Intertree competition in uneven-aged ponderosa pine stands

2003 ◽  
Vol 33 (9) ◽  
pp. 1719-1726 ◽  
Author(s):  
C W Woodall ◽  
C E Fiedler ◽  
K S Milner

Intertree competition indices and effects were examined in 14 uneven-aged ponderosa pine (Pinus ponderosa var. scopulorum Engelm.) stands in eastern Montana. Location, height, diameter at breast height (DBH), basal area increment, crown ratio, and sapwood area were determined for each tree (DBH >3.8 cm) on one stem-mapped plot (0.2-0.4 ha) in each sample stand. Based on tree locations, various competition indices were derived for each sample tree and correlated with its growth efficiency by diameter class. In addition, trends in individual tree attributes by diameter class and level of surrounding competition were determined. For trees with a DBH <10 cm, growth efficiency was most strongly correlated with the sum of surrounding tree heights within 10.6 m. The index most highly correlated for larger trees was the sum of surrounding basal area within 6.1 m. Regardless of tree size, individual tree growth efficiency, basal area increment, and crown ratio all decreased under increasing levels of competition, with the effect more pronounced in smaller trees. These results suggest that individual trees in uneven-aged stands experience competition from differing sources at varying scales based on their size, with response to competition diminishing as tree size increases.

2004 ◽  
Vol 34 (11) ◽  
pp. 2217-2229 ◽  
Author(s):  
Douglas B Mainwaring ◽  
Douglas A Maguire

Basal area and height growth were analyzed for individual trees in uneven-aged ponderosa pine (Pinus ponderosa Dougl. ex Laws.) and lodgepole pine (Pinus contorta Dougl. ex. Loud.) stands in central Oregon. Basal area growth was modeled as a function of other stand and tree variables to address three general objectives: (1) to compare the predictive ability of distance-dependent versus distance-independent stand density variables; (2) to determine the degree to which small trees negatively affect the growth of overstory trees; and (3) to test for differences in growth efficiency between species and between indices of spatial occupancy used to define efficiency (area potentially available, crown projection area, and a surrogate for total tree leaf area). Distance-dependent variables were found to improve growth predictions when added to models with only distance-independent variables, and small trees were found to have a quantifiably negative effect on the growth of larger trees. While volume growth efficiency declined with increasing levels of spatial occupancy for lodgepole pine, ponderosa pine volume growth efficiency was greatest at the highest levels of crown base sapwood area and crown projection area. The behavior in ponderosa pine resulted from the previously recognized correlation between tree height and total leaf area or crown size. The final statistical models distinguished between the positive effect of relative height and the negative effect of increasing tree size.


2003 ◽  
Vol 33 (3) ◽  
pp. 435-443 ◽  
Author(s):  
Daniel Mailly ◽  
Sylvain Turbis ◽  
David Pothier

A current trend in the development of forest stand models is to use spatially explicit, individual-tree information to simulate forest dynamics with increased accuracy. By adding spatial information, such as tree coordinates, crown shape, and size, it is hypothesized that the computation of the model's driving function is improved over traditional competition indices, especially when simulating multistoried stands. In this paper, we want to test whether computationally demanding competition indices outperform traditional indices in predicting mean basal area increment. The study was undertaken in old, uneven-aged black spruce (Picea mariana (Mill.) BSP) stands in northeastern Quebec, Canada. The predictability of individual tree growth rates was related to crown dimensions and other stand and tree variables measured in the field. Data were collected from 90 trees coming from stands of varying site quality (range 9.6–16.5 m height at 50 years, age taken at 1 m) and age (range 66–257 years). Hegyis's distance-dependent competition index was found to be the most strongly correlated competition measure (r = 0.57) with mean basal area growth of the last 20 years. This value, 12% higher than the value obtained from the best distance-independent competition index (r = 0.45), clearly shows that precision gains can be achieved when estimating basal area increment with spatial indices in black spruce stands. Using indices computed from virtual hemispherical images did not prove superior to simpler distance-dependent indices based on their individual correlations with basal area increment. When included in a basal area increment model for the last 20 years of growth, however, the gains in precision were comparable to Hegyi's competition index. This indicates that indices derived from a hemispherical approach have some value in spatially explicit forest simulations models but that further tests using younger stands are needed to confirm this result in black spruce stands.


2013 ◽  
Vol 43 (4) ◽  
pp. 311-320 ◽  
Author(s):  
Jianwei Zhang ◽  
Martin W. Ritchie ◽  
Douglas A. Maguire ◽  
William W. Oliver

We analyzed 45 years of data collected from three ponderosa pine (Pinus ponderosa Douglas ex P. Lawson & C. Lawson) levels-of-growing-stock installations in Oregon (OR) and northern California (CA), USA, to determine the effect of stand density regimes on stand productivity and mortality. We found that periodic annual increment (PAI) of diameter, basal area (BA), volume, and aboveground dry mass were significantly related to stand density index (SDI) and stand age at start of the period; the quadratic trends varied among sites. Precipitation departure from the normal for each period explained a significant amount of residual variation in all PAI variables except diameter. BA production did not change significantly as SDI exceeded 270 trees·ha−1 at the OR sites and 320 trees·ha−1 at the CA site. Stand productivity was the highest at Elliot Ranch (CA) and the least at Blue Mountains (OR). A similar trend held in growth efficiency under lower stand densities (SDI < 600). Most of the mortality was caused by Dendroctonus bark beetles in stands that exceeded SDI of 500 trees·ha−1. Limiting SDI was about 900 trees·ha−1, although plots at Elliot Ranch reached much higher than that. The results demonstrate that silvicultural control of stand density can be a powerful tool for reducing bark beetle caused mortality without sacrificing stand productivity.


2007 ◽  
Vol 37 (11) ◽  
pp. 2096-2105 ◽  
Author(s):  
Kelsey Sherich ◽  
Amy Pocewicz ◽  
Penelope Morgan

Trees respond to edge-to-interior microclimate differences in fragmented forests. To better understand tree physiological responses to fragmentation, we measured ponderosa pine ( Pinus ponderosa Dougl. ex P. & C. Laws) and Douglas-fir ( Pseudotsuga menziesii (Mirbel) Franco) leaf area, crown ratios, sapwood area, basal area (BA) growth rates, and BA growth efficiency at 23 long-established (>50 year) forest edges in northern Idaho. Trees located at forest edges had more leaf area, deeper crowns, higher BA growth rates, and more sapwood area at breast height than interior trees. Ponderosa pine had significantly higher BA growth efficiency at forest edges than interiors, but Douglas-fir BA growth efficiency did not differ, which may relate to differences in photosynthetic capacity and drought and shade tolerance. Edge orientation affected BA growth efficiency, with higher values at northeast-facing edges for both species. Edge effects were significant even after accounting for variation in stand density, which did not differ between the forest edge and interior. Although edge trees had significantly greater canopy depth on their edge-facing than forest-facing side, sapwood area was evenly distributed. We found no evidence that growing conditions at the forest edge were currently subjecting trees to stress, but higher leaf area and deeper crowns could result in lower tolerance to future drought conditions.


2007 ◽  
Vol 37 (2) ◽  
pp. 343-355 ◽  
Author(s):  
Nate G. McDowell ◽  
Henry D. Adams ◽  
John D. Bailey ◽  
Thomas E. Kolb

We examined the response of growth efficiency (GE), leaf area index (LAI), and resin flow (RF) to stand density manipulations in ponderosa pine ( Pinus ponderosa Dougl. ex Laws.) forests of northern Arizona, USA. The study used a 40 year stand density experiment including seven replicated basal area (BA) treatments ranging from 7 to 45 m2·ha–1. Results were extended to the larger region using published and unpublished datasets on ponderosa pine RF. GE was quantified using basal area increment (BAI), stemwood production (NPPs), or volume increment (VI) per leaf area (Al) or sapwood area (As). GE per Al was positively correlated with BA, regardless of numerator (BAI/Al, NPPs/Al, and VI/Al; r2 = 0.84, 0.95, and 0.96, respectively). GE per As exhibited variable responses to BA. Understory LAI increased with decreasing BA; however, total (understory plus overstory) LAI was not correlated with BA, GE, or RF. Opposite of the original research on this subject, resin flow was negatively related to GE per Al because Al/As ratios decline with increasing BA. BAI, and to a lesser degree BA, predicted RF better than growth efficiency, suggesting that the simplest measurement with the fewest assumptions (BAI) is also the best approach for predicting RF.


2021 ◽  
Vol 13 (12) ◽  
pp. 2297
Author(s):  
Jonathon J. Donager ◽  
Andrew J. Sánchez Meador ◽  
Ryan C. Blackburn

Applications of lidar in ecosystem conservation and management continue to expand as technology has rapidly evolved. An accounting of relative accuracy and errors among lidar platforms within a range of forest types and structural configurations was needed. Within a ponderosa pine forest in northern Arizona, we compare vegetation attributes at the tree-, plot-, and stand-scales derived from three lidar platforms: fixed-wing airborne (ALS), fixed-location terrestrial (TLS), and hand-held mobile laser scanning (MLS). We present a methodology to segment individual trees from TLS and MLS datasets, incorporating eigen-value and density metrics to locate trees, then assigning point returns to trees using a graph-theory shortest-path approach. Overall, we found MLS consistently provided more accurate structural metrics at the tree- (e.g., mean absolute error for DBH in cm was 4.8, 5.0, and 9.1 for MLS, TLS and ALS, respectively) and plot-scale (e.g., R2 for field observed and lidar-derived basal area, m2 ha−1, was 0.986, 0.974, and 0.851 for MLS, TLS, and ALS, respectively) as compared to ALS and TLS. While TLS data produced estimates similar to MLS, attributes derived from TLS often underpredicted structural values due to occlusion. Additionally, ALS data provided accurate estimates of tree height for larger trees, yet consistently missed and underpredicted small trees (≤35 cm). MLS produced accurate estimates of canopy cover and landscape metrics up to 50 m from plot center. TLS tended to underpredict both canopy cover and patch metrics with constant bias due to occlusion. Taking full advantage of minimal occlusion effects, MLS data consistently provided the best individual tree and plot-based metrics, with ALS providing the best estimates for volume, biomass, and canopy cover. Overall, we found MLS data logistically simple, quickly acquirable, and accurate for small area inventories, assessments, and monitoring activities. We suggest further work exploring the active use of MLS for forest monitoring and inventory.


1973 ◽  
Vol 3 (4) ◽  
pp. 495-500 ◽  
Author(s):  
James A. Moore ◽  
Carl A. Budelsky ◽  
Richard C. Schlesinger

A new competition index, modified Area Potentially Available (APA), was tested in a complex unevenaged stand composed of 19 different hardwood species. APA considers tree size, spatial distribution, and distance relationships in quantifying intertree competition and exhibits a strong correlation with individual tree basal area growth. The most important characteristic of APA is its potential for evaluating silvicultural practices.


1989 ◽  
Vol 19 (1) ◽  
pp. 113-121 ◽  
Author(s):  
L. E. Frelich ◽  
J. G. Bockheim ◽  
J. E. Leide

Basal-area increment and chemical composition of xylem wood were measured in three old-growth (ca. 75–100 years) white pine (Pinusstrobus L.) and three sugar maple (Acersaccharum) Marsh.) stands across a pH and SO4 gradient in precipitation in Wisconsin. In 1986 the volume-weighted mean pH and SO4 content of precipitation ranged from 4.5 to 5.0 and from 21 to 11 kg • ha−1, respectively, from southeastern to northwestern Wisconsin. With one exception (a white pine site at Point Beach in eastern Wisconsin), basal-area increment increased from the 1890s until the 1950s (sugar maple) or 1970s (white pine), then levelled off. Growth efficiency, estimated as the ratio of basal area to exposed crown area or crown volume for the 1980–1985 period, was similar for sugar maple across the gradient; however, growth efficiency of white pine was lower at Point Beach than at the two northern Wisconsin sites. Lead concentrations in xylem wood of both species have increased with time, except at Crotte Creek in northwestern Wisconsin, and Pb concentrations in xylem wood of both species were significantly greater in southeastern than in northwestern Wisconsin. Sulfur concentrations in xylem wood of white pine have increased since the 1960s at Point Beach and at one site in north central Wisconsin; S concentrations are significantly greater for both species in southeastern than in northern Wisconsin. Concentrations of Ca, Mg, and K in xylem wood of sugar maple have decreased over the past century. Whereas xylem wood concentrations of Mn and Zn generally show no age-related trends, Fe and P concentrations have increased markedly at all sites, particularly during the past decade. Although additional research is needed to determine the potential of dendrochemistry in evaluating the consequences of environmental pollution, the age- and site-related trends in chemical composition of xylem wood of white pine and sugar maple appear to be related to vehicular emissions (Pb), air pollution (S), migration along ray paths during conversion of sapwood into heartwood (P, Fe, Ca, K, Mg), and possibly reallocation of nutrients from the labile soil pool to perennial tree tissues during stand development (Ca, K, Mg).


1990 ◽  
Vol 5 (3) ◽  
pp. 79-82 ◽  
Author(s):  
William W. Oliver

Abstract Growth and stand development of ponderosa pine (Pinus ponderosa) were monitored for 20 years after planting at five different square spacings (6, 9, 12, 15, and 18 ft) in the presence or absence of competing shrubs on the westside Sierra Nevada. Mean tree size was positively correlated and stand values negatively correlated with spacing in the absence of competing shrubs. Trees growing with competing shrubs attained 76% of the diameter, 80% of the height, and 58% of the cubic volume of trees free of shrub competition when all spacings were combined. This study suggests that the major effect of shrub competition in ponderosa pine plantations on good sites is to lengthen the rotation. West. J. Appl. For. 5(3):79-82, July 1990.


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