Influence of stocking on radial and longitudinal variation in modulus of elasticity, microfibril angle, and density in a 24-year-old Pinus radiata thinning trial

2011 ◽  
Vol 41 (7) ◽  
pp. 1422-1431 ◽  
Author(s):  
Michael S. Watt ◽  
Branislav Zoric ◽  
Mark O. Kimberley ◽  
Jonathan Harrington
Keyword(s):  
Wood Density ◽  
Pinus Radiata ◽  
Modulus Of Elasticity ◽  
Significant Interaction ◽  
Graphical Model ◽  
Microfibril Angle ◽  
Wood Properties ◽  
Tree Age ◽  
Longitudinal Variation ◽  
Maximum Difference

Detailed radial measurements of wood properties at four heights (0, 1.4, 5, and 20 m) were taken from 24-year-old Pinus radiata D. Don growing at four final crop stockings (200, 350, 500, and 1100 stems·ha–1). Using these measurements, the objectives of the study were to examine pith-to-bark trends at several heights to (i) determine how stocking influenced modulus of elasticity (MoE), wood density, and microfibril angle (MFA), (ii) quantify the relations among these properties and age at different stocking levels, and (iii) develop a graphical model for MoE across the stocking range. The influence of stocking on all wood properties was primarily expressed through a highly significant interaction between age and stocking. Wood properties in the highest stocking treatment diverged from those in the lowest stocking treatment at tree age 5 to reach a maximum difference of 92 kg·m–3 (488 vs. 580 kg·m–3) at tree age 18 for density, –5.7° (29.2° vs. 23.5°) at tree age 10 for MFA, and 5.1 GPa (12.1 vs. 17.2 GPa) at tree age 20 years for MoE. Graphical predictions from the model show greatest gains in MoE at high final crop stocking to occur over the lower part of the stem.

Determining the main and interactive effect of age and clone on wood density, microfibril angle, and modulus of elasticity for Pinus radiata

2010 ◽  
Vol 40 (8) ◽  
pp. 1550-1557 ◽  
Author(s):  
Michael S. Watt ◽  
Charles Sorensson ◽  
Dave J. Cown ◽  
Heidi S. Dungey ◽  
Robert Evans
Keyword(s):  
Pinus Radiata ◽  
Modulus Of Elasticity ◽  
Interactive Effect ◽  
Microfibril Angle ◽  
Wood Properties ◽  
Radiata Pine ◽  
Linear Increase ◽  
Interactive Effects ◽  
Tree Age ◽  
Clonal Variation

Detailed radial measurements of wood properties, taken at breast height, were obtained from control pollinated seedlings and a selection of 13 year old radiata pine ( Pinus radiata D. Don) clones. Using these data the key objectives of this study were to determine (i) the magnitude of mean clonal variation in modulus of elasticity (MOE) and properties affecting MOE (density and microfibril angle (MFA)) and (ii) whether there is a significant age × clone interaction for these traits. All wood properties were significantly affected by the main and interactive effects of age and clone. There was a relatively linear increase in both MOE and density with tree age, while MFA declined linearly with tree age. Values of density and MOE diverged between the clonal extremes from age 3 to age 12. After diverging markedly up to age 6, differences in MFA between clonal extremes remained relatively constant to age 12. At age 12, values for density, MFA, and MOE varied between clonal extremes by, respectively, 194 kg·m–3 (465–659 kg·m–3), 11.3° (9.6–20.9°), and 11.2 GPa (10.4–21.6 GPa). The seedling material had a relatively intermediate ranking, across the age range, for all traits considered.

Inheritance of density, microfibril angle, and modulus of elasticity in juvenile wood of Pinus radiata at two locations in Australia

2007 ◽  
Vol 37 (11) ◽  
pp. 2164-2174 ◽  
Author(s):  
Brian S. Baltunis ◽  
Harry X. Wu ◽  
Mike B. Powell
Keyword(s):  
Genetic Correlation ◽  
Pinus Radiata ◽  
Modulus Of Elasticity ◽  
Juvenile Wood ◽  
Wood Quality ◽  
Genetic Correlations ◽  
Microfibril Angle ◽  
Wood Properties ◽  
Quality Traits ◽  
Selection For

A total of 1640 increment cores from 343 radiata pine ( Pinus radiata D. Don) families were sampled at two second-generation progeny trials, aged 6 and 7 years, for a detailed genetic study of juvenile wood quality traits. Density, microfibril angle (MFA), and modulus of elasticity (MOE) were determined from pith to bark using SilviScan® technology. Heritability was greatest for area-weighted density at the two sites (0.63 and 0.77, respectively), and the lowest for growth traits (<0.23). Genotype by environment interaction was low for all three wood quality traits. A positive genetic correlation between density and MOE (0.43), and a highly negative, and therefore, favourable genetic correlation between MFA and MOE (–0.92) were observed, implying that improvement of multiple juvenile wood properties is possible. The genetic correlations between whole-core wood quality traits and individual-ring measurements suggest that improvement for juvenile wood properties across the entire profile of the corewood including the innermost rings can be achieved. However, density, MFA, and MOE had unfavourable genetic correlations with diameter growth suggesting that selection for increased density and MOE, and reduced MFA in the absence of selection for growth will result in a genetic loss for growth rate.

Direct Effects of Wood Characteristics on Pulp and Handsheet Properties of Eucalyptus globulus

Holzforschung ◽  
2002 ◽  
Vol 56 (3) ◽  
pp. 244-252 ◽  
Author(s):  
R. Wimmer ◽  
G.M. Downes ◽  
R. Evans ◽  
G. Rasmussen ◽  
J. French
Keyword(s):  
Wood Density ◽  
Eucalyptus Globulus ◽  
Strong Predictor ◽  
Fibre Length ◽  
Microfibril Angle ◽  
Wood Properties ◽  
Bending Stiffness ◽  
Pulp Yield ◽  
Paper Properties ◽  
Handsheet Properties

Summary Fibre length, microfibril angle and wood density were measured in eight-year-old Eucalyptus globulus clones planted on three different sites. Samples were chipped and pulped, and the pulps beaten prior to preparation of 60g/m2 handsheets. Eleven physical handsheet properties, together with pulp freeness, were measured using standard methods. Direct relationships between wood properties and pulp/paper properties were studied using path analysis. Fibre length had a strong, direct effect on tear index, bending stiffness, freeness, pulp yield and active alkali consumption. Wood density was a strong predictor of most handsheet properties: high density woods produced bulkier, more porous sheets with higher bending stiffness, while lower density woods produced smoother, denser sheets with higher tensile strength.The effect of microfibril angle was small and more evident in handsheets made from beaten pulp.

Implications of selection history on genetic architecture of growth, form, and wood-quality traits in Pinus radiata

2008 ◽  
Vol 38 (9) ◽  
pp. 2372-2381 ◽  
Author(s):  
S. Kumar ◽  
R. D. Burdon ◽  
G. T. Stovold ◽  
L. D. Gea
Keyword(s):  
Wood Density ◽  
Pinus Radiata ◽  
Wood Quality ◽  
Genetic Correlations ◽  
Wood Properties ◽  
Radiata Pine ◽  
Single Pair ◽  
Narrow Sense Heritability ◽  
Tree Form ◽  
Selection History

Clonal trials of Pinus radiata D. Don (radiata pine), representing two populations (or breeds), one selected for growth and form (GF) and the other selected for high wood density as well as growth and form (HD), were replicated on two low-altitude New Zealand sites: Tarawera (pumice soil, 38°08′S) and Woodhill (coastal dune, 36°42′S). The GF material comprised 33 pair-crosses (19 parents) × 10 clones, and the HD material comprised 19 single-pair crosses (35 parents) × 10 clones, with six ramets per clone per site. Diameter (DBH), two tree-form variables, and needle retention (NRA) were assessed 8 years after planting, and wood density (DEN), acoustic velocity, and collapse were assessed 9 years after planting. The site differences were generally expressed more strongly in the GF population. Estimated genetic parameters were mostly similar for the two breeds, except that genotypic correlation between DBH and DEN was apparently zero in the HD population. Estimated broad-sense heritabilities (H2) were generally markedly higher than narrow-sense heritability estimates (h2), except with DEN. Estimated between-site type-B clonal genotypic correlations were generally high (>0.8) for wood properties. Overall, DBH showed adverse genetic correlations with wood properties. The Elite/Breed strategy appeared to be helpful in combating adverse genetic correlations.

scholarly journals Effect of number of annual rings and tree ages on genomic predictive ability for solid wood properties of Norway spruce 

2020 ◽  
Author(s):  
Linghua Zhou(Former Corresponding Author) ◽  
Zhiqiang Chen ◽  
Lars Olsson ◽  
Thomas Grahn ◽  
Bo Karlsson ◽  
...  
Keyword(s):  
Norway Spruce ◽  
Microfibril Angle ◽  
Predictive Ability ◽  
Tree Breeding ◽  
Wood Properties ◽  
Tree Age ◽  
Breeding Cycle ◽  
Standing Tree ◽  
Increment Core ◽  
The Impact

Abstract BACKGROUND: Genomic selection (GS) or genomic prediction is considered as a promising approach to accelerate tree breeding and increase genetic gain by shortening breeding cycle, but the efforts to develop routines for operational breeding are so far limited. We investigated the predictive ability (PA) of GS based on 484 progeny trees from 62 half-sib families in Norway spruce (Picea abies (L.) Karst.) for wood density, modulus of elasticity (MOE) and microfibril angle (MFA) measured with SilviScan, as well as for measurements on standing trees by Pilodyn and Hitman instruments. RESULTS: GS predictive abilities were comparable with those based on pedigree-based prediction. Marker-based PAs were generally 25-30% higher for traits density, MFA and MOE measured with SilviScan than for their respective standing tree-based method which measured with Pilodyn and Hitman. Prediction accuracy (PC) of the standing tree-based methods were similar or even higher than increment core-based method. 78-95% of the maximal PAs of density, MFA and MOE obtained from coring to the pith at high age were reached by using data possible to obtain by drilling 3-5 rings towards the pith at tree age 10-12. CONCLUSIONS: This study indicates standing tree-based measurements is a cost-effective alternative method for GS. PA of GS methods were comparable with those pedigree-based prediction. The highest PAs were reached with at least 80-90% of the dataset used as training set. Density could be conducted at an earlier age than for MFA and MOE. Operational breeding can also be optimized by training the model at an earlier age or using 3 to 5 outermost rings at tree age 10 to 12 years, thereby shortening the cycle and reducing the impact on the tree.

Anatomical properties of Hibiscus macrophyllus and its mature wood development

IAWA Journal ◽  
2021 ◽  
Vol 42 (4) ◽  
pp. 475-485
Author(s):  
Efrida Basri ◽  
Ratih Damayanti ◽  
Atmawi Darwis ◽  
Saefudin ◽  
Imam Wahyudi
Keyword(s):  
Wood Density ◽  
Fibre Length ◽  
Microfibril Angle ◽  
Mature Wood ◽  
Tree Age ◽  
Vessel Element ◽  
Plantation Forest ◽  
X Ray Diffraction ◽  
Wood Development ◽  
Anatomical Properties

Abstract The Hibiscus macrophyllus tree is widely planted in Indonesia especially on Java Island. It has several advantages to be developed commercially as a community or plantation forest compared to the famous introduced species Falcataria moluccana and Anthocephalus spp., including faster growth, higher wood density, and better stem morphology (straighter, more rounded, and lesser branches). However, information about the basic properties of this wood grown in plantations is limited. This study aimed to investigate the anatomical properties of H. macrophyllus and their variation at three ages (8, 12 and 16 years old), as well as to predict the mature wood development by using radial variation in fiber length, microfibril angle (MFA), and wood density from pith toward the bark as the indicators. The wood samples were obtained from a community forest area at Ciamis Regency, West Java Province. Furthermore, anatomical characteristics were examined through wood slides following the IAWA List, while fibre and vessel element dimensions were measured through macerated specimens prepared by modified Franklin’s method. The MFA was determined by X-Ray Diffraction, while wood density was measured in line with British Standard 373-57. The results showed that the anatomical structures were not influenced by tree age, except for wood porosity, and fibre and vessel element dimensions. The 16-year-old tree tended to be semi-ring-porous, the younger trees were diffuse-porous, while the fiber and vessel element length, as well as the diameter, were decreased. Meanwhile, the wall thickness was increased. The fibre length, MFA, and wood density were useful indicators for wood maturity that seemed to be developed at about 11 years of age.

Intra-specific competition and the radial development of wood density, microfibril angle and modulus of elasticity in plantation-grown Eucalyptus nitens

Trees ◽  
2012 ◽  
Vol 26 (6) ◽  
pp. 1771-1780 ◽  
Author(s):  
Jane Medhurst ◽  
Geoff Downes ◽  
Maria Ottenschlaeger ◽  
Chris Harwood ◽  
Rob Evans ◽  
...  
Keyword(s):  
Wood Density ◽  
Modulus Of Elasticity ◽  
Microfibril Angle ◽  
Eucalyptus Nitens

Evidence for associations between SilviScan-measured wood properties and intraring checking in a study of twenty-nine 6-year-old Pinus radiata

2005 ◽  
Vol 35 (5) ◽  
pp. 1156-1172 ◽  
Author(s):  
Roderick D Ball ◽  
Mike S McConchie ◽  
Dave J Cown
Keyword(s):  
Cell Wall ◽  
Wood Density ◽  
Wall Thickness ◽  
Pinus Radiata ◽  
A Priori ◽  
Wood Property ◽  
Wood Properties ◽  
Good Evidence ◽  
Cell Wall Thickness ◽  
Tree Level

Wood anatomical characteristics of twenty-nine 6-year-old Pinus radiata D. Don trees, selected to represent the extremes of intraring checking, were assessed with SilviScan. Derived ring-level summary variables were calculated from the SilviScan pith-to-bark wood property traces, based on ring means, standard deviations, and quantiles. Incidence of checking was assessed on discs that had been oven dried using a standardized method, and evidence for associations between wood characteristics and checking was evaluated at the tree and ring level using Bayesian statistical methods. Bayes factors of 39.1, 14.9, and 7.8 were obtained at the tree level, representing moderate to good evidence for associations between ring medians of wood density, tracheid radial diameter, and cell wall thickness, respectively. Increasing wood density, decreasing tracheid radial diameter, and increasing cell wall thickness were associated with reduced incidence of checking. These are parameters expected a priori to contribute to tracheid cells' resistance to collapse under drying stresses. A generalized linear model with radial diameter and cell wall thickness as independent variables had a concordance of 83% for predicting the checking status of trees.

Temporal variation in δ13C, wood density and microfibril angle in variously irrigated Eucalyptus nitens

2009 ◽  
Vol 36 (1) ◽  
pp. 1 ◽  
Author(s):  
David M. Drew ◽  
E. Detlef Schulze ◽  
Geoffrey M. Downes
Keyword(s):  
Water Stress ◽  
Temporal Variation ◽  
Wood Density ◽  
Stable Carbon Isotope ◽  
Microfibril Angle ◽  
Wood Property ◽  
Wood Properties ◽  
Eucalyptus Nitens ◽  
Stable Carbon ◽  
Over Time

Wood can serve as a record of past climate, recording tree responses to changing conditions. It is also valuable in understanding tree responses to environment to optimise forest management. Stable carbon isotope ratios (δ13C), wood density and microfibril angle (MFA) are potentially useful wood property parameters for these purposes. The goal of this study was to understand how δ13C varied over time in response to cycles of soil drying and wetting and to variation in temperature in Eucalyptus nitens Deane & Maiden, in concert with wood density and MFA. δ13C increases did not necessarily occur when water stress was highest, but, rather, when it was relieved. Our hypothesis is that this was a result of the use of previously fixed carbohydrate reserves when growth and metabolic activity was resumed after a period of dormancy. MFA in particular showed concomitant temporal variation with δ13C. A peak in δ13C may not coincide temporally with an increase in water stress, but with a decrease, when higher growth rates enable the final incorporation of earlier stored photosynthate into mature wood. This has implications for using δ13C as a tool to understand past environmental conditions using radial measurements of wood properties. However, interpreting this data with other wood properties may be helpful for understanding past tree responses.

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