

1985 ◽  
Vol 15 (1) ◽  
pp. 156-162 ◽  
Author(s):  
K. Van Cleve ◽  
F. Harrison
Keyword(s):  
Forest Floor ◽  
Black Spruce ◽  
Forest Types ◽  
P Deficiency ◽  
Interior Alaska ◽  
Rooting Medium ◽  
Forest Floors ◽  
Work Supports ◽  
Plant Bioassays

This paper considers the extent to which phosphorus (P) supply for plant use is controlled by the chemical quality of forest floor organic matter, independent of climate. Using plant bioassays, forest floor materials from representative examples of each of the major forest types in interior Alaska were examined for nutrient supplying power. The work supports conclusions reached in earlier studies which indicated that black spruce forest floors were highly nutrient limited compared with those of other interior Alaska forest types. In addition, floodplain white spruce forests may experience marked P deficiency because of dilution of the element by periodic siltation. Potential phosphorus supply for seedling growth was best described by P concentration of the rooting medium. The supply also was related to the concentrations of lignin and tannin which control forest floor decomposition and recycling of P within the microbial population.

Characterization of soil P in coastal forest chronosequences of southern Vancouver Island: effects of climate and harvesting disturbance

2000 ◽  
Vol 80 (4) ◽  
pp. 633-647 ◽  
Author(s):  
C. M. Preston ◽  
J. A. Trofymow
Keyword(s):  
Tree Growth ◽  
Mineral Soil ◽  
Vancouver Island ◽  
Available P ◽  
Organic P ◽  
Coastal Forest ◽  
East Side ◽  
The West ◽  
West Side ◽  
Total P

Limitation of tree growth due to inadequate P supply has been found for young plantations following harvesting of old-growth in high rainfall areas of coastal British Columbia. To understand the reasons for P limitation, we investigated P chemistry in mineral soil to 50 cm depth in sites from the Coastal Forest Chronosequence project on Vancouver Island. This allowed comparison of biogeoclimatic subzone (higher rainfall on west than east coast sites) and of time from harvesting disturbance (seral stage). Available (Bray 1) P was significantly higher (P < 0.001) on the drier east side (up to 50 mg kg−1), than on the west side (<5 mg kg−1), although total P values were less divergent (694 mg kg−1, east and 534 mg kg−1, west). There were no significant seral stage effects on total and available P. Extraction with 0.5 M NaOH recovered 50–60% of total P, except for samples from 10–30 cm depth on the west side, for which only 20% was recovered, an effect not found for C. Analysis of the NaOH extracts by 31P nuclear magnetic resonance (NMR) spectroscopy showed much higher proportions of orthophosphate P on the east side. West side extracts were higher in organic P forms, especially diesters, typical of forest ecosystems with restricted nutrient cycling and high precipitation. On the west side, low concentrations of available P, higher proportions of organic P in NaOH extracts, and depression of NaOH extractability at 10–30 cm are consistent with P being a limiting nutrient for tree growth, a problem that may be exacerbated by harvesting disturbance. Key words: Forest chronosequences, harvesting disturbance, 31P NMR, P cycling, organic P

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