The thermal ecology of burying beetles: temperature influences reproduction and daily activity in Nicrophorus marginatus

2021 ◽  
Author(s):  
Maranda L. Keller ◽  
Daniel R. Howard ◽  
Carrie L. Hall
2020 ◽  
Vol 98 (9) ◽  
pp. 591-602
Author(s):  
K.W. Burke ◽  
J.D. Wettlaufer ◽  
D.V. Beresford ◽  
P.R. Martin

The coexistence of closely related species plays an important role in shaping local diversity. However, competition for shared resources can limit the ability of species to coexist. Many species avoid the costs of coexistence by diverging in habitat use, known as habitat partitioning. We examine patterns of habitat use in seven co-occurring species of burying beetles (genus Nicrophorus Fabricius, 1775), testing the hypothesis that Nicrophorus species partition resources by occupying distinct habitats. We surveyed Nicrophorus abundance and 54 habitat characteristics at 100 random sites spanning an environmentally diverse region of southeastern Ontario, Canada. We found that three species occupied distinct habitat types consistent with habitat partitioning. Specifically, Nicrophorus pustulatus Herschel, 1807, Nicrophorus hebes Kirby, 1837, and Nicrophorus marginatus Fabricius, 1801 appear to be specialists for forest canopy, wetlands, and open fields, respectively. In contrast, Nicrophorus orbicollis Say, 1825, Nicrophorus sayi Laporte, 1840, and Nicrophorus tomentosus Weber, 1801 appear to be generalists with wide breadths of habitat use. We were unable to identify the habitat associations of Nicrophorus defodiens Mannerheim, 1846. Our findings are consistent with habitat acting as an important resource axis along which some Nicrophorus species partition; however, divergence along other resource axes (e.g., temporal partitioning) also appears important for Nicrophorus coexistence.


Diversity ◽  
2021 ◽  
Vol 13 (12) ◽  
pp. 662
Author(s):  
Mark C. Belk ◽  
Peter J. Meyers ◽  
J. Curtis Creighton

The cost of reproduction hypothesis suggests that allocation to current reproduction constrains future reproduction. How organisms accrue reproductive costs and allocate energy across their lifetime may differ among species adapted to different resource types. We test this by comparing lifetime reproductive output, patterns of reproductive allocation, and senescence between two species of burying beetles, Nicrophorus marginatus and N. guttula, that differ in body size, across a range of carcass sizes. These two species of burying beetles maximized lifetime reproductive output on somewhat different–sized resources. The larger N. marginatus did better on large and medium carcasses while the smaller N. guttula did best on small and medium carcasses. For both species, reproduction is costly and reproduction on larger carcasses reduced lifespan more than reproduction on smaller carcasses. Carcass size also affected lifetime reproductive strategies. Each species’ parental investment patterns were consistent with terminal investment on carcasses on which they performed best (optimal carcass sizes). However, they exhibited reproductive restraint on carcass sizes on which they did not perform as well. Reproductive senescence occurred largely in response to carcass size. For both species, reproduction on larger carcasses resulted in more rapid senescence. These data suggest that whether organisms exhibit terminal investment or reproductive restraint may depend on type and amount of resources for reproduction.


2018 ◽  
Vol 2018 ◽  
pp. 1-7
Author(s):  
Ashlee N. Smith ◽  
Mark C. Belk

We tested whether brood parasitism could be successful between two co-occurring species of burying beetles, Nicrophorus guttula and Nicrophorus marginatus, and whether these species exhibit an adaptive response to brood parasitism by detecting and removing parasites. We cross-fostered larvae between broods of the two species and created mixed-species broods to simulate the addition of brood parasites. Brood parasites survived in both species’ broods. Nicrophorus marginatus culled 86% of brood parasites compared to 56% of their own larvae, and N. guttula culled 50% of brood parasites compared to 22% of their own larvae. Additionally, N. guttula brood parasites were significantly smaller than N. guttula that were raised by N. guttula parents, but N. marginatus brood parasites were significantly larger than N. marginatus that were raised by N. marginatus parents. This paper provides the first evidence that burying beetles can discriminate between their own larvae and other species’ larvae. We suggest that brood parasitism may be the selective force responsible for this ability.


2019 ◽  
Vol 20 (2) ◽  
pp. 283-298
Author(s):  
Kholid Mawardi ◽  
Cucu Nurzakiyah

The results of the study found that the responsibility of religious education of children in the family of Tablighi Jama'ah differed in terms of several conditions, namely first, when parents were not going to khuruj where both parents were responsible for children's education; secondly, when the father goes khuruj, then the mother is responsible for everything including children's education; third, when both parents go khuruj, then the responsibility of the child is left to other family members such as grandparents or their first adult children; and fourth, when the child goes to khuruj, where parents are responsible for children's religious education both mother and father. The pattern of the religious education in the Tablighi Jama'ah family in the village of Bolang is formed from several similarities held in the implementation of religious education, one of which is the daily activity that is carried out by the Tablighi Jama'at family. Al-Qur'an becomes one of the material given to children in the ta'lim. Children are taught how to read the Qur'an and memorize short letters such as Surat al-Falaq, al-Ikhlas, and so on. In addition to al-Qur'an, in this ta'lim there is a special study in the Tablighi Jama'ah, which is reading the book of fadhilah ‘amal, and the last is mudzakarah six characteristics.


2010 ◽  
Vol 19 (1-2) ◽  
pp. 127-147 ◽  
Author(s):  
Krister Hertting

Leading with Pedagogical Tact- a Challenge in Children's Sports in Sweden The purpose of this article is to elucidate and problemize meetings between children and leaders in children's sport. The competitive sport is high valuated in the Swedish society and sport for children is central in the Swedish youth politics. The foundation in Swedish sport, as well as in the other Nordic countries, has for a long time relied on voluntary commitment. Approximately 650 000 people are voluntary engaged as leaders in sport in Sweden and 70% of children between 7 and 14 years compete in sports clubs. There is, however, a tension in the Swedish sport system. The sports for children has double missions - ‘association nurturing’ and ‘competition nurturing’, missions which are not always in harmony. In the daily activity it is the voluntary leaders who have to deal with these missions, which creates a field of tension. In this article I argue for a bridge between these missions by a leadership based on pedagogical tact. The empirical outlook is a narrative based on statements from leaders, children and parents in a study dealing with voluntary leadership within children's football. In the end I argue that focusing on this bridge is a win-win situation, both for children and sports.


2017 ◽  
Vol 3 (3) ◽  
pp. 140
Author(s):  
Suci Rahmatia ◽  
Putri Wulandari ◽  
Nurul Khadiko ◽  
Fitria Gani Sulistya

<p><em>Abstrak </em><strong> - Antena merupakan alat pemancar yang akrab dengan aktifitas sehari-hari dan mudah sekali dijumpai, di rumah, di gedung, bahkan pada alat komunikasi yang digunakan. Salah satu antena yang sering digunakan adalah antena televisi. Antena televisi yang sering digunakan adalah Yagi-Uda yang biasanya dipakai sebagai outdoor antena dan antena dipole yang biasanya digunakan untuk indoor antena. Masing – masing jenis antena memiliki kriteria dan keuntungan berdasarkan dari kebutuhan penggunaannya. Baik antena dipole maupun antena Yagi-Uda memiliki perbedaan diantaranya adalah besar bandwidth, nilai gain, dan pola radiasi. Pada paper ini dapat diketahui bahwa bandwidth yang dimiliki antena yagi-uda lebih besar daripada antena dipole yakni 0.39943 MHz untuk antena yagi-uda dan 0.16569 MHz untuk antena dipole. Begitupula dengan besar Gain yang dimiliki antena Yagi-Uda (6.64 dBi) lebih besar dibandingkan dengan gain dari antena dipole (2.29 dBi). Perbedaan ini dikarenakan faktor elemen director dan ketebalannya.</strong></p><p><strong><br /></strong></p><p><strong><em>Kata Kunci</em></strong> – <em>Atena Televisi, Atena Yagi-Uda, Atena Dipole, Gain, Bandwidth</em></p><p><em> </em></p><p><em>Abstract</em> <strong>- Antenna is a transmitter tool that is familiar with daily activity and easy to find at home, in the building, even on the communication tool used. One of antenna that is often used is a television antenna. Television antennas are often used is Yagi-Uda which is usually used as an outdoor antenna and dipole antenna that is usually used for indoor antennas. Each type of antenna has the criteria and advantages based on the needs of its use. Both dipole antennas and Yagi-Uda antennas have differences among them are bandwidth, gain, and radiation pattern. In this paper it can be seen that the bandwidth of yagi-uda antenna is bigger than dipole antenna that is 0.39943 MHz for Yagi-Uda antenna and 0.16569 MHz for dipole antenna. Neither the large Gain of the Yagi-Uda antenna (6.64 dBi) is greater than the gain of the dipole antenna (2.29 dBi). This difference is due to element factor of director and its thickness.</strong></p><p><strong><br /></strong></p><p><strong><em>Keywords</em></strong><strong> – </strong><em>Television Antenna, Yagi-Uda Antenna, Dipole Antenna, Gain, Bandwidth</em><strong> </strong></p>


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