Field evaluation of ecological differentiation of dwarf mistletoe on shore pine and western hemlock

1976 ◽  
Vol 6 (2) ◽  
pp. 225-228 ◽  
Author(s):  
R. B. Smith ◽  
E. F. Wass
Keyword(s):  
British Columbia ◽  
Field Evaluation ◽  
Field Studies ◽  
Western Hemlock ◽  
Lake Area ◽  
Dwarf Mistletoe ◽  
Varied Composition ◽  
Coastal British Columbia ◽  
Ecological Races ◽  
Shore Pine

Hemloclc dwarf mistletoe (Arcenthobiiuntsugense (Rosend.) G. N. Jones) principally infects western hemlock (Tsugaheteropliylla (Raf.) Sarg.), but it is also damaging to shore pine (Pinuscontorta Dougl. ex Loud.) growing in coastal British Columbia. Stands of varied composition were studied in the Home Lake area of Vancouver Island to compare the levels of infection in shore pine and western hemlock. Infection of shore pine occurred whether or not infected hemlock trees were present in the stand. Observations and measurements from these field studies support indications from earlier artificial inoculation trials that two ecological races of A. tsugense exist.

Epidemiology of dwarf mistletoe in western hemlock stands in south coastal British Columbia

1985 ◽  
Vol 15 (5) ◽  
pp. 909-913 ◽  
Author(s):  
R. I. Alfaro ◽  
W. J. Bloomberg ◽  
R. B. Smith ◽  
A. J. Thomson
Keyword(s):  
British Columbia ◽  
Spatial Distribution ◽  
Distribution Pattern ◽  
Tree Mortality ◽  
Infection Intensity ◽  
Western Hemlock ◽  
Intensity Level ◽  
Dwarf Mistletoe ◽  
Infection Index ◽  
Coastal British Columbia

Surveys of seven western hemlock (Tsugaheterophylla) stands in south coastal British Columbia using fixed-radius plots assessed dwarf mistletoe (Arceuthobiumtsugense) infection intensity level and spatial distribution patterns. Regression analysis indicated a close relationship between the plot infection index (average dwarf mistletoe rating of trees in a plot) and the percentage of infected trees. Plot infection index reached 4.0 when all trees in a plot were infected. Infected trees were either associated with widely spaced infection centers or were more or less evenly distributed throughout the stand. Spatial distribution pattern and spread rate were related to (i) the severity and distribution pattern of the initial inoculum sources, (ii) the manner of stand regeneration and resulting stand structure, and (iii) the presence of barriers to spread such as a high nonhost component and drastic slope increases. Tree mortality from dwarf mistletoe averaged 0.6% (range, 0–1.8%) and was much lower than mortality from other causes. On average, 9.3% of the trees had large dwarf mistletoe caused swellings on the lower third of the bole.

scholarly journals Estimating the Number of Filled Seeds per Cone of Western Hemlock from Coastal British Columbia

1996 ◽  
Vol 11 (2) ◽  
pp. 44-49
Author(s):  
M. D. Meagher
Keyword(s):  
British Columbia ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Linear Relationships ◽  
The Face ◽  
Predetermined Level ◽  
Exponential Factors ◽  
The Relationship ◽  
Coastal British Columbia ◽  
Filled Seeds

Abstract Unopened western hemlock (Tsuga heterophylla) cones from coastal British Columbia were tested for the relationship between total filled seeds per cone and the number of sound seeds exposed by slicing through the center of the cone's long axis, and with cone length. Predictive linear relationships of total number of filled seeds per cone (TFS), based on number of filled seeds cut on the face through the cone's long axis (CC), were found in a cone sample bulked from a number of trees. The regression based on CC explained about 63% of the variation in TFS. More accurate estimates of TFS were found where cone length, and exponential factors of CC and cone length, were included in the analysis. Analyses of cones from seven parents did not find statistically significant trends of TFS on CC in all cases, and differed in slope in most cases. Sample sizes (cones) to estimate TFS to a predetermined level of precision were much larger than cones needed to accept or reject a tree for study. West. J. Appl. For. (11)2:44-49.

Development of dwarf mistletoe in western hemlock regeneration in southeast Alaska

1982 ◽  
Vol 12 (3) ◽  
pp. 482-488 ◽  
Author(s):  
Charles G. Shaw III
Keyword(s):  
British Columbia ◽  
Disease Control ◽  
Control Strategies ◽  
Severe Infection ◽  
Host Tissue ◽  
Western Hemlock ◽  
Dwarf Mistletoe ◽  
Local Conditions ◽  
Southeast Alaska ◽  
Understory Trees

Western hemlock (Tsugaheterophylla (Raf.) Sarg.) regenerated within 9.14 m of old-growth hemlock severely infected with dwarf mistletoe, Arceuthobiumtsugense ((Rosendahl) G.N. Jones), and left standing on cutover sites in southeast Alaska were felled and examined for infection. The percentage of understory trees infected among the 3429 examined averaged 9, 7, 5, and 17, respectively, in 17-, 19-, 35-, and 43-year-old stands. Ninety-two percent of all infections in the 17-, 19-, and 35-year-old stands were alive, but only 51% were alive in the 43-year-old stand. In all stands few trees had more than two live infections and few had any infections in their middle or upper crown. In all stands, advanced regeneration was more frequently infected and accounted for a significantly greater proportion of crop trees than new reproduction. A significantly higher proportion of hemlock crop trees were infected than non crop trees, but the number of infections on crop trees was consistently low. Most infections were established on host tissue 5 years or younger, but tissue up to 13 years old was infected. At a comparable age, young stands in Alaska appear to be less severely affected by A. tsugense than similar stands in Washington, Oregon, and British Columbia. Disease control strategies, developed for stands experiencing more severe infection intensities than those now known to prevail in southeast Alaska, will require modification to meet local conditions.

scholarly journals Western redcedar population dynamics in old-growth forests: Contrasting ecological paradigms using tree rings

2003 ◽  
Vol 79 (3) ◽  
pp. 517-530 ◽  
Author(s):  
Lori D Daniels
Keyword(s):  
British Columbia ◽  
Population Dynamics ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Silver Fir ◽  
Old Growth ◽  
Western Redcedar ◽  
Old Growth Forests ◽  
Gap Phase ◽  
Coastal British Columbia

In coastal British Columbia, late-successional forests dominated by western redcedar (Thuja plicata Donn ex D. Don) are structurally complex, with deep multi-layered canopies, large trees that are > 250 years old, and abundant coarse woody debris (CWD). These forests are presumed to be "old-growth" forests in which fine-scale gaps are the dominant disturbance regime, accounting for their structural diversity. In this study, I have used tree-ring analyses to investigate western redcedar regeneration dynamics in these old-growth forests. Western redcedar dominates canopies of many stands, but is rare in the understorey although it tolerates shade. The traditional interpretation is that western redcedar depends on catastrophic disturbance to regenerate and that it is replaced through succession by western hemlock (Tsuga heterophylla (Raf.) Sarg.) and Pacific silver fir (Abies amabilis (Dougl. ex Loud.) Dougl. ex J. Forbes), which are abundant in the understorey. Dominant trees are perceived to represent an even-aged, post-disturbance cohort and the lack of regeneration indicates a population decline in the absence of catastrophic disturbances. In this paper, I investigate four assumptions underlying this interpretation: (1) Tree size indicates age. (2) Populations establish as even-aged, post-disturbance cohorts. (3) Abundant CWD represents recent mortality. (4) Regeneration is insufficient to maintain canopy dominance. Using tree-ring evidence, I show that population dynamics of western redcedar are a combination of gap-phase establishment and a continuous mode of recruitment from the sub-canopy to the canopy. Specifically, size is a poor surrogate for tree age. Age distributions from 15 sites revealed uneven-aged populations, rather than single post-disturbance cohorts. Both logs and snags of western redcedar may persist more than 270 years; they do not represent recent mortality that is disproportionate to the number of live western redcedar in canopy. The regeneration niches of western redcedar and western hemlock overlap. For both species, gap-phase disturbances result in substrate suitable for successful seedling establishment. Preliminary results from dendroecological analysis of radial growth rates of trees in the subcanopy and canopy strata suggest that western hemlock and Pacific silver fir depend on gaps to recruit to the upper canopy, but recruitment of western redcedar may be independent of canopy gaps. I propose that differences in mode of recruitment to the canopy may explain the differences in population structures between western redcedar, western hemlock, and Pacific silver fir in the old-growth forest. These results provide an ecological precedent for use of a range of silvicultural systems, including clearcuts through single-tree harvesting and protection forests, when managing western redcedar in coastal British Columbia. Key words: Coastal British Columbia, disturbance regimes, regeneration dynamics, Thuja plicata, variable retention silviculture

Overturning resistance of western redcedar and western hemlock in mixed-species stands in coastal British Columbia

2007 ◽  
Vol 37 (5) ◽  
pp. 931-939 ◽  
Author(s):  
Kenneth E. Byrne ◽  
Stephen J. Mitchell
Keyword(s):  
British Columbia ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Thuja Plicata ◽  
Specific Information ◽  
Mixed Species ◽  
Stem Form ◽  
Western Redcedar ◽  
Applied Forces ◽  
Coastal British Columbia

Specific information about the applied forces that cause trees to fail is required to validate mechanistic models of windthrow in different forest types. Static tree-pulling tests were conducted to examine the overturning resistance of western redcedar ( Thuja plicata Donn ex D. Don) and western hemlock ( Tsuga heterophylla (Raf.) Sarg.) in a mixed species second-growth stand in coastal British Columbia. Although widely used, tree-pulling techniques are not standardized. Data from three inclinometers were used to estimate stem deflection, which was found to increase with tree slenderness. Differing methods of fitting stem curvature had a small effect on estimates of self-loading at failure. The distance of the pivot point from the centre of the stem base increased with tree diameter. Accounting for the correct self-loading at failure produced a small difference in the overall turning moment regressions but did not improve the fit of these regressions. However, this difference increased with tree size and warrants consideration in future tree-pulling tests with large or plate-rooted trees. The stem mass – overturning resistance relationship had the best fit and was not significantly different for these species in spite of their differences in wood density and stem form.

EcologyDevelopment of Dwarf Mistletoe (Arceuthobium) Infections on Western Hemlock, Shore Pine, and Western Larch

1971 ◽  
Vol 1 (1) ◽  
pp. 35-42 ◽  
Author(s):  
R. B. Smith
Keyword(s):  
Seed Dispersal ◽  
Life Span ◽  
Western Hemlock ◽  
First Year ◽  
Mature Fruit ◽  
Dwarf Mistletoe ◽  
Western Larch ◽  
The Third ◽  
Second Year ◽  
Shore Pine

Newly dispersed seeds, established infections, and individually tagged shoots of hemlock dwarf mistletoe on western hemlock and shore pine, and established infections and individual shoots of larch mistletoe on western larch were examined periodically for up to 7 years, Infections on hemlock first appeared as swellings, some of which were visible during the first year after seed dispersal. Aerial shoots appeared in the second year. By the end of the third year most infections exhibited both swellings and aerial shoots. On all three hosts, mature mistletoe fruit were produced occasionally in the fourth year but generally not until the fifth year. Most female aerial shoots bore at least one crop of flowers during their existence, however, only half produced mature fruit. Aerial shoots varied in their life span from less than 1 year up to 7 years. The average was 2 to 3 years. One female shoot on larch bore five successive crops of flowers. The fastest longitudinal growth of mistletoe endophytic system occurred on western larch and the slowest was on shore pine. All infections were nonsystemic in that the endophytic system was largely confined to the swollen portions of the hosts.

Vanterpoolia, a new genus of sporodochial hyphomycetes

1982 ◽  
Vol 60 (6) ◽  
pp. 973-975 ◽  
Author(s):  
A. Funk
Keyword(s):  
British Columbia ◽  
New Genus ◽  
Single Species ◽  
Tsuga Heterophylla ◽  
Western Hemlock ◽  
Coastal British Columbia

Vanterpoolia, a new genus of sporodochial hyphomycetes, is erected for a fungus occurring on necrotic, newly flushed needles of western hemlock (Tsuga heterophylla (Raf.) Sarg.) in coastal British Columbia, Canada. The genus is characterized by triradiate conidia or rodlike, appendaged conidia produced in acropetal chains from simple, septate conidiophores arising from a pseudoparenchymatous stroma. A single species, Vanterpoolia tsugae n.sp., is described.

The Rate of Upward Advance and Intensification of Dwarf Mistletoe on Immature Western Hemlock

1972 ◽  
Vol 2 (3) ◽  
pp. 313-316 ◽  
Author(s):  
K. S. Richardson ◽  
B. J. van der Kamp
Keyword(s):  
British Columbia ◽  
Height Growth ◽  
Western Hemlock ◽  
Dwarf Mistletoe ◽  
Rate Of Spread ◽  
History Of ◽  
Dense Stand ◽  
Individual Trees ◽  
Crown Closure

The rate of upward advance and intensification of dwarf mistletoe on immature coastal western hemlock were determined by reconstructing the infection history of individual trees from data on age, height above ground, and sex of mistletoe infections present at the time of observation. The vertical rate of spread was 30 ± 4 cm/year in a dense stand (750 trees per hectare, 19 m in height) and 65 ± 4 cm/year in an open stand (250 trees per hectare, 26 m in height). This compares with height growth of the trees of 33 and 58 cm/year in the two stands, respectively. The number of new infections per year increased logarithmically with time, doubling roughly every 4 years. An asymtotic stabilization of the rate of intensification over the last 6 years probably reflects crown closure. The results suggest that dwarf mistletoe may not be a serious threat to thrifty, immature, dense, coastal hemlock stands in southern British Columbia, unless such stands are overtopped by infected residual trees.

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