Crown stratification by species in even-aged mixed stands of Douglas-fir – western hemlock

1979 ◽  
Vol 9 (1) ◽  
pp. 1-9 ◽  
Author(s):  
Charles A. Wierman ◽  
Chadwick Dearing Oliver

The pattern of vertical stratification in mixed Douglas-fir (Pseudotsugamenziesii (Mirb.) Franco) and western hemlock (Tsugaheterophylla (Raf.) Sarg.) stands was documented by observing tree interaction patterns on temporary plots in stands between 35 and 80 years old. It was found that Douglas-fir predictably dominated the hemlock. Several possible causes of the stratification were studied. They were differences in ages between species, differences in mortality patterns, and differences in growth rates. Increment borings of trees on sample plots in the 35- to 80-year stands with the observed stratification revealed the overtopping Douglas-firs and suppressed hemlocks to be essentially of the same age. Searches for dead or dying Douglas-fir in groups of mixed stands of 35 to 80 years, 22 to 35 years, and less than 12 years revealed very little Douglas-fir mortality by suppression. Therefore, the possibility of death of intermediate and suppressed Douglas-firs was not an important factor leading to Douglas-firs being found primarily in the upper crown positions. Height and diameter growth patterns of closely growing codominant Douglas-firs, co-dominant hemlocks, and suppressed hemlocks were documented by stem analyses. Douglas-fir became significantly taller than the hemlocks after about 20 years and suppressed competing hemlocks. The possibility of stratification in mixed planted stands is less certain, since only young planted stands were available for study. Management of such mixed stands appears to improve the yield and quantity of wood and reduce or eliminate costly precommercial thinnings.

2006 ◽  
Vol 36 (6) ◽  
pp. 1484-1496 ◽  
Author(s):  
M M Amoroso ◽  
E C Turnblom

We studied pure and 50/50 mixtures of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and western hemlock (Tsuga heterophylla (Raf.) Sarg.) plantations to compare attained total yields between mixed-species stands as opposed to monocultures of equal densities. Whether overall stand density influences this outcome has not been adequately investigated, and to address this we included three density levels (494, 1111, and 1729 trees/ha) in the analysis. At age 12, as components of the mixed stands, Douglas-fir exhibited greater height, diameter, and individual-tree volume than western hemlock at all densities. At 494 and 1111 trees/ha the monocultures had a higher volume per hectare than the mixed stand, but at 1729 trees/ha the mixed stand appeared to be just as productive as the pure stands. The increase in productivity by the mixture at high densities seems to have resulted from the partial stratification observed and most likely also from better use of the site resources. Because of this, less interspecific competition was probably experienced in the mixed stand than intraspecific competition in the pure stands. This study shows the important role density plays in the productivity of mixed stands and thus in comparing mixed and pure stands.


1986 ◽  
Vol 16 (2) ◽  
pp. 367-372 ◽  
Author(s):  
Bruce C. Larson

Stand development patterns and growth rates of even-aged mixed stands of Douglas-fir (Pseudotsugamensiesii Mirb. Franco) and grand fir (Abiesgrandis Dougl. Lindl.) were investigated by stand reconstruction in eastern Washington. Although Douglas-fir dominated the stands early, grand fir of the same age eventually overtook the tallest Douglas-firs. Dominant trees of both species were found to have highly variable taper form (based on basal area to height ratios), although a linear relationship existed in other crown classes. Stem volume growth rates were determined by reconstructing past diameters and heights and were found comparable to site class IV Douglas-fir in western Washington. Intensive management, especially planting and thinning, could improve upon the observed mean annual increments of 800 board feet per acre at 80 years of age.


2019 ◽  
Vol 20 (6) ◽  
Author(s):  
RIGNOLDA DJAMALUDDIN

Abstract. Djamaluddin R. 2019. Growth pattern in tropical mangrove trees of Bunaken National Park, North Sulawesi, Indonesia. Biodiversitas 20: 1713-1720. Seasonal diameter growth patterns in mangrove are often related to rainfall, temperature and moisture regime. At any localities, specific environmental factors may influence growth rate of individual trees. I asked whether stem growth of tropical mangrove in BNP is constant over a year, and whether stem growth rates are different by sites, species, and trees of the same species. Dendrometer bands were installed on trees from twelve different sites in BNP to measure stem growth rates. Measurements were made at two months intervals from July 1999 to June 2001 and March 2014 to December 2016. Growth rates measured in trees at the twelve sites varied significantly from 0.83 ± 0.27 to 1.71 ± 0.31 mm month-1. Growth rates were higher on Sonneratia alba (1.65 ± 0.69 mm month-1), low on Rhizophora stylosa, Xylocarpus moluccensis, Avicennia marina, Ceriops tagal (0.82 ± 0.16, 0.82 ± 0.18, 0.85 ± 0.18, 0.88 ± 0.28 mm month-1, respectively), and medium on Rhizophora apiculata, Bruguiera parviflora, Bruguiera gymnorrhiza, Rhizophora mucronata (1.19 ± 0.16, 1.22 ± 0.69, 1.25 ± 0.49, 1.31 ± 0.22 mm month-1, respectively). Statistically, growth rates were higher in trees with initial girths more than 50 cm compared to trees with initial girth less than 50 cm on B. gymnorrhiza, C. tagal, and R. mucronata, but these were slower for R. apiculata and S. alba. Between individual trees in six species tested, the differences in diameter growth rate were statistically significant. Growth rates varied among different site conditions, and the effect of soil water salinity on these was significant at higher intertidal sites. Growth rates exhibited seasonal patterns, and these were correlated positively with rainfall and negatively with temperature. The effect of the 2015-2016 El Nino was significant on reduced growth rates.  


2004 ◽  
Vol 34 (2) ◽  
pp. 384-395 ◽  
Author(s):  
Richard Kabzems ◽  
Oscar García

The trembling aspen (Populus tremuloides Michx.) – white spruce (Picea glauca (Moench) Voss) mixed woods near Fort Nelson are distinguished by the large size of individual trees, longevity, and the low occurrence of internal decay in trembling aspen. The development of these forest ecosystems has had limited documentation and may be significantly different than those described in other portions of the boreal forest. At five study stands, stem analysis techniques were used to examine the patterns of height and radial growth over time according to species and structure type. There were two patterns of species establishment that were consistent with the stand structure. In codominant stands, recruitment periods for trembling aspen and white spruce overlapped. The stratified stands were consistently associated with a 29- to 58-year lag in white spruce recruitment. Spruce that were codominant with aspen at the time of sampling had sustained periods of rapid height and diameter growth. White spruce that were later to establish on site had slower rates of height and diameter growth. White spruce ages indicated that a dominant recruitment episode was more common than continuous recruitment. Height and diameter growth of trembling aspen were similar in both stand types. The differences in trembling aspen growth patterns between stands were due to site quality. The white spruce in codominant stands did not appear to go through a period of suppression and then release associated with stand-level trembling aspen mortality, as commonly described for other boreal mixedwoods. The vigor and longevity of trembling aspen in Fort Nelson appear to prolong the period of trembling aspen domination of mixed stands well beyond the time periods observed in other boreal ecosystems.


1955 ◽  
Vol 31 (1) ◽  
pp. 45-59 ◽  
Author(s):  
G. S. Allen ◽  
I. K. Barber ◽  
Ian Mahood

An area of 738 acres about 20 miles north and west of Port Alberni on Vancouver Island was seeded by helicopter to Douglas fir (1/2 pound per acre) and western hemlock (1/5 pound per acre) in November 1951. One week earlier this area and a surrounding buffer strip, totalling 1760 acres, had been baited by helicopter using wheat treated with thallous sulphate and "1080".Baiting was successful and the roden:trap ratio dropped from 1:4.6 to 1:94.3 as a result. By April the ratio had climbed again to 1:18.7. By the first week of June 1952, a total of about 1400 Douglas fir and 840 hemlock had germinated per acre. By November 1952, this had dropped to 1000 Douglas fir and 198 hemlock per acre. A final extensive survey in June 1954, showed a total of 730 Douglas fir and 57 hemlock of the 1952 crop per acre with a milacre stocking of 42.0 percent for Douglas fir and 43.9 percent for Douglas fir and hemlock. The 4-milacre stocking due to the 1952 Douglas fir seedlings was 78.3 percent. Practically all of the seedlings that germinated in 1952 are attributed to the seeding operation because of the relatively uniform distribution of seedlings and the lack of seedlings on the control area.The seeding operation in itself restocked 628 acres to B.C. Forest Service minimum standards (31 percent by 1-milacres). It restocked 708 acres to U.S. "medium" or "good" standards. With natural trees included, only 28 acres are below B.C. Forest Service standards, and 6 acres below U.S. "medium" stocking. Total stocking, including natural trees, is 59.3 percent by 1-milacres and 88.6 percent by 4-milacres.The baiting and seeding operation can be considered highly successful. The surviving Douglas firs are healthy and vigorous and show little deer or grouse damage to date. Distribution of the seedlings is good and most gaps that occurred are believed due to uneven local distribution of seed. Considering that this experiment was carried out on a rather steep south exposure and that the first summer following seeding was unusually hot and dry, the satisfactory results suggest that direct seeding following rodent control has a definite place in West Coast forestry practice. It may well take the place of more expensive and troublesome planting on many areas that are slow to restock.


Mycologia ◽  
1971 ◽  
Vol 63 (3) ◽  
pp. 567-574 ◽  
Author(s):  
A. Funk ◽  
R. A. Shoemaker
Keyword(s):  

1997 ◽  
Vol 75 (9) ◽  
pp. 1424-1435 ◽  
Author(s):  
D. Mailly ◽  
J. P. Kimmins

Silvicultural alternatives that differ in the degree of overstory removal may create shady environments that will be problematic for the regeneration of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco). Gradients of light in the field were used to compare mortality, growth, and leaf morphological acclimation of two conifer species of contrasting shade tolerances: Douglas-fir and western hemlock (Tsuga heterophylla (Raf.) Sarg.). Results after two growing seasons indicated that Douglas-fir mortality occurred mainly at relative light intensity (RLI) below 20%, while western hemlock mortality was evenly distributed along the light gradient. Height, diameter, and biomass of the planted seedlings increased with increasing light for both species but at different rates, and maximum biomass accumulation always occurred in the open. Douglas-fir allocated more resources to stem biomass than western hemlock, which accumulated more foliage biomass. Increases in specific leaf area for Douglas-fir seedlings occurred at RLI ≤ 0.4 and red/far red (R/FR) ratio ≤ 0.6, which appear to be the minimal optimum light levels for growth. Conversely, western hemlock seedlings adjusted their leaf morphology in a more regular pattern, and changes were less pronounced at low light levels. These results, along with early mortality results for Douglas-fir, suggest that the most successful way to artificially regenerate this species may be by allowing at least 20% of RLI for ensuring survival and at least 40% RLI for optimum growth. Key words: light, light quality, leaf morphology, acclimation.


1972 ◽  
Vol 2 (3) ◽  
pp. 256-263 ◽  
Author(s):  
D. G. W. Edwards ◽  
Patricia E. Olsen

Coating Douglas-fir and western hemlock seeds with R-55, a rodent repellent, seriously reduced normal germination. Acidifying the latex sticker, or adding graphite powder, had no effect. If seeds were coated first with latex, more R-55 could be applied without significantly increasing germination losses. Both stratification and dry storage of R-55 coated seeds increased germination losses. Treatment differences detected by normal germination percentages were also reflected in germination values. Observations based on total (normal + abnormal) germination obscured the effects of the various coatings.


Nature ◽  
2015 ◽  
Vol 531 (7595) ◽  
pp. 538-538
Author(s):  
Gregory M. Erickson ◽  
Kristina Curry Rogers ◽  
Scott A. Yerby
Keyword(s):  

2019 ◽  
Vol 433 ◽  
pp. 105-110
Author(s):  
Matthew E. Hane ◽  
Andrew J. Kroll ◽  
Aaron Springford ◽  
Jack Giovanini ◽  
Mike Rochelle ◽  
...  

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