Productivity of aspen forests in northeastern Minnesota, U.S.A., as related to stand composition and canopy structure

2001 ◽  
Vol 31 (6) ◽  
pp. 1019-1029 ◽  
Author(s):  
Christopher B Edgar ◽  
Thomas E Burk
Keyword(s):  
Populus Tremuloides ◽  
Vertical Structure ◽  
Canopy Structure ◽  
Abies Balsamea ◽  
Basal Area ◽  
Balsam Fir ◽  
Annual Increment ◽  
Tree Species Composition ◽  
Residual Variation ◽  
Paper Birch

Relationships among stand composition, stemwood productivity, and canopy structure were investigated for 55 study areas in northeastern Minnesota. Tree species composition among study areas was deliberately allowed to vary. Aspen, primarily quaking (Populus tremuloides Michx.) and to a lesser degree bigtooth (Populus grandidentata Michx.), was a significant component of every study area. The two most common associates were paper birch (Betula papyrifera Marsh.) and balsam fir (Abies balsamea (L.) Mill.). Productivity was defined as mean annual increment of tree stemwood volume and ranged from 3.3 to 12.6 m3·ha–1·year–1. A multiple-regression approach was used to investigate the relationships between productivity, stand composition, and canopy structure. Base models relating productivity to aspen site index, aspen cohort age, and total basal area were developed. Measures of stand composition and canopy vertical structure were added to the base models, and their significance in explaining residual variation in productivity was tested. Productivity was found to be negatively correlated with stand composition and canopy vertical structure, with all other factors held constant. Pure aspen and single-canopied stands were expected to be the most productive. Exceptions were present in the data: the two most productive stands were vertically stratified, aspen – balsam fir – paper birch mixtures.

The effect of light availability and basal area on cone production in Abies balsamea and Picea glauca

2002 ◽  
Vol 80 (4) ◽  
pp. 370-377 ◽  
Author(s):  
David F Greene ◽  
Christian Messier ◽  
Hugo Asselin ◽  
Marie-Josée Fortin
Keyword(s):  
Populus Tremuloides ◽  
Picea Glauca ◽  
Light Availability ◽  
Abies Balsamea ◽  
Basal Area ◽  
White Spruce ◽  
Balsam Fir ◽  
Forest Edges ◽  
Cone Production ◽  
Effect Of Light

Mean annual seed production is assumed to be proportional to basal area for canopy trees, but it is not known if subcanopy trees produce fewer seeds than expected (given their size) because of low light availability. Ovulate cone production was examined for balsam fir (Abies balsamea (L.) Mill.) and white spruce (Picea glauca (Moench) Voss) in 1998 and for balsam fir in 2000 in western Quebec using subcanopy stems, near or far from forest edges, or (at one site) planted white spruce trees in fully open conditions. A very simple light model for transmission through mature trembling aspen (Populus tremuloides Michx.) crowns and through boles near forest edges was developed to account for the effect of light receipt on cone production. The enhanced light near forest edges (e.g., recent clearcuts) leads to about a doubling of cone production for subcanopy stems. The minimum subcanopy height for cone production far from an edge is about 10 m for balsam fir and 14 m for white spruce, with these minima decreasing near edges. By contrast, the minimum height for white spruce in a plantation (full light) is about 3 m. Accounting for light receipt leads to an increase in the explained variance.Key words: balsam fir, cone production, light model, regressions, subcanopy stems, white spruce.

scholarly journals Développement de sapinières éclaircies exposées à une épidémie de tordeuse des bourgeons de l'épinette

1998 ◽  
Vol 74 (1) ◽  
pp. 91-99 ◽  
Author(s):  
David Pothier
Keyword(s):  
Choristoneura Fumiferana ◽  
Abies Balsamea ◽  
Basal Area ◽  
Stand Density ◽  
Spruce Budworm ◽  
Balsam Fir ◽  
Growth And Yield ◽  
Annual Increment ◽  
Diameter Increment ◽  
Outbreak Period

In 1968, three intensities of low thinning were applied in 33-year-old balsam fir (Abies balsamea [L.] Mill.) stands of eastern Québec in order to remove 0, 20 and 30% of the total basal area. Sample plots were located in two distinct areas, one of which was annually sprayed with insecticides between 1978 and 1982 while the other was never protected against a spruce budworm (Choristoneura fumiferana [Clem.]) outbreak that occurred between 1976 and 1983. A total of 32 0.04-ha sample plots were periodically measured between 1968 and 1994. Low thinnings mostly removed non merchantable trees (DBH < 9.1 cm) and left 5 000 to 7 000 trees/ha while stand density of control plots reached 10 000 stems/ha. From 1968 to 1976, net annual increment in merchantable volume was around 8 m3/ha but no differences were detected between thinning levels. During the period of severe defoliation caused by spruce budworm (1976-1985), net annual increment in merchantable volume of sample plots sprayed with insecticides decreased by 50% but get back to its initial level after 1985. Net annual increment of sample plots that were never protected against spruce budworm was strongly negative during the outbreak period until the extinction of all trees was completed. While thinning level only slightly affected merchantable yield of sample plots during the 26-year period, mean annual increment in DBH was inversely proportional to stand density. This advantage of thinned plots could have been greater if stand density had been reduced further at the beginning of the study. Indeed, the analysis of the evolution of stand density as a function of mean volume of trees shows that mortality rate of all sample plots was similar to that given by the self-thinning rule. This suggests that even trees of most heavily thinned plots suffered from strong intra-specific competition. Key words: balsam fir, low thinning, spruce budworm, growth and yield, diameter increment, self-thinning rule

Effet d'une coupe d'ensemencement et du milieu de germination sur la régénération des sapinières boréales riches de seconde venue du Québec

2000 ◽  
Vol 76 (4) ◽  
pp. 643-652 ◽  
Author(s):  
Patricia Raymond ◽  
Jean-Claude Ruel ◽  
Marius Pineau
Keyword(s):  
Picea Glauca ◽  
Forest Cover ◽  
Seedling Survival ◽  
Mineral Soil ◽  
Abies Balsamea ◽  
Basal Area ◽  
White Spruce ◽  
Balsam Fir ◽  
Paper Birch ◽  
Cutting System

Stand regeneration failures, sometimes observed in rich second growth balsam fir stands, prompted a study in 1991 to assess the effects of the shelterwood cutting system on regeneration of balsam fir (Abies balsamea (L.) Mill.), white spruce (Picea glauca (Moench) Voss) and paper birch (Betula papyrifera Marsh). The factorial experiment design (split-plot) includes forest cover reduction (0% and 25% of basal area) in main plots and germination substrate (mineral soil, litter removed and control) in subplots. Results of five growing seasons showed that germination substrate was the main factor determining first-year seedling establishment, and that cover reduction became important for seedling survival and long-term establishment of the three species. A 25% canopy reduction combined with mineral seedbed treatment resulted in the best regeneration densities of the three species. Thus far, the results demonstrate the importance of combining humus disturbance to seed cutting in order to achieve regeneration goals set for the future stand. Key words: shelterwood cutting system, balsam fir, white spruce, paper birch, regeneration, seed cutting, germination substrate

Attributes of forest strips used by snowshoe hare in winter within clear-cut boreal landscapes

2005 ◽  
Vol 35 (10) ◽  
pp. 2521-2527 ◽  
Author(s):  
François Potvin ◽  
Normand Bertrand ◽  
Jean Ferron
Keyword(s):  
Abies Balsamea ◽  
Basal Area ◽  
Balsam Fir ◽  
Snowshoe Hare ◽  
Forest Patches ◽  
Clear Cut ◽  
Important Prey ◽  
South Central ◽  
Snowshoe Hares ◽  
Riparian Strips

The snowshoe hare (Lepus americanus Erxleben) is an important prey for many predators in the boreal forest. In this biome, clear-cut landscapes are generally large and consist of aggregated cutting blocks separated by narrow forest strips (typically 60–100 m wide). To identify attributes of forest strips that are important for snowshoe hares, we measured the use of strips using track counts over two winters in six clear-cut landscapes (23–256 km2) in south-central Quebec. Surveys were conducted in 20 riparian strips (RS), 20 upland strips (US), and 15 control sites (CO) at the periphery of clear-cut landscapes. Overall, 392 signs of hare presence were recorded along 50 km of transects. Snowshoe hares were present in one-third of the strips surveyed and were five times less abundant in US and RS than in CO. The species avoided strip edges. Hares were more common in the wider strips (>100 m), in the strips adjacent to residual forest patches (≥25 ha), or in those having a denser shrub canopy, which is often associated with a greater basal area in balsam fir (Abies balsamea (L.) Mill.). To maintain snowshoe hare at moderate densities in large clear-cut landscapes, we suggest leaving uncut forest strips >100 m wide in areas having a good shrub cover with presence of balsam fir.

AN ANALYSIS OF MEASURES OF DENSITY IN EVEN-AGED BALSAM FIR AND JACK PINE STANDS

1964 ◽  
Vol 40 (4) ◽  
pp. 474-481 ◽  
Author(s):  
P. E. Vezina
Keyword(s):  
Pinus Banksiana ◽  
Abies Balsamea ◽  
Basal Area ◽  
Stand Density ◽  
Jack Pine ◽  
Balsam Fir ◽  
Aerial Photographs ◽  
Growing Stock ◽  
The One ◽  
Crown Closure

The concept of stand density in relation to thinning is examined and its development over the years is discussed. Present difficulties of objectively measuring stand density are recognized and probable future trends towards the development of better formulae to express stand density are outlined. Researchers should continue to collect information on interrelationships among stand variables. Certain merits accrue from description of stand density in terms of variables, such as crown closure, that can be measured with some precision from aerial photographs. Conversely, valid estimates of crown closure which are often difficult to obtain by means of devices from the ground, could be predicted from stand density. Three stand variables, used as expressions of stand density, were tested in crown closure simple regressions in even-aged, unmanaged stands of balsam fir (Abies balsamea (L.) Mill.) and jack pine (Pinus banksiana Lamb.). These are: total number of trees, number of trees 4 inches and up, and basal area per acre. The strongest relationship found was the one where crown closure is compared with basal area; it was stronger for jack pine than for balsam fir. This was explained by differences in tolerance among the two species. The significance of these relationships for the stand development, and the feasibility of using height-and diameter-based indices as measures of growing stock in studies of yield are discussed.

Thinning ponderosa pine (Pinus ponderosa) stands reduces mortality while maintaining stand productivity

2013 ◽  
Vol 43 (4) ◽  
pp. 311-320 ◽  
Author(s):  
Jianwei Zhang ◽  
Martin W. Ritchie ◽  
Douglas A. Maguire ◽  
William W. Oliver
Keyword(s):  
Pinus Ponderosa ◽  
Ponderosa Pine ◽  
Bark Beetles ◽  
Basal Area ◽  
Stand Density ◽  
Growth Efficiency ◽  
Stand Age ◽  
Annual Increment ◽  
Stand Productivity ◽  
Residual Variation

We analyzed 45 years of data collected from three ponderosa pine (Pinus ponderosa Douglas ex P. Lawson & C. Lawson) levels-of-growing-stock installations in Oregon (OR) and northern California (CA), USA, to determine the effect of stand density regimes on stand productivity and mortality. We found that periodic annual increment (PAI) of diameter, basal area (BA), volume, and aboveground dry mass were significantly related to stand density index (SDI) and stand age at start of the period; the quadratic trends varied among sites. Precipitation departure from the normal for each period explained a significant amount of residual variation in all PAI variables except diameter. BA production did not change significantly as SDI exceeded 270 trees·ha−1 at the OR sites and 320 trees·ha−1 at the CA site. Stand productivity was the highest at Elliot Ranch (CA) and the least at Blue Mountains (OR). A similar trend held in growth efficiency under lower stand densities (SDI < 600). Most of the mortality was caused by Dendroctonus bark beetles in stands that exceeded SDI of 500 trees·ha−1. Limiting SDI was about 900 trees·ha−1, although plots at Elliot Ranch reached much higher than that. The results demonstrate that silvicultural control of stand density can be a powerful tool for reducing bark beetle caused mortality without sacrificing stand productivity.

One and two years impact of commercial thinning on spruce budworm feeding ecology and host tree foliage production and chemistry

1996 ◽  
Vol 72 (4) ◽  
pp. 393-398 ◽  
Author(s):  
Éric Bauce
Keyword(s):  
Choristoneura Fumiferana ◽  
Soluble Sugars ◽  
Abies Balsamea ◽  
Basal Area ◽  
Spruce Budworm ◽  
Balsam Fir ◽  
First Year ◽  
Commercial Thinning ◽  
Mill Stand ◽  
Second Year

Field rearing experiments of spruce budworm, Choristoneura fumiferana (Clem.), were conducted in conjunction with foliar chemical analyses, one and two years after a commercial thinning (removal of 25% stand basal area) in a 50-year-old balsam fir, Abies balsamea (L.) Mill., stand. The first year after thinning, spruce budworm larvae reared on the residual trees developed five days faster and removed 43% more foliage than those reared on control trees, but in the second year they developed two days faster and removed 37% more foliage. The increase in larval development rate was related to an increase in foliar soluble sugars while a reduction in foliar monoterpenes caused by the thinning apparently accounted for the greater amount of foliage ingested by the larvae. The first year after thinning, trees were more vulnerable to spruce budworm because there was no increase in foliage production and the trees were more heavily defoliated. However, in the second year trees were less vulnerable to the insect because there was an increase in foliage production that exceeded the increase in defoliation, hence a net gain in foliage. Results from this study showed that commercial thinning could reduce the vulnerability of balsam fir trees to spruce budworm if thinning is conducted two years prior to budworm outbreak, but the same silvicultural procedure could increase the vulnerability to the insect if it is conducted during an outbreak. Key words: spruce budworm, balsam fir, chemistry, thinning, defoliation

scholarly journals Contrasting Release Approaches for a Mixed Paper Birch (Betula papyrifera)–Quaking Aspen (Populus tremuloides) Stand

2008 ◽  
Vol 25 (3) ◽  
pp. 124-132 ◽  
Author(s):  
Eric K. Zenner ◽  
Klaus J. Puettmann
Keyword(s):  
Populus Tremuloides ◽  
Basal Area ◽  
Complete Removal ◽  
Quaking Aspen ◽  
Betula Papyrifera ◽  
Mixed Stands ◽  
Early Release ◽  
Paper Birch ◽  
Area And Volume

Abstract Early release from competitors can be used to influence the species composition, quality, and rate of development of young stands. Release strategies can vary in intensity, ranging from complete removal of competitors and infrequent future entries (early, heavy, infrequent [EHI]) to lighter entries that are repeated more frequently (early, light, often [ELO]). It is unclear, however, which strategy is more successful for producing high-quality birch sawtimber (Betula papyrifera Marsh.) in mixed stands with aspen (Populus tremuloides Michx.). We evaluated the effects of various release intensities on the growth and mortality of a 16–18-ft-tall natural aspen–paper birch stand in Minnesota following density reductions from 1,500–3,000 trees ac−1 (trees per acre [TPA]) to 750 (ELO750), 500 (EHI500), and 250 (EHI250) TPA. After 6 years, paper birch was overtopped by aspen and contributed only 14% of basal area in control plots, but it occupied all diameter classes and contributed 77–87% of basal area in release plots. The basal area and volume of all paper birch and of only paper birch crop trees (100 largest TPA) were highest in lightly released ELO750 and lowest in control plots. Growth of mean quadratic diameter, basal area, and volume of paper birch was 2–3 times higher in release plots but independent of release intensity. Early release is necessary to maintain paper birch dominance, but there is no short-term advantage among treatment intensities. Long-term growth simulations using the Forest Vegetation Simulator suggest that merchantable timber production was unaffected by release strategy but that the EHI250 strategy produced the most birch sawtimber (40 times as much as in ELO750).

scholarly journals Mortality Patterns Following Spruce Budworm Infestation in Unprotected Spruce-Fir Forests in Maine

2003 ◽  
Vol 20 (4) ◽  
pp. 148-153 ◽  
Author(s):  
Dale S. Solomon ◽  
Lianjun Zhang ◽  
Thomas B. Brann ◽  
David S. Larrick
Keyword(s):  
Abies Balsamea ◽  
Basal Area ◽  
Growth Function ◽  
Red Spruce ◽  
Balsam Fir ◽  
Stem Density ◽  
Cumulative Mortality ◽  
Time Period ◽  
Composition Classes ◽  
Annual Mortality

Abstract Cumulative and annual mortality of red spruce (Picea rubens Sarg.) and balsam fir [Abies balsamea (L.) Mill.] were examined over a 10 yr period to follow the mortality patterns in unprotected spruce-fir forests in northern Maine. Different mortality patterns were determined based on stand composition classes and merchantability classes. In general, balsam fir was more vulnerable to budworm attack, and reached 92–100% basal area mortality and 84–97% stem density mortality 12 yr after the start of the outbreak. Red spruce, in contrast, had approximately 32–59% basal area mortality and 30–66% stem density mortality during the same time period. Balsam fir mortality started 1 to 2 yr before spruce, while spruce mortality continued 2 to 3 yr after fir mortality was completed. Higher mortality was found in smaller trees than sawtimber-sized trees. Stands with hardwood components (30–70% in basal area) had the lowest mortality rate for both species. Furthermore, Schnute growth function (Schnute 1981) was used to characterize the cumulative mortality trajectories after the defoliation of spruce and fir by stand composition classes. The models estimated the time when annual mortality achieved maximum, the cumulative mortality at that time, and the asymptotic mortality over a long time period after the start of the attack. The information can provide guidelines for predicting protection strategies and scheduling salvage harvests.

Refining tree recruitment models

2003 ◽  
Vol 33 (1) ◽  
pp. 41-46 ◽  
Author(s):  
C Calogeropoulos ◽  
D F Greene ◽  
C Messier ◽  
S Brais
Keyword(s):  
Wind Speed ◽  
Populus Tremuloides ◽  
Abies Balsamea ◽  
Basal Area ◽  
Horizontal Wind ◽  
Dimensional Model ◽  
Forest Environment ◽  
Horizontal Wind Speed ◽  
One Dimensional ◽  
Cone Production

We used a micrometeorological dispersal model to simulate seed and seedling distributions derived from subcanopy balsam fir (Abies balsamea (L.) Mill.) source trees in a trembling aspen (Populus tremuloides Michx.) dominated forest. Our first objective was to determine the effect of substituting basal area for cone production as a proxy for seed output. The results showed that the r2 from the regression of predicted versus observed densities increased by ~5% for seeds and ~15% for seedling simulations. Our second objective was to determine the effects of changing the median horizontal wind speed. The median speed in this forest environment varies according to the proportion of leaves abscised. For values of the median expected wind speed between the extremes of leafless and full-canopy forests, the r2 of predicted versus observed varied between 0.35 and 0.49 for seeds and between 0.33 and 0.62 for seedling simulations. We demonstrated that the simple one-dimensional model can have added precision if the dispersal parameters are chosen so as to allow more fine-scale variation.

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